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Middle Jurassic Coptoclavidae (Insecta: Coleoptera: Dytiscoidea) from China: a Good Example of Mosaic Evolution
作者姓名:WANG Bo  Alexander G. PONOMARENKO and ZHANG Haichun
作者单位:WANG Bo,HANG Haichun(State Key Laboratory of Palaeobiology and Stratigraphy,Nanjing Institute of Geology and Palaeontology,Chinese Academy of Sciences,Nanjing 210008,China);A.G.PONOMARENKO(Paleontological Institute,Russian Academy of Sciences,ul.Profsoyuznaya 123,Moscow,117647,Russia) 
基金项目:This research has been financially supported by the National Natural Science Foundation of China,the Major Basic Research Program of China,Grant RFFI 07-04-92105-GFEN_a
摘    要:<正>Adults of the aquatic coptoclavid beetles(Coleoptera:Adephaga:Dytiscoidea),described from four Middle Jurassic fossil localities in Inner Mongolia and Liaoning in northeastern China,are attributed to Daohugounectes primitives Wang,Ponomarenko and Zhang,2009,which was previously proposed after study of larvae.The generic name Timarchopsis Brauer,Redtenbacher and Ganglbauer,1889 is proposed as a substitute for the preoccupied and junior homonym Necronectes Ponomarenko,1977,non Milne-Edwards,1881. Furthermore,the subfamily name Necronectinae Ponomarenko,1977 is substituted by the available name Timarchopsinae.Daohugounectes is placed into Timarchopsinae because its adults have long,slightly apically widened tibiae and small femoral plates.The adults of this genus differ from those of other Timarchopsinae in the following features:antennae short and widened in the middle part;basal segments of protarsi not cut apically; metaventrite with a triangular plate.The larvae look like somewhat primitive forms in the subfamily Timarchopsinae.In contrast to these primitive larvae,the adults with some advanced characters can be regarded as among the most advanced forms in the subfamily Timarchopsinae,and probably represent a transition between the Timarchopsinae and Charanoscaphinae.Such mosaic evolution within Daohugounectes indicates that the evolutionary process of aquatic beetles is far more complex than previously thought.

关 键 词:dytiscoid  beetles    Coptoclavidae    Coleoptera    mosaic  evolution    Jurassic    Daohugou    China
收稿时间:2009/12/20 0:00:00
修稿时间:2010/2/28 0:00:00

Middle Jurassic Coptoclavidae (Insecta:Coleoptera:Dytiscoidea) from China:a Good Example of Mosaic Evolution
Authors:WANG Bo  AGPONOMARENKO  ZHANG Haichun State Key Laboratory of Palaeobiology  Stratigraphy  Nanjing Institute of Geology  Palaeontology  Chinese Academy of Sciences  Nanjing  China Paleontological Institute  Russian Academy of Sciences  ulProfsoyuznaya  Moscow  Russia
Institution:WANG Bo~(1,*),A.G.PONOMARENKO~2 and ZHANG Haichun~1 1 State Key Laboratory of Palaeobiology and Stratigraphy,Nanjing Institute of Geology and Palaeontology,Chinese Academy of Sciences,Nanjing 210008,China 2 Paleontological Institute,Russian Academy of Sciences,ul.Profsoyuznaya 123,Moscow,117647,Russia
Abstract:Adults of the aquatic coptoclavid beetles(Coleoptera:Adephaga:Dytiscoidea),described from four Middle Jurassic fossil localities in Inner Mongolia and Liaoning in northeastern China,are attributed to Daohugounectes primitives Wang,Ponomarenko and Zhang,2009,which was previously proposed after study of larvae.The generic name Timarchopsis Brauer,Redtenbacher and Ganglbauer,1889 is proposed as a substitute for the preoccupied and junior homonym Necronectes Ponomarenko,1977,non Milne-Edwards,1881. Furthermore,...
Keywords:dytiscoid beetles  Coptoclavidae  Coleoptera  mosaic evolution  Jurassic  Daohugou  China  
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