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This review provides an up-to-date synthesis of the matrilineal phylogeography of a uniquely well-studied Holarctic mammal, the brown bear. We extend current knowledge by presenting a DNA sequence derived from one of the earliest known fossils of a polar bear (dated to 115 000 years before present), a species that shares a paraphyletic mitochondrial association with brown bears. A molecular clock analysis of 140 mitochondrial DNA sequences, including our new polar bear sequence, provides novel insights into the times of origin for different brown bear clades. We propose a number of regional biogeographic scenarios based on genetic data, divergence time estimates and paleontological records. The case of the brown bear provides an example for researchers working with less well-studied taxa: it shows clearly that phylogeographic models based on patterns of modern genetic variation alone can be substantially improved by including data on historical patterns of genetic diversity in the form of ancient DNA sequences derived from accurately dated samples and by using an approach to divergence-time estimation that suits the data under analysis. Using such approaches it has been possible to (i) establish that the processes shaping modern genetic diversity in brown bears acted recently, within the last three glacial cycles; (ii) distinguish among hypotheses concerning species’ responses to climatic oscillations in accordance with the lack of phylogeographic structure that existed in brown bears prior to the last glacial maximum (LGM); (iii) reassess theories linking monophyletic brown bear populations to particular LGM refuge areas; and (iv) identify vicariance events and track analogous patterns of migration by brown bears out of Eurasia to North America and Japan.  相似文献   
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The aurora and other phenomena in near Earth space are becoming a considerable part of the science curriculum in upper secondary school (high school) in Norway. Introducing scientific methods to the young students is an important objective of the education, but experimental experience is mainly restricted to simple laboratory exercises under controlled conditions; observations of uncontrollable natural phenomena are generally left to academic scientists and researchers. The Space Physics Group and The Science Education and Outreach Group at The Department of Physics and Technology, University of Bergen, are constructing a Space Science Suitcase with a set of simple versions of instruments for monitoring solar and geophysical activity in near Earth space. The instruments will be lent to physics classes in upper secondary schools.  相似文献   
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Crucian carp were exposed to three combinations of pH and inorganic aluminium concentrations for 25 days. Mortality, plasma chloride and haematocrit, and gill aluminium content were investigated. No mortality was observed in any of the three exposures. Crucian carp survived acidic Al-rich water (pH 5.17) with a high concentration of low-molecular weight inorganic Al species (242 µgl–1) for 25 days. These fish had lowered plasma chloride levels and high amounts of aluminium on their gills compared to fish exposed to acidic Al-poor water (pH 5.16 and 12 µgl–1 Ali) and control water (pH 6.29 and 4 µgl–1 Ali). Haematocrit was the same in all three exposures. Because aluminium was not acutely toxic to the anoxia tolerant crucian carp, the present results give support to the hypothesis that Al polymerization with subsequent hypoxia is of high importance for the mechanism of acute Al toxicity to fish.  相似文献   
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The conditions for energy flux, momentum flux and the resulting streaming velocity are analysed for standing waves formed in front of a fully reflecting wall. The exchange of energy between the outer wave motion and the near bed oscillatory boundary layer is considered, determining the horizontal energy flux inside and outside the boundary layer. The momentum balance, the mean shear stress and the resulting time averaged streaming velocities are determined. For a laminar bed boundary layer the analysis of the wave drift gives results similar to the original work of Longuet–Higgins from 1953. The work is extended to turbulent bed boundary layers by application of a numerical model. The similarities and differences between laminar and turbulent flow conditions are discussed, and quantitative results for the magnitude of the mean shear stress and drift velocity are presented. Full two-dimensional simulations of standing waves have also been made by application of a general purpose Navier–Stokes solver. The results agree well with those obtained by the boundary layer analysis. Wave reflection from a plane sloping wall is also investigated by using the same numerical model and by physical laboratory experiments. The phase shift of the reflected wave train is compared with theoretical and empirical models.  相似文献   
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Affluence drives the global displacement of land use   总被引:2,自引:0,他引:2  
Increasing affluence is often postulated as a main driver for the human footprint on biologically productive areas, identified among the main causes of biodiversity loss, but causal relationships are obscured by international trade. Here, we trace the use of land and ocean area through international supply chains to final consumption, modeling agricultural, food, and forestry products on a high level of resolution while also including the land requirements of manufactured goods and services. In 2004, high-income countries required more biologically productive land per capita than low-income countries, but this connection could only be identified when land used to produce internationally traded products was taken into account, because higher-income countries tend to displace a larger fraction of land use. The equivalent land and ocean area footprint of nations increased by a third for each doubling of income, with all variables analyzed on a per capita basis. This increase came largely from imports, which increased proportionally to income. Export depended mostly on the capacity of countries to produce useful biomass, the biocapacity. Our analysis clearly shows that countries with a high biocapacity per capita tend to spare more land for nature. Biocapacity per capita can be increased through more intensive production or by reducing population density. The net displacement of land use from high-income to low-income countries amounted to 6% of the global land demand, even though high-income countries had more land available per capita than low-income countries. In particular, Europe and Japan placed high pressure on ecosystems in lower-income countries.  相似文献   
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