Late Quaternary climatic history of the Chilean Channels based on fossil pollen and beetle analyses,with an analysis of the modern vegetation and pollen rain

Fossil beetles and pollen were examined from an intermorainal bog at Puerto Edén, Isla Wellington, Chile (latitude 49°08'S, longitude 74°25'W). Wood from near the base of the section has an age of 12 960 ± 150 yr BP. Occurrence of flightless beetle species in the basal peat sample is evidence that some members of the biota survived the last glacial maximum in refugia. The assumption that the Chilean Channels were entirely ice-covered is incorrect. Plants and insects that invaded the deglaciated terrain were those of an Empetrum heathland in which patches of Nothofagus forest were restricted to sheltered locations. The climate supporting the heathland is inferred to have been windier and probably drier than that of the present day. From 13 000 yr BP to 9500 yr BP Nothofagus forest expanded, possibly in response to less windiness and more available moisture. Neither the fossil beetle nor pollen data support a return to significantly colder conditions between 11 000 and 10 000 yr BP at the time of the Younger Dryas Stade. From 9500 to 5500 yr BP the climate was as wet as that of the present day, based on an increased representation of the pollen of moorland plants and of aquatic beetle species. From 5500 to 3000 yr BP the climate was drier, as indicated by the expansion of Empetrum heath and the reduction in mesic habitats. From 3000 yr BP to the present-day mesic habitats dominated as the climate returned to a wetter mode. The alternatively wetter and drier episodes are attributed to latitudinal shifts in the position of storm tracks in the belt of Southern Westerlies.     

Glacial forest refugium in Howard Valley,South Island,New Zealand

Fossils of forest habitat beetles and leaves of Nothofagus menziesii provide evidence of a forest refugium at times between ca. 34 000 and ca. 18 500 cal. a BP at an upland site in Howard Valley, located adjacent to glaciated valleys in South Island, New Zealand. The stratigraphy of the glacial‐aged terrace sequence of organic‐rich silts and fluvial sand/gravels indicates that soil development occurred episodically for around 15 000 a. Fifty‐four beetle taxa represent seven habitat types: forest, forest or scrub, riparian and aquatic, litter, grass/tussock, marshland and moss habitats. Leaf and beetle fossils indicate that forest dominated by N. menziesii persisted at the site for most of the time period represented, and tree line taxa such as Taenarthrus sp. 1 (Carabidae) and Podocarpus sp. (Podocarpaceae) indicate that the site may represent the upper tree limit for full‐glacial time. The finding of forest at this elevated site adds to the growing fossil evidence for multiple forest refugia in New Zealand during the last glaciation and is consistent with the pollen records, which have consistently indicated the presence of forest species during the last glaciations. Copyright © 2009 John Wiley & Sons, Ltd.     

Cretocrenis burmanicus,the first Mesozoic amber inclusion of a water scavenger beetle (Coleoptera: Hydrophilidae)

Cretocrenis burmanicus, gen. et sp. nov. is described from Burmese amber and represents the oldest known amber inclusion of the family Hydrophilidae (Coleoptera: Polyphaga). The new genus resembles the small-bodied modern genera of the tribe Anacaenini and the subfamily Acidocerinae, and the morphology of the venter and male genitalia suggest that it belongs to the Horelophus+Crenitis clade of the tribe Anacaenini. The fossil is described in detail, illustrated, and compared with extant hydrophilid genera.     

Abundance, diversity and succession of aquatic Coleoptera and Heteroptera in a cluster of artificial ponds in the North German Lowlands

Our study describes and evaluates environmental influences on assemblages of aquatic Coleoptera and Heteroptera in artificial ponds situated near Lake Steinhude in Lower Saxony (Germany). We determined temporal dynamics and colonization patterns for 14 ponds of different age. In total, we recorded 4941 individuals that represented 87 species of aquatic beetles and bugs. Between 30 and 40 species were found in most of the ponds. Heteropteran species of the families Corixidae and Notonectidae acted as pioneer species in new ponds, while aquatic coleopterans predominated in older ponds. The results of Canonical Correspondence Analyses (CCA) showed that among the key factors affecting community structure were land use, vegetation cover, water chemistry and the age of the ponds. We found that the distribution of adjacent ponds on areas with different land use has a positive influence on the diversity and abundance of the aquatic insect fauna.     

Prophaenognatha, a New Aclopinae Genus from the Yixian Formation, China and Its Phylogenetic Position Based on Morphological Characters (Coleoptera: Scarabaeidae)

Prophaenognatha robusta gen. et sp. nov. (Scarabaeoidea: Scarabaeidae: Aclopinae), the best-preserved aclopine fossil so far, is described and illustrated from the Upper Jurassic–Lower Cretaceous Yixian Formation of the Jehol Biota, western Liaoning Province, NE China. The key to extinct and extant genera of Aclopinae is given and the monophyly of extant and fossil Aclopinae lineages is supported by five character states. The new taxon provides evidence about the evolution of Scarabaeoidea with its phylogenetic position inferred based on 68 morphological characters.     

Mid-Wisconsin seasonal temperatures reconstructed from fossil beetle assemblages in eastern North America: comparisons with other proxy records from the Northern Hemisphere

Mutual climatic range (MCR) analysis was applied to 15 North American beetle assemblages spanning the interval from > 52 000 to 17 200 yr BP, bracketing a Mid-Wisconsin interstadial interval. The analyses yielded estimates of mean July (TMAX) and mean January (TMIN) temperatures. The oldest assemblage (> 52 ka) yielded TMAX values 7.5–8°C lower than present and TMIN values 15–18°C lower than present. A Mid-Wisconsin interstadial warming dating from 43.5–39 ka was rapid and intense. At the peak of the warming event, about 42 ka, TMAX values were only 1–2°C lower than modern. This level of amelioration apparently lasted only about 2000–3000 yr. By 23.7 ka, TMAX values declined to 11.5–10°C lower than modern, but another, small-scale amelioration is indicated by assemblages dating from 20.5 to 19.7 ka. The interstadial event recorded from the site at Titusville, Pennsylvania closely matches the timing and intensity of the climate change estimated from British beetle faunas in the Upton Warren interstadial. Another warm interval (ca. 31–32.5 ka) has been documented from fossil beetle assemblages in Europe and North America. Copyright © 1999 John Wiley & Sons, Ltd.     

Middle Jurassic Coptoclavidae (Insecta:Coleoptera:Dytiscoidea) from China:a Good Example of Mosaic Evolution

<正>Adults of the aquatic coptoclavid beetles(Coleoptera:Adephaga:Dytiscoidea),described from four Middle Jurassic fossil localities in Inner Mongolia and Liaoning in northeastern China,are attributed to Daohugounectes primitives Wang,Ponomarenko and Zhang,2009,which was previously proposed after study of larvae.The generic name Timarchopsis Brauer,Redtenbacher and Ganglbauer,1889 is proposed as a substitute for the preoccupied and junior homonym Necronectes Ponomarenko,1977,non Milne-Edwards,1881. Furthermore,the subfamily name Necronectinae Ponomarenko,1977 is substituted by the available name Timarchopsinae.Daohugounectes is placed into Timarchopsinae because its adults have long,slightly apically widened tibiae and small femoral plates.The adults of this genus differ from those of other Timarchopsinae in the following features:antennae short and widened in the middle part;basal segments of protarsi not cut apically; metaventrite with a triangular plate.The larvae look like somewhat primitive forms in the subfamily Timarchopsinae.In contrast to these primitive larvae,the adults with some advanced characters can be regarded as among the most advanced forms in the subfamily Timarchopsinae,and probably represent a transition between the Timarchopsinae and Charanoscaphinae.Such mosaic evolution within Daohugounectes indicates that the evolutionary process of aquatic beetles is far more complex than previously thought.     

Palaeoenvironment and biogeography of a late MIS 3 fossil beetle fauna from South Taranaki,New Zealand

Fossil beetles from a last glacial landslide‐generated lake‐forest sequence aged 33 480–34 410 cal. yr BP (late Marine Isotope Stage 3 (MIS 3)) are identified from the Waitotara Valley in South Taranaki, North Island, New Zealand. The stratigraphy indicates that the landslide caused changes to local hydrology, resulting in the formation of a lake‐swamp environment and subsequent transition to forest. Fossil leaves suggest a forest dominated by Nothofagus menziesii, and radiocarbon ages indicate the site was forested for around 4000 yr. A fossil beetle‐based temperature estimate using the maximum likelihood envelope method indicates the climate was cooler than present day. The distributions of the fossil beetle taxa are examined and compared with the modern ecological patterns. The fossil fauna is very typical of a modern‐day Nelson (northern South Island) fauna. None of the beetle species is present in the modern South Taranaki fauna and many taxa such as Platypus caviceps, Alema paradoxa, Rhyzobius consors, Syrphetodes ater, Cyclaxyra impressa and species of Grynoma and Pycnomerus are now absent from part or all of the lower North Island. This is important because the lower North Island is currently an area of low diversity and endemism and these results suggest this biogeographical pattern stems from the last glaciation. Copyright © 2008 John Wiley & Sons, Ltd.     

Late Pleistocene Palaeoenvironments of the Southern Lake Agassiz Basin,USA

Macroscopic plant remains, pollen, insect and mollusc fossils recovered from a cut bank on the Red River in North Dakota, USA, provide evidence that an extensive wetland occupied the southern basin of Lake Agassiz from 10 230 to 9900 ¹⁴C yr BP. Marsh‐dwelling plants and invertebrates had colonised the surface of a prograding delta during the low‐water Moorhead Phase of Lake Agassiz. A species of Salix (willow) was abundant along distributary channels, and stands of Populus tremuloides (aspen), Ulmus sp. (elm), Betula sp. (birch), and Picea sp. (spruce) grew on the better‐drained sand bars and beach ridges. Most of the species of plants, insects, and molluscs represented as fossils are within their existing geographic ranges. Based on a few species with more northerly distributions, mean summer temperature may have been about 1–2°C lower than the present day. No change in species composition occurred in the transition from the Younger Dryas to Preboreal. At the time that the wetland existed, Lake Agassiz was draining either eastward to the North Atlantic Ocean or northwestward to the Arctic Ocean. The wetland was drowned during the Emerson Phase transgression that resulted in meltwater draining southward to the Gulf of Mexico after 9900 ¹⁴C yr BP. Copyright © 2005 John Wiley & Sons, Ltd.     

Devensian organic interstadial deposits and ice sheet extent in Buchan,Scotland

Pre-Late Devensian organic deposits in the Buchan area of northeast Scotland were investigated for their geomorphological and palaeoecological (pollen, plant macrofossils, coleoptera) properties. Close ecological agreement exists between fossil indicators and allows the inference that the environment in the vicinity of the deposits was a dwarf shrub tundra of the type met today in high latitude areas of Scandinavia and arctic Russia. The latest in a series of radiocarbon dates from the site produced determinations beyond the limits of the method, although the geomorphological and fossil evidence appears to point to an interstadial date within Oxygen Isotope Stages 5a or 5c. The site has special significance for arguments concerning the much-debated concept of ‘Moraineless Buchan’; indeed, evidence is presented which supports the concept of extensive ice sheet glaciation during the Late Devensian for this crucial geographical area. If Buchan is to be seen as a further casualty amongst other disputed ice-free enclaves, then a return to earlier models of extensive ice sheet glaciation in the Late Devensian of Scotland would seem to be necessary. © 1998 John Wiley & Sons, Ltd.