干旱对青藏高原腹地高寒草地生态系统净CO2交换的影响

干旱对草地生态系统NEE有深刻影响。基于涡度相关技术提供的碳通量及小气候数据,研究了2009年当雄高寒草地生态系统的碳交换特征及其主控因子,同时分析了干旱的可能影响。5—7月初及9月发生的干旱导致草地GLAI、ALB和GPP较低,6月中旬到7月初碳吸收一度下降。干旱使6、7月份NEE日变化进程发生改变。同时,NEE和GPP的季节变化也受到干旱影响。由于干旱导致生态系统吸收能力降低,75]3日出现NEE日净碳排放最高值（0．9gCm-2d-1）。5-7月的NEE月总量均大于0,且逐月增加。该草地2009年的GPP和NEE分别为-158．1和52．4gCm。日均0〈01时,0成为影响白天NEE变化的主控因子。GLAI、r和目是3个对NEE季节变异影响最大的指标,且其影响程度依次降低。GPP季节变化的主控因子是GLAI、θ、PPT、VPD和瓦,生态系统水分状况（0、PPT或VPD）对GPP的影响大于T20。Rcco主要受控于t、GLAI、PAR和PPT,且其影响力依次降低。GLAI的季节变化可解释NEE和GPP变异的60．7％和76．1％。当雄高寒草地生态系统水分条件的年际变化可能是影响NEE年际变异的主要因子。     

干旱半干旱地区草地碳循环关键过程对降雨变化的响应

未来全球气候变化背景下,全球或局部地区的降雨量及降雨时间分布将发生一定的变化.草地生态系统在全球碳收支中作用显著,对气候变化的反馈起着重要的作用.但由于草地多处于干旱半干旱地区,受到水分条件的限制,对降雨变化响应敏感,其碳源汇功能表现出很大的不确定性.为了更好地预测未来全球气候变化背景下草地的碳源汇功能及其对气候变化的进一步反馈,有必要深入研究决定草地碳源汇功能的两大碳循环关键过程--净初级生产力和土壤呼吸对降雨变化的响应特征及机制.本文对国内外有关草地生产力和土壤呼吸如何对降雨量、降雨强度、降雨频率和间隔时间进行响应的相关研究成果进行了综述,在此基础上指出了目前研究存在的不足,并对未来相关的重点研究方向进行了探讨和展望.     

内蒙古草地生态系统碳源/汇时空格局及其与气候因子的关系

草地净生态系统生产力(NEP)能够表征草地生态系统的固碳能力,直接定性定量地描述草地生态系统的碳源/汇性质和大小.因此,研究区域尺度草地生态系统NEP具有重要的实践意义.基于卫星遥感资料,地面气象观测资料及实地采样数据,结合光能利用率模型估算了2001-2012年内蒙古草地生态系统净初级生产力(NPP).同时,应用土壤呼吸模型估算了逐月平均土壤呼吸量(Rs),进而估算内蒙古草地净生态系统生产力(NEP).研究揭示了2001-2012年内蒙古草地生态系统NPP,NEP年际变化规律,气候因子的年际变化规律,以及草地NPP,NEP与主要气候因子的关系.结果表明:2001年以来,内蒙古草地生态系统整体发挥碳汇效应,净碳汇总量达到0.55 Pg C,年均固碳率约为0.046 Pg C/a;研究区大部分草地NPP,NEP与降水均呈正相关关系,与温度相关性不显著,内蒙古草地生态系统仍有巨大的固碳潜力.     

温度和水分对科尔沁沙质草地土壤氮矿化的影响

土壤氮矿化对陆地生态系统初级生产力起决定性作用,但其影响因素较多,其中温度和水分最为重要。研究沙质草地土壤氮矿化对温度和水分的响应,对预测全球变化对沙质草地生态系统结构和功能的影响具有重要作用。因此,通过开顶式气室(OTC)模拟增温和人工调控田间持水量的方法对科尔沁沙质草地的土壤进行原位培养,分析温度和水分对土壤氮矿化作用的影响。结果表明:无论温度如何变化,科尔沁沙质草地土壤氮净矿化/硝化速率随着田间持水量的增加而明显提高。净硝化速率和净矿化速率在田间持水量为9.5%时最大,田间持水量达到时12.5%明显下降。增温使沙质草地土壤氮矿化显著变化,但增温的效应与田间持水量存在一定的关联。在相对适宜的田间持水量条件下(田间持水量为6.5%~12.5%),OTC增温可以使科尔沁沙质草地的土壤氮矿化/硝化速率显著提高;但是在田间持水量处于相对较低或者过高的状态下,该地区土壤的净氮净矿化/硝化速率对温度增加的响应不明显。     

温度和水分对科尔沁沙质草地土壤氮矿化的影响北大核心CSCD

土壤氮矿化对陆地生态系统初级生产力起决定性作用,但其影响因素较多,其中温度和水分最为重要。研究沙质草地土壤氮矿化对温度和水分的响应,对预测全球变化对沙质草地生态系统结构和功能的影响具有重要作用。因此,通过开顶式气室（OTC）模拟增温和人工调控田间持水量的方法对科尔沁沙质草地的土壤进行原位培养,分析温度和水分对土壤氮矿化作用的影响。结果表明：无论温度如何变化,科尔沁沙质草地土壤氮净矿化/硝化速率随着田间持水量的增加而明显提高。净硝化速率和净矿化速率在田间持水量为9.5%时最大,田间持水量达到时12.5%明显下降。增温使沙质草地土壤氮矿化显著变化,但增温的效应与田间持水量存在一定的关联。在相对适宜的田间持水量条件下（田间持水量为6.5%~12.5%）,OTC增温可以使科尔沁沙质草地的土壤氮矿化/硝化速率显著提高;但是在田间持水量处于相对较低或者过高的状态下,该地区土壤的净氮净矿化/硝化速率对温度增加的响应不明显。     

放牧对退化草地近地面辐射的影响

随着环境变化,草原地表状态也会改变,尤其是近地面能量收支过程变化更明显。为深入理解半干旱草原地表辐射能量平衡过程对放牧的响应特征,利用内蒙古正镶白旗典型退化草原2020年生长季（6—10月）的辐射通量观测资料,对比分析了禁牧、放牧草地太阳总辐射、地表反射辐射、大气长波辐射、地表长波辐射、净辐射以及地表反照率的日变化和生长季变化规律的差异。结果表明：在生长季,太阳总辐射随着时间推进逐月递减;禁牧区的地表反射辐射总体小于放牧区;各月大气长波辐射日变化幅度很小,处于130—370 W·m^-2;禁牧、放牧条件下地表长波辐射存在明显的季节变化规律,但是二者之间的差异甚微。内蒙古典型退化草原生长季近地面辐射通量有显著单峰型日变化特征。禁牧、放牧草地地表反照率都呈现“U”型日变化规律。生长季放牧草地的反照率明显高于禁牧草地。在禁牧区,辐射分量（地表反射辐射、大气长波辐射）和植被指数（归一化差异植被指数）对净辐射的影响是正向极显著的;而地表反照率和另一辐射分量（地表长波辐射）对净辐射有显著的负向作用。在放牧区,地表反射辐射和大气长波辐射对净辐射有极显著的正向作用;而地表反照率和地表长波辐射对净辐射的作用则是负向显著的。植被状况是影响内蒙古典型退化草原近地表辐射能量收支过程的首要因子。     

干旱对河西走廊植被净初级生产力的影响

干旱对生态系统碳循环具有重要影响,随着气候变暖,全球干旱事件频率不断上升,研究干旱对植被净初级生产力的影响具有重要意义。提高植被净初级生产力（NPP）的时间分辨率是认识干旱对其影响机制的重要途径。基于5天NDVI遥感数据,以河西走廊为研究区,利用CASA模型估算2010—2015年5天步长尺度的NPP,将5天降水为零定义为干旱基本单元,利用标准分数来探究干旱对植被NPP的影响。结果表明：基于高时间分辨率遥感数据能有效探测干旱对NPP的影响机制,研究期间河西走廊年干旱频次6—44次,干旱对该地NPP有负效应,对5天均值的影响为-0.23 g·m^-2,影响幅度为-1.93%,整体影响较小。干旱对林地NPP影响最大,下降3.70%,草地次之,荒漠最小。随着干旱持续时间的增加,NPP变化趋势呈现为先降低,然后降低趋势增加,最后趋于稳定的过程。干旱对夏季的NPP影响最大,冬季最小,秋季干旱对林地影响最大。     

京津冀地区生态系统服务权衡的多尺度特征与影响因素解析

对2000年和2015年京津冀地区四类生态系统服务(净初级生产力、粮食生产、水源涵养、土壤保持)进行估算,采用多层次空间统计法、相关分析法和玫瑰图法对生态系统服务权衡关系进行多尺度分析,利用多元回归模型定量揭示土地利用因素、社会因素、经济因素对生态系统服务权衡与协同关系的影响。结果表明:(1)净初级生产力(NPP)、水源涵养、土壤保持高值区集中分布于京津冀北部和西北部山区,低值区分布于东南部平原区,粮食生产呈东南高西北低的空间格局。2000—2015年仅粮食生产服务上升,净初级生产力、水源涵养、土壤保持服务均下降。(2)粮食生产与水源涵养、粮食生产与土壤保持表现出权衡关系,水源涵养与土壤保持表现为协同关系,这3对生态系统服务的权衡与协同关系均呈增强趋势。(3)耕地的粮食生产服务上升,土壤保持服务下降,各地类的净初级生产力均呈下降趋势。(4)生态系统服务权衡与协同关系的增强主要归因于土地利用因子,耕地减少、建设用地大幅增加、林地和草地增加是最直接的影响因素。     

祁连山干旱山地草地生物量对水分条件的响应

以祁连山排露沟流域干旱山地为研究对象,对海拔2 700~3 000 m典型草地群落的草本种类、高度和生物量等进行调查,并同步测定样地内的土壤水分,分析草地生物量随海拔高度的季节性变化特征以及草本生物量和土壤水分的关系。结果表明：（1）草地地上生物量平均值为135.36 g·m^-2,并随海拔升高呈先增加后降低的"单峰"变化模式,在海拔2 900 m时最高,为176.79±28.37 g·m^-2。地下生物量平均值为946.13 g·m^-2,并随海拔升高生物量呈递增趋势,在海拔3 000 m时最高,为1 301.19 ±68.24 g·m^-2。（2）草地地上、地下生物量在不同海拔高度间差异性显著（P<0.05）;该流域干旱山地草地根冠比在4.14~11.95之间变化。（3）在生长季5~9月份,干旱山地草地土壤含水量在9.23%~31.31%之间波动,平均值为14.94%。（4）草本地上、地下生物量与土壤平均含水量均呈显著正相关（P<0.05）,相关性系数分别为0.7784和0.7843。在不同海拔草地群落中,不同土层含水量对草地生物量的贡献不尽相同,但60 cm以上根系主要分布层内的水分对草地生物量具有重要的意义。     

黑河流域植被净初级生产力的遥感估算

利用光能利用率模型C-FIX,高时空分辨率的SPOT/VEGETATION遥感数据,全球格网化气象再分析资料以及黑河流域土地利用图,估算了1998—2002年黑河流域不同生态系统净初级生产力(NPP)的年总量和平均生产力,输出了黑河流域NPP年累积量空间分布格局图、NPP季节动态图,分析了近5a来黑河流域NPP时空变化特征,评价了不同土地利用类型下的生态系统生产力水平差异。其结果表明,黑河流域陆地生态系统NPP空间分布及其季相变化特征是流域自然环境、地貌、气候以及人类生产活动长期共同作用和影响的结果,其中水分条件是控制黑河流域陆地生态系统NPP空间格局的决定因子。1998—2002年黑河流域山区高覆盖度草地和下游荒漠区净初级生产力持续下降,反映出这些地区生态环境恶化严重。这些研究结果可以为黑河流域的水-生态-经济系统的合理设计与有效实施提供科学数据集。     

Implications of local-scale productivity on compensatory growth in a semi-arid shrubland

Grazing is a dominant determinant of aboveground net primary production (ANPP) and composition of plant communities. However, ANPP can be maintained following grazing due to compensatory growth, the level of which might depend on additional environmental factors, such as precipitation and edaphic conditions. Several studies have shown that along large-scale regional gradients, community-level compensatory ability is positively correlated with ANPP and soil resource availability. However, community-level responses to grazing are also expected to be affected by local-scale heterogeneity in ANPP, particularly under low primary productivity typical to arid environments. Here, we studied the effect of local-scale variations in ANPP on the compensatory growth of an annual community in a semi-arid region. For two consecutive years, ANPP was evaluated following shoot damage in sites with different primary productivity. The results demonstrated that annual ANPP varied significantly among sites and among plots within sites; however, compensatory ability was negatively correlated with annual ANPP, with overcompensation in the least productive patches and under-compensation in the most productive patches. This pattern contradicts the positive correlation between ANPP and compensatory ability commonly found along large-scale productivity ecoclines, suggesting that the effects of ANPP on compensatory ability might be scale-dependent.     

Soil as a capacitor: Considering soil water content improves temporal models of productivity

Above-ground net primary production (ANPP) in arid and semiarid ecosystems is mainly explained by precipitation (ppt). However, when this relationship is evaluated taking into consideration data from different years in the same site (i.e. temporal models of productivity) the relation is weak, and sometimes it does not exist. In spite of this, the inclusion of previous year's ppt and/or ANPP frequently improves temporal models. In this study we analyze if considering NDVI and mean annual soil water content or transpiration (instead of different combinations of current-year ppt and previous year's ppt or ANPP) improves temporal models of productivity in the Southern Monte (Argentina). Current-year ppt only explained 39.7% of variation in ANPP, while mean soil water content explained 85.3%. The remaining models, which include current-year ppt together with previous-year's ppt or previous-year's ANPP, improve the first model; but the explanatory power of the model based only on mean soil water content is never reached. Our results also show that water losses exceed annual ppt in dry years, whereas the opposite occurs during years with above-average ppt. This carryover effect of soil water indicates that soil acts as a capacitor, accumulating water during wettest years and releasing it during following years.     

Spatiotemporal variations of water use efficiency and its driving factors in Inner Mongolia from 2001 to 2020

Water use efficiency (WUE) is an important variable to explore coupled relationships in carbon and water cycles. In this study, we first compared the spatial variations of annual gross primary productivity (GPP) and evapotranspiration (ET) using four GPP and ET products. Second, we selected the products closest to the flux towers data to estimate WUE. Finally, we quantitatively analyzed the impact of climate change and soil water content on WUE. The results showed that: (1) Four GPP and ET products provided good performance, with GOSIF-GPP and FLDAS-ET exhibiting a higher correlation and the smallest errors with the flux tower data. (2) The spatial pattern of WUE is consistent with that of GPP and ET, gradually decreasing from the northeast to the southwest. Higher WUE values appeared in the northeast forest ecosystem, and lower WUE values occurred in the western Gobi Desert, with a value of 0.28 gC m^?2 mm^?1. The GPP and ET products showed an increasing trend, while WUE showed a decreasing trend (55.15%) from 2001 to 2020. (3) The spatial relationship between WUE and driving factors reveal the variations in WUE of Inner Mongolia are mainly affected by soil moisture between 0 and 10 cm (SM0-10cm), vapor pressure deficit (VPD), and precipitation, respectively. (4) In arid regions, VPD and precipitation exhibit a major influence on WUE. An increase in VPD and precipitation has a negative and positive effect on WUE, with threshold values of approximately 0.36 kPa and 426 mm, respectively. (5) In humid regions, SM0-10cm, VPD, SM10-40cm, and SM40-100cm exert a significant impact on WUE, especially SM0-10cm, and weakens with increasing soil depths, these differences may be related to physiological structure and living characteristics of vegetation types in different climate regimes. Our results emphasize the importance of VPD and soil moisture in regional variability in WUE.     

Responses of Grassland Net Primary Productivity to Environmental Variables in Northern China

Various environmental factors affect net primary productivity (NPP) of grassland ecosystem. Extensive reports on the effects of environmental variables on NPP can be found in literature. However, the agreement on the relative importance of various factors in shaping the spatial pattern of grassland NPP has not yet been reached. Here a grassland in situ NPP database comprising 602 samples in northern China for 1980-1999 was developed based on a literature review of published biomass and forage yield field measurements. Correlation analyses and dominance analysis were used to quantify the separate and combined effects of environmental variables (climate, topography and soil) on spatial variation in NPP separately. Grassland NPP ranged from 4.76 g C m^-2a^-1 to 975.94 g C m^-2a^-1, showing significant variations in space. NPP increased with annual precipitation and declined with annual mean temperature significantly. Specifically, precipitation had the greatest impact on deserts, followed by steppes and meadows. Grassland NPP decreased with increasing altitude because of water limitation, and positively correlated with slope, but weakly correlated with aspect. Soil quality showed positive effects on NPP. Annual precipitation was the dominant factor affecting the spatial variability of net primary productivity, followed by elevation.     

Nitrogen limitation in arid-subhumid ecosystems: A meta-analysis of fertilization studies

Evidence supporting water limitation in arid-semiarid ecosystems includes strong correlations between aboveground net primary production (ANPP) and annual precipitation as well as results from experimental water additions. Similarly, there is evidence of N limitation on ANPP in low precipitation ecosystems, but is this a widespread phenomenon? Are all arid-semiarid ecosystems equally limited by nitrogen? Is the response of N fertilization modulated by water availability?We conducted a meta-analysis of ANPP responses to N fertilization across arid to subhumid ecosystems to quantify N limitation, using the effect-size index R which is the ratio of ANPP in fertilized to control plots. Nitrogen addition increased ANPP across all studies by an average of 50%, and nitrogen effects increased significantly (P = 0.03) along the 50-650 mm yr⁻¹ precipitation gradient. The response ratio decreased with mean annual temperature in arid and semiarid ecosystems but was insensitive in subhumid systems. Sown pastures showed significant (P = 0.007) higher responses than natural ecosystems. Neither plant-life form nor chemical form of the applied fertilizer showed significant effects on the primary production response to N addition. Our results showed that nitrogen limitation is a widespread phenomenon in low-precipitation ecosystems and that its importance increases with annual precipitation from arid to subhumid regions. Both water and N availability limit primary production, probably at different times during the year; with frequency of N limitation increasing and frequency of water limitation decreasing as annual precipitation increases. Expected increase N deposition, which could be significant even in arid ecosystems, would increase aboveground net primary production in water-limited ecosystems that account for 40% of the terrestrial surface.     

退耕还林还草工程对黄土高原植被总初级生产力的影响

采用Landsat解译的2000年和2015年土地利用/覆盖数据和VPM模型（Vegetation Photosynthesis Model）模拟的2000~2016年总初级生产力（Gross Primary Productivity, GPP）数据,识别出了近16 a黄土高原退耕还林还草的空间范围,并估算了GPP的年际变化趋势。在此基础上,对比分析了退耕区和未退耕区GPP年际变化的差异,从而揭示退耕还林还草工程对GPP年际变化的影响。结果显示,2000~2015年,黄土高原退耕还林还草面积约3.5万km ²,占2000年耕地面积的16.8%。期间,GPP呈增加趋势,GPP显著上升区域占全区面积的67.3%,平均增速24.1 g/(m ²?a)（以C计,下同）。虽然退耕区多年平均GPP低于未退耕区,但退耕区GPP年际增速和相对变化率明显高于未退耕区,分别提高了5.9 g/(m ²?a)和1.5%。     

Declining Precipitation Enhances the Effect of Warming on Phenological Variation in a Semiarid Tibetan Meadow Steppe

Vegetation phenology is a sensitive indicator of global warming, especially on the Tibetan Plateau. However, whether climate warming has enhanced the advance of grassland phenology since 2000 remains debated and little is known about the warming effect on semiarid grassland phenology and interactions with early growing season precipitation. In this study, we extracted phenological changes from average NDVI in the growing season (GNDVI) to analyze the relationship between changes in NDVI, phenology and climate in the Northern Tibetan Damxung grassland from 2000 to 2014. The GNDVI of the grassland declined. Interannual variation of GNDVI was mainly affected by mean temperature from late May to July and precipitation from April to August. The length of the growing season was significantly shortened due to a delay in the beginning of the growing season and no advancement of the end of the growing season, largely caused by climate warming and enhanced by decreasing precipitation in spring. Water availability was the major determinant of grass growth in the study area. Warming increased demand for water when the growth limitation of temperature to grass was exceeded in the growing season. Decreased precipitation likely further exacerbated the effect of warming on vegetation phenology in recent decades due to increasing evapotranspiration and water limitations. The comprehensive effects of global warming and decreasing precipitation may delay the phenological responses of semiarid alpine grasslands.     

草原化荒漠植被草本层片植物对人工施加氮素的响应

水分是干旱区生态过程的第一非生物限制因子,土壤氮由于直接影响着生物生产力和土壤生态过程而被认为是仅次于水分的限制因子。利用人工施加氮肥的控制试验研究了荒漠植被草本层片植物在多度、物种丰富度、高生长以及地上部分生物量等群落学特征对不同施肥处理的响应。结果表明,在施肥量分别为12.5、25、50 g·m-2时,样方植物多度和物种丰富度在施肥当年和第二年较对照均有不同程度的降低,且施肥量越高,降低越明显,这一结果支持来自典型草地和森林植被生物多样性对氮素增加或氮降沉增加响应的结论,即氮素的增加会使生物多样性减少。地上部分生物量对施氮肥的响应在施肥当年(年降水量为多年平均值的1.4倍)和第二年(年降水量为多年平均值的0.7倍)表现出相反的规律,即施肥当年随着施肥量的增加生物量显著增加,而在第二年则显著减小,说明水肥耦合同样有利于荒漠生态系统生物生产力的提高。氮素对植物高生长的影响不明显,各施肥梯度与对照相比均无显著性差异(P>0.05),这可能主要取决于荒漠草本植物本身固有的生物学特性。     

宁夏农牧交错区（盐池）草地生产力对气候变化的响应

用宁夏农牧交错区(盐池县)1954—2004年的气候资料,分析了该地区51 a来气温、降水的变化趋势及其草地气候生产力的变化。结果表明：从1954年以来的51 a内,盐池气温呈明显上升趋势;春、夏季降水量和年降水量略呈增加趋势,秋、冬季降水量略呈减少的态势,但趋势不明显;草地气候生产力呈增加趋势。草地气候生产力与年降水量关系密切,水分是制约草地气候生产力的关键因子。未来“暖湿型”气候对盐池草地的干物质生产最有利,平均增产幅度为10% ,而“冷干型”气候对草地的干物质生产最不利,平均减产幅度为10%。若气温升高1～2 ℃,降水量增加10%～20% ,则盐池草地的气候生产力将增加10%～20%。