Phylogenetic systematics of chelonioid sea turtles

Abstract Chelonioid sea turtles (Order Testudinata; Superfamily Chelonioidea) first appear in the early Cretaceous (Aptian-Albian). Their long fossil record is excellent compared to most marine tetrapods. Nonetheless, there has been no inclusive attempt to provide an analysis of cladistic relationships among this group. In this paper, the following chelonioid classification is proposed, based on 76 characters among 20 fossil and six living well- represented genera:

• 1 Family Cheloniidae (Coniacian to Recent); Toxochelys, Ctenochelys, Osteopygis, Erquelinnesia, Allopleuron, Argillochelys, Puppigerus, Eochelone, Syllomus, Natator, Eretmochelys, Chelonia, Caretta, Lepidochelys
• 2 Family Protostegidae (Aptian to Maastrichtian?); ‘Santana new protostegid’,Rhinochelys, Notochelone, Desmatochelys, Chelosphargis, Protostega, Archelon
• 3 Family Dermochelyidae (Santonian? to Recent); Corsochelys, ‘HMG new dermo-chelyid’, Eosphargis, Psephophorus, Dermochelys.

     

Calcareous nannofossil biostratigraphic study of forearc basin sediments: Lower to Upper Cretaceous Budden Canyon Formation (Great Valley Group), northern California,USA

The results of a calcareous nannofossil biostratigraphic investigation of the North Fork Cottonwood Creek section of the Budden Canyon Formation (BCF; Hauterivian–Turonian) in northern California are summarized using the Boreal – cosmopolitan Boreal Nannofossil Biostratigraphy (BC) – Upper Cretaceous Nannofossil Biostratigraphy (UC) nannofossil zonal schemes of Bown et al. and Burnett et al. Sixteen intervals, ranging from the BC15 to UC8 zones, were established in the section. Combined biostratigraphic and magnetostratigraphic studies suggest a Hauterivian to mid‐Turonian age for the studied sequence. The Hauterivian–Barremian, Barremian–Aptian, Aptian–Albian, Albian–Cenomanian, and Cenomanian–Turonian stage boundaries were delineated near the top of the Ogo Member, below the Huling Sandstone Member, within the upper Chickabally Member, in the upper portion of the Bald Hills Member and within the Gas Point Member, respectively. Unconformities probably exist at the base of the Huling Sandstone Member and the upper part of the upper Chickabally Member. The nannofossil assemblage in the North Fork Cottonwood Creek suggests that the study area was under the influence of cold‐water conditions during the Barremian to Lower Aptian interval, shifting to tropical/warm‐water conditions during the Albian to Turonian interval as a result of the mid‐Cretaceous global warming. Although oceanic anoxic events have not yet been reported in the BCF, preliminary total organic carbon, along with nannofossil data, suggest the presence of the global Cenomanian–Turonian boundary oceanic anoxic event 2.     

Triploporella ? edgelli n. sp., a new Dasycladalean alga from the Lower Cretaceous of Lebanon

The Dasycladalean assemblage of the Jezzinian strata (uppermost Barremian–lowermost Aptian) of Lebanon consists of two organo‐genera and eight genera with eleven species, including eight Triploporellaceae. Although Triploporella marsicana (Praturlon, 1964) was reported by Saint‐Marc from the same interval, it is not found in our material. However, a lookalike, which is herein described as Triploporella ? edgelli n. sp., is identified. This new species has cyst‐containers within the primary segments of its laterals but, because it lacks calcified secondary segments, it is left in open nomenclature.     

Effect of human‐controlled hydrological regime on the source,transport, and flux of particulate organic carbon from the lower Huanghe (Yellow River)

Evaluating the role of fluvial transfer of terrestrial organic carbon (OC) and subsequent burial in the global carbon cycle requires the sources and fluxes of fluvial OC to be assessed, which remains poorly constrained in the Huanghe (Yellow River). Here, we report the elemental, stable isotopic, and radiocarbon activity of particulate organic carbon (POC) sampled at the outlet of Huanghe in 2012–2013. We show that the Huanghe riverine POC can be explained by binary mixing of fossil (POC_fossil) and non‐fossil (POC_non‐fossil) components, the former may reach ~40% of the total POC. The Huanghe POC_non‐fossil is mostly sourced from C3 plants, with a mean residence time of c. 2200 years. The current human‐controlled hydrological regime strongly influenced the POC sources, transport modes, and fluxes. In 2012–2013, the Huanghe delivered 0.73 Tg (1 Tg = 10¹² g) of POC to the sea, and about 28% of the annual POC flux occurred within a short human induced flood event. Globally, the Huanghe should be one of the largest rivers in the transfer and re‐burial of fossil OC. However, the fate of Huanghe fossil OC is still unconstrained and needs to be further investigated. Copyright © 2015 John Wiley & Sons, Ltd.     

Fourth‐ to third‐order cycles in the Hakobuchi Formation: Shallow‐marine Campanian final deposition of the Yezo Group,Nakagawa area,northern Hokkaido,Japan

The Yezo Group has a wide longitudinal distribution across Hokkaido, northern Japan. It represents a Cretaceous (Early Aptian–Late Maastrichtian) and Late Paleocene forearc basin‐fill along the eastern margin of the paleo‐Asian continent. In the Nakagawa area of northern Hokkaido, the uppermost part of the Yezo Group consists of the Hakobuchi Formation. Along the western margin of the Yezo basin, 24 sedimentary facies (F) represent 6 facies associations (FA), suggesting prevailing storm‐dominated inner shelf to shoreface environments, subordinately associated with shoreface sand ridges, outer shelf, estuary and fluvial environments. The stacking patterns, thickness and facies trends of these associations allow the discrimination of six depositional sequences (DS). Inoceramids Sphenoceramus schmidti and Inoceramus balticus, and the ammonite Metaplacenticeras subtilistriatum, provide late Early to Late Campanian age constraints to this approximately 370‐m thick final stage of deposition and uplift of the Yezo forearc basin. Six shallow‐marine to subordinately non‐marine sandstone‐dominated depositional sequences include four 10 to 110‐m thick upward‐coarsening regressive successions (FS1), occasionally associated with thin, less than 10‐m thick, upward‐fining transgressive successions (FS2). The lower DS1–3, middle DS4–5 and upper DS6 represent three depositional sequential sets (DSS1–3). These eastward prograding and westward retrograding recurring shallow‐marine depositional systems may reflect third‐ and fourth‐order relative sealevel changes, in terms of sequence stratigraphy.     

Detrital zircon geochronology of the Cretaceous Sindong Group,Southeast Korea: Implications for depositional age and Early Cretaceous igneous activity

The Sindong Group forms the lowermost basin‐fill of the Gyeongsang Basin, the largest Cretaceous nonmarine basin located in southeastern Korea, and comprises the Nakdong, Hasandong, and Jinju Formations with decreasing age. The depositional age of the Sindong Group has not yet been determined well and the reported age ranges from the Valanginian to Albian. Detrital zircons from the Sindong Group have been subjected to U–Pb dating using laser ablation inductively coupled plasma mass spectrometry. The Sindong Group contains noticeable amounts of detrital magmatic zircons of Cretaceous age (138–106 Ma), indicative of continuous magmatic activity prior to and during deposition of the Sindong Group. The youngest detrital zircon age of three formations becomes progressively younger stratigraphically: 118 Ma for the Nakdong Formation, 109 Ma for the Hasandong Formation, and 106 Ma for the Jinju Formation. Accordingly, the depositional age of the Sindong Group ranges from the late Aptian to late Albian, which is much younger than previously thought. Lower Cretaceous magmatic activity, which supplied detrital zircons to the Sindong Group, changed its location spatially through time; it occurred in the middle and northern source areas during the early stage, and then switched to the middle to southern source areas during the middle to late stages. This study reports first the Lower Cretaceous magmatic activity from the East Asian continental margin, which results in a narrower magmatic gap (ca 20 m.y.) than previously known.     

Depositional environments of well‐sorted detrital limestone from the Minatogawa Formation in the southern part of Okinawa Island,the Ryukyu Archipelago,Japan

Well‐sorted detrital limestone is one of the typical lithofacies of the latest interval of the Pleistocene Ryukyu Group, which is exposed in the Ryukyu Archipelago in southwestern Japan. The depositional environments of the limestone are interpreted to be extremely shallow and to include back‐reef lagoons or moats and subaerial sand dunes. However, detailed micropaleontological analyses have not been performed on this limestone. In this study, the interpretation of the depositional environments and paleo‐water depths was improved by quantitative examination of foraminiferal assemblages for the well‐sorted detrital limestone of the Minatogawa Formation in the southern part of Okinawa Island. Thin sections of limestone collected from the Minatogawa (Horikawa) quarry were subjected to sedimentological and foraminiferal analyses. Comparison with modern foraminiferal distribution within the Ryukyu Archipelago indicates that back‐reef and fore‐reef dwelling foraminifers characterize the fossil assemblages from the well‐sorted detrital limestone (bioclastic grainstone). Three ratios of indicator foraminiferal taxa (ratios of back‐reef to fore‐reef taxa, planktonic foraminifers to Amphistegina lobifera and Amphistegina lessonii, and Calcarina gaudichaudii to other Calcarina species), as well as multivariate analyses suggest that the well‐sorted detrital limestone was deposited in fore‐reef setting shallower than 40 m in water depth. A comparable depth range was reconstructed from the coral assemblage in the associated coral limestone, suggesting that the Minatogawa Formation was deposited in a gently inclined ramp setting with patch reefs and/or fringing reefs. Stratigraphic changes in paleo‐water depth, together with evidence of several unconformities associated with paleosol layers suggest that there were repeated transgressions and regressions, with an amplitude up to several tens of meters, when the Minatogawa Formation was deposited.     

Depositional ages and characteristics of Middle–Upper Jurassic and Lower Cretaceous lacustrine deposits in southeastern Mongolia

Lower Cretaceous lacustrine oil shales are widely distributed in southeastern Mongolia. Due to the high organic carbon content of oil shale, many geochemical studies and petroleum exploration have been conducted. Although most of the oil shales are considered to be Early Cretaceous in age, a recent study reveals that some were deposited in the Middle Jurassic. The present study aims at establishing depositional ages and characteristics of the Jurassic and Cretaceous lacustrine deposits in Mongolia. The Lower Cretaceous Shinekhudag Formation is about 250 m thick and composed of alternating beds of shale and dolomite. The Middle Jurassic Eedemt Formation is about 150 m thick and composed of alternating beds of shale, dolomitic marl, and siltstone. The alternations of shale and dolomite in both formations were formed by lake level changes, reflecting precipitation changes. Shales were deposited in the center of a deep lake during highstand, while dolomites were formed by primary precipitation during lowstand. Based on the radiometric age dating, the Shinekhudag Formation was deposited between 123.8 ±2.0 Ma and 118.5 ±0.9 Ma of the early Aptian. The Eedemt Formation was deposited at around 165–158 Ma of Callovian–Oxfordian. The calculated sedimentation rate of the Shinekhudag Formation is between 4.7 ±2.6 cm/ky and 10.0 ±7.6 cm/ky. Shales in the Shinekhudag Formation show micrometer‐scale lamination, consisting of algal organic matter and detrital clay mineral couplets. Given the average thickness of micro‐laminae and calculated sedimentation rate, the micro‐lamination is most likely of varve origin. Both Middle–Upper Jurassic and Lower Cretaceous lacustrine oil shales were deposited in intracontinental basins in the paleo‐Asian continent. Tectonic processes and basin evolution basically controlled the deposition of these oil shales. In addition, enhanced precipitation under humid climate during the early Aptian and the Callovian–Oxfordian was another key factor inducing the widespread oil shale deposition in Mongolia.     

Preferred orientation of the trace fossil Entradichnus meniscus in eolian dune strata (Djadokhta Formation) at Tugrikiin Shiree,southern Mongolia and its paleoecological implications

Eolian sand dune deposits of the Upper Cretaceous Djadokhta Formation at Tugrikiin Shiree, southern Mongolia, yield not only dinosaur skeletal remains but also numerous trace fossils produced by invertebrates. This paper describes the trace fossil Entradichnus meniscus, a long unlined and unbranched trail that is filled with meniscate laminae and occurs characteristically in positive epirelief. The trail is straight to gently meandering, parallel to the foreset laminae of the eolian dunes, and exhibits a significant preferred orientation parallel to the depositional dip of the cross‐stratification laminae. In addition, almost all the crescentic internal laminae of the trail show concave down‐dips. These features indicate that the trails were produced beneath the slipface of eolian dunes by the downward burrowing of the trace‐makers. This occurrence mode of E. meniscus of the Mongolian Cretaceous is very similar to that described from the Jurassic eolian dune deposits in North America. Hence, the downward burrowing of the E. meniscus animal might be a common feature in arid eolian dune deposits at least during the Jurassic and Cretaceous, and possibly reflecting a behavioral response to the morphology of large sand dunes under an arid climate.     

New discovery of Palaeontinid fossils from the Middle Jurassic in Daohugou, Inner Mongolia (Homoptera, Palaeontinidae)

Three new species of fossil Palaeontinidae are described from Daohugou Village, Inner Mongolia, China: Daohugoucossus shii sp. nov., D. parallelivenius sp. nov., D. lii sp. nov. The diagnosis of Dao- hugoucossus Wang, Zhang and Fang, 2006 is revised. These materials are the first complete fossil palaeontinids in the Middle Jurassic of the world. Based on observation of these new specimens, Sc of this new genus has the following combined characters: joined with R at base, diverged near M separa- tion, coalesced with R before the nodal line, extended beyond the nodal line and fused with R1 before distal end. Due to the Sc characteristics mentioned above, the genus Liaocossus Ren, Yin and Dou, 1998 can be well distinguished from “Ilerdocossus-complex”.     

Phylogenetic relationships of the Chinese fossil species of the genusEquus (Perissodactyla, Equidae)

The Chinese fossils ofEquus began to appear from the beginning of the Quaternary at 2.5 Ma B.P., and the extant species ofEquus still live in China under natural conditions at the present, which is unique in the world. 12 valid fossil species ofEquus have been discovered in China, including the stenonid, caballoid and hemione representatives ofEquus. The origin and evolution of the Chinese fossil species and other relevant species ofEquus are discussed in detail. The pmposed phylogenetic relationships and temporal distribution sequences including all the Chinese fossil species ofEquus are established. Some mistakes in the earlier researches about the Chinese fossils ofEquus are corrected. Project supported by the French-Chinese Cooperation (Grant No. 203720C).     

Zircon U–Pb ages from tuff beds of the upper Mesozoic Tetori Group in the Shokawa district,Gifu Prefecture,central Japan†

Abstract The upper Mesozoic Tetori Group contains numerous fossils of plants and marine and non‐marine animals. The group has the potential to provide key information to improve our understanding of the Middle Jurassic to Early Cretaceous biota of East Asia. However, the depositional age of the Tetori Group remains uncertain, and without good age constraints, accurate correlation with other areas is very difficult. As a first step in obtaining reliable ages for the formations within the Tetori Group, we used laser ablation‐inductively coupled plasma–mass spectrometry to measure the U–Pb ages of zircons collected from tuff beds in the Shokawa district, Takayama City, Gifu Prefecture, central Japan. The youngest reliable U–Pb ages from the tuff beds of the Ushimaru, Mitarai and Okurodani Formations are 130.2 ± 1.7, 129.8 ± 1.0 and 117.5 ± 0.7 Ma, respectively (errors represent 2 SE). These results indicate that the entire Tetori Group in the Shokawa district, which was previously believed to be correlated to the Upper Jurassic to Lower Cretaceous, is in fact correlated to the Lower Cretaceous. The maximum ages of the Ushimaru, Mitarai and Okurodani Formations are late Hauterivian to Barremian, late Hauterivian to Barremian and Barremian to Aptian, respectively.     

The Cretaceous of the East Asian continental margin: Stratigraphy,paleogeography, and paleoclimate

Lithostratigraphic correlation of a 6–10‐km‐thick Aptian–Maastrichtian terrigenous sequence of the East Asian continental margin and Sakhalin and Hokkaido Islands has revealed the existence of a single marine basin. This basin was populated by mixed Tethyan–Boreal fauna and sloped eastward until the Middle Cenomanian. Intense volcanic and tectonic processes caused the uplift of the continental margin in the mid‐Albian to Cenomanian and eastward migration of the shoreline. Paleobotanical studies have discovered a number of climatic changes. Relatively warm conditions existed in the Aptian, changing to cooler conditions in the Early Albian. The maximum warming occurred from the Late Albian to Cenomanian when large‐leaved flowering plants dominated the population. In the Late Cretaceous, the East Asian volcanic belt created a mountain edifice up to 3000 m high, which controlled longitudinal climatic and floral zonation. This control was more efficient than the latitudinal control. A wide development of flowering Platanaceae flora in the Turonian points to a relative cooling. The floral assemblage shows a temperate climate from the Early Coniacian onwards, with an optimum in the Campanian that is consistent with global transgression. As a result of the warm climate, the early Maastrichtian is characterized by highly diverse biota. Furthermore, in the middle Maastrichtian floral and faunal diversity decreased and the seasonal prevalence increased. Cooling occurred in the latest Maastrichtian. Five periods of coal accumulation are recognized throughout the Late Cenomanian to Late Maastrichtian. Shelf, deltaic, and slope turbidite facies of the continental margin contain organic carbon ranging from 0.3% to 2.2%, which together with direct evidence for oil and gas, are believed to be prolific for hydrocarbon exploration.     

Ultrasonic velocities of North Sea chalk samples: influence of porosity, fluid content and texture

We have studied 56 unfractured chalk samples of the Upper Cretaceous Tor Formation of the Dan, South Arne and Gorm Fields, Danish North Sea. The samples have porosities of between 14% and 45% and calcite content of over 95%. The ultrasonic compressional‐ and shear‐wave velocities (V_P and V_S) for dry and water‐saturated samples were measured at up to 75 bar confining hydrostatic pressure corresponding to effective stress in the reservoir. The porosity is the main control of the ultrasonic velocities and therefore of the elastic moduli. The elastic moduli are slightly higher for samples from the South Arne Field than from the Dan Field for identical porosities. This difference may be due to textural differences between the chalk at the two locations because we observe that large grains (i.e. filled microfossils and fossil fragments) that occur more frequently in samples from the Dan Field have a porosity‐reducing effect and that samples rich in large grains have a relatively low porosity for a given P‐wave modulus. The clay content in the samples is low and is mainly represented by either kaolinite or smectite; samples with smectite have a lower P‐wave modulus than samples with kaolinite at equal porosity. We find that ultrasonic V_P and V_S of dry chalk samples can be satisfactorily estimated with Gassmann's relationships from data for water‐saturated samples. A pronounced difference between the V_P/V_S ratios for dry and water‐saturated chalk samples indicates promising results for seismic amplitude‐versus‐offset analyses.     

First‐order exchange coefficient coupling for simulating surface water–groundwater interactions: parameter sensitivity and consistency with a physics‐based approach

Distributed hydrologic models capable of simulating fully‐coupled surface water and groundwater flow are increasingly used to examine problems in the hydrologic sciences. Several techniques are currently available to couple the surface and subsurface; the two most frequently employed approaches are first‐order exchange coefficients (a.k.a., the surface conductance method) and enforced continuity of pressure and flux at the surface‐subsurface boundary condition. The effort reported here examines the parameter sensitivity of simulated hydrologic response for the first‐order exchange coefficients at a well‐characterized field site using the fully coupled Integrated Hydrology Model (InHM). This investigation demonstrates that the first‐order exchange coefficients can be selected such that the simulated hydrologic response is insensitive to the parameter choice, while simulation time is considerably reduced. Alternatively, the ability to choose a first‐order exchange coefficient that intentionally decouples the surface and subsurface facilitates concept‐development simulations to examine real‐world situations where the surface‐subsurface exchange is impaired. While the parameters comprising the first‐order exchange coefficient cannot be directly estimated or measured, the insensitivity of the simulated flow system to these parameters (when chosen appropriately) combined with the ability to mimic actual physical processes suggests that the first‐order exchange coefficient approach can be consistent with a physics‐based framework. Copyright © 2009 John Wiley & Sons, Ltd.     

Sustainable Chemical Technologies in Production of Clean Fuels from Fossil Fuels

Some aspects of the present and the possible future role of sustainable chemical technologies in the production of clean liquid and gaseous fuels from fossil fuels are discussed. The state‐of‐the‐art and the vision of possible sources and alternative routes which may lead to clean fuels from fossil fuels due to the progress in crude oil, natural gas and coal processing are briefly presented. The possible future role of the Fischer‐Tropsch synthesis, methanol synthesis, dimethylether synthesis, and a group of methanol transformation processes is also discussed.     

Middle and early cretaceous magnetic stratigraphy from the Cismon section,northern Italy

The pelagic limestones exposed in the valley of the Cismon river (near Feltre) appear to represent continuous deposition from Valanginian to Campanian, apart from a short hiatus in the Early Albian. Detrital magnetite is the carrier of remanence in these predominantly white-grey limestones, and a well-defined magnetic stratigraphy has been obtained. The Cretaceous quiet zone at Cismon is totally normal in polarity and stretches from Early Aptian to Early Campanian. Below the Lower Aptian, the Early Cretaceous mixed polarity interval is tentatively correlated with the sequence of geomagnetic reversals derived from the oceanic magnetic anomalies.     

Control strategy for variable damping element considering near‐future excitation influence

A control strategy is proposed for variable damping elements (VDEs) used together with auxiliary stiffness elements (ASEs) that compose a time‐varying non‐linear Maxwell (NMW) element, considering near‐future excitation influence. The strategy first composes a state equation for the structural dynamics and the mechanical balance in the NMW elements. Next, it establishes a cost function for estimating future responses by the weighted quadratic norms of the state vector, the controlled force and the VDEs' damping coefficients. Then, the Euler equations for the optimum values are introduced, and also approximated by the first‐order terms under the autoregressive (AR) model of excitation information. Thus, at each moment t_k, the strategy conducts the following steps: (1) identify the obtained seismic excitation information to an AR model, and convert it to a state equation; and (2) determine VDEs' damping coefficients under the initial conditions at t_k and the final state at t_k+L, using the first‐order approximation of the Euler equations. The control effects are examined by numerical experiments. Copyright © 2000 John Wiley & Sons, Ltd.     

Intrinsic Qp seismic attenuation from the rise time of microearthquakes: a local scale application at Rio‐Antirrio,Western Greece

We determine the attenuation structure of a three‐dimensional medium based on first pulse‐width measurements from microearthquake data. Ninety‐five microearthquakes from a seventy stations local network were considered in this study. Measurements of the first half cycle of the wave, the so‐called rise time τ were carried out on high quality velocity seismograms and inverted to estimate the P‐waves intrinsic quality factor Q_p. The results of this investigation indicate that first pulse width data from a local microearthquake network permit retrieval with sufficient accuracy of the heterogeneous Q_p structure. The inferred attenuation variability corresponds to the known geological formations in the region.     

Adsorption Kinetic Modeling of Safranin onto Rice Husk Biomatrix Using Pseudo‐first‐ and Pseudo‐second‐order Kinetic Models: Comparison of Linear and Non‐linear Methods

Batch kinetic studies were carried out for the removal of safranin from aqueous solution using a biomatrix prepared from rice husk. The adsorption kinetic data were modeled using the pseudo‐first‐order and pseudo‐second‐order kinetic equations. The linear and non‐linear forms of these two widely used kinetic models were compared in this study. In order to determine the best‐fitting equation, the coefficient of determination (r²), the sum of the squares of the errors (SSE), sum of the absolute errors (SAE), average relative error (ARE), hybrid fractional error function (HYBRID), Marquardt's percent standard deviation (MPSD), and the Chi‐squared test (χ²) were used as error analysis methods. Results showed that the non‐linear forms of pseudo‐first‐order and pseudo‐second‐order models were more suitable than the linear forms for fitting the experimental data. Non‐linear method is thus more appropriate for estimating the kinetic parameters and should primarily be used to describe adsorption kinetics.