贵州南部中三叠世鱼鳞蛤组合序列

鱼鳞蛤(Daonella)是中三叠世双壳类特征性化石。它在世界上分布广泛、演化迅速,具有重要地层对比意义。这一属在我国分布于广西、贵州、云南、川南和藏南。滇、黔、桂三省交界地区,产鱼鳞蛤的中三叠世地层十分发育,在这里,鱼鳞蛤常常是地层划分的重要古生物依据。因此,研究鱼鳞蛤的种群分布规律是这一地区地质工作中的一项重要任务。笔者最近对黔南的鱼鳞蛤层进行了调查,发现有许多种在纵横分布上均十分稳定。     

Biostratigraphy,microfacies and depositional environments of upper Viséan limestones from the Burren region,County Clare,Ireland

The Burren region in western Ireland contains an almost continuous record of Viséan (Middle Mississippian) carbonate deposition extending from Chadian to Brigantian times, represented by three formations: the Chadian to Holkerian Tubber Formation, the Asbian Burren Formation and the Brigantian Slievenaglasha Formation. The upper Viséan (Holkerian–Brigantian) platform carbonate succession of the Burren can be subdivided into six distinct depositional units outlined below. (1) An Holkerian to lower Asbian unit of skeletal peloidal and bryozoan bedded limestone. (2) Lower Asbian unit of massive light grey Koninckopora‐rich limestone, representing a shallower marine facies. (3) Upper Asbian terraced limestone unit with minor shallowing‐upward cycles of poorly bedded Kamaenella‐rich limestone with shell bands and palaeokarst features. This unit is very similar to other cyclic sequences of late Asbian age in southern Ireland and western Europe, suggesting a glacio‐eustatic origin for this fourth‐order cyclicity. (4) Lower Brigantian unit with cyclic alternations of crinoidal/bryozoan limestone and peloidal limestone with coral thickets. These cycles lack evidence of subaerial exposure. (5) Lower Brigantian bedded cherty dark grey limestone unit, deposited during the maximum transgressive phase of the Brigantian. (6) Lower to upper Brigantian unit mostly comprising cyclic bryozoan/crinoidal cherty limestone. In most areas this youngest unit is truncated and unconformably overlain by Serpukhovian siliciclastic rocks. Deepening enhanced by platform‐wide subsidence strongly influenced later Brigantian cycle development in Ireland, but localized rapid shallowing led to emergence at the end of the Brigantian. A Cf5 Zone (Holkerian) assemblage of microfossils is recorded from the Tubber Formation at Black Head, but in the Ballard Bridge section the top of the formation has Cf6 Zone (Asbian) foraminiferans. A typical upper Asbian Rugose Coral Assemblage G near the top of the Burren Formation is replaced by a lower Brigantian Rugose Coral Assemblage H in the Slievenaglasha Formation. A similar change in the foraminiferans and calcareous algae at this Asbian–Brigantian formation boundary is recognized by the presence of upper Asbian Cf6γ Subzone taxa in the Burren Formation including Cribrostomum lecomptei, Koskinobigenerina sp., Bradyina rotula and Howchinia bradyana, and in the Slievenaglasha Formation abundant Asteroarchaediscus spp., Neoarchaediscus spp. and Fasciella crustosa of the Brigantian Cf6δ Subzone. The uppermost beds of the Slievenaglasha Formation contain a rare and unusual foraminiferal assemblage containing evolved archaediscids close to tenuis stage indicating a late Brigantian age. Copyright © 2006 John Wiley & Sons, Ltd.     

Conodont Biostratigraphy and Age Determination of the Lower–Middle Triassic Boundary in South Guizhou Province, China

The demarcation of the Lower–Middle Triassic boundary is a disputed problem in global stratigraphic research. Lower–Middle Triassic strata of different types, from platform to basin facies, are well developed in Southwest China. This is favorable for the study of the Olenekian–Anisian boundary and establishing a stratotype for the Qingyan Stage. Based on research at the Ganheqiao section in Wangmo county and the Qingyan section in Guiyang city, Guizhou province, six conodont zones have been recognized, which can be correlated with those in other regions, in ascending order as follows: 1, Neospathodus cristagalli Interval-Zone; 2, Neospathodus pakistanensis Interval-Zone; 3, Neospathodus waageni Interval-Zone; 4, Neospathodus homeri-N. triangularis Assemblage-Zone; 5, Chiosella timorensis Interval-Zone; and 6, Neogongdolella regalis Range-Zone. An evolutionary series of the Early–Middle Triassic conodont genera Neospathodus-Chiosella-Neogongdolella discovered in the Ganheqiao and Qingyan sections has an intermediate type named Neospathodus qingyanensis that appears between Neospathodus homeri and Chiosella timorensis in the upper part of the Neospathodus homeri-N. triangularis Zone, showing an excellent evolutionary relationship of conodonts near the Lower–Middle Triassic boundary. The Lower–Middle Triassic boundary is located at 1.5 m below the top of the Ziyun Formation, where Chiosella timorensis Zone first appears in the Qingyan section, whereas this boundary is located 0.5 m below the top of the Ziyun Formation, where Chiosella timorensis Zone first appears in the Ganheqiao section. There exists one nearly 6-m thick vitric tuff bed at the bottom of the Xinyuan Formation in the Ganheqiao section, which is usually regarded as a lithologic symbol of the Lower–Middle Triassic boundary in South China. Based on the analysis of high-precision and high-sensitivity Secondary Ion Mass Spectrum data, the zircon age of this tuff has a weighted mean 206Pb/238U age of 239.0±2.9Ma (2s), which is a directly measured zircon U-Pb age of the Lower–Middle Triassic boundary. The Ganheqiao section in Wangmo county can therefore provide an excellent section through the Lower–Middle Triassic because it is continuous, the evolution of the conodonts is distinctive and the regionally stable distributed vitric tuff near the Lower–Middle Triassic boundary can be regarded as a regional key isochronal layer. This section can be regarded not only as a standard section for the establishment of the Qingyan Stage in China, but also as a reference section for the GSSP of the Lower–Middle Triassic boundary.     

Paleoenvironmental and biostratigraphic significance of uvigerinids and other foraminifera from the Bhuban Formation,Assam-Arakan Basin,Mizoram

Planktic and benthic foraminifera including uvigerinids are documented from the Upper Bhuban Formation, exposed at Thingdawl village, Kolasib district, Mizoram. The foraminiferal assemblage is poorly preserved and consists of index fossils useful for precise biochronology and interpretation of the depositional environment. A total of ten benthic and six planktic foraminiferal species are described. Six species belong to the genus Uvigerina, four to Ammonia, one species each of Globorotalia, Globigerinoides, Clavatorella, Praeorbulina and two species of the genus Orbulina. Based on lithological and foraminiferal assemblage, middle neritic to upper part of outer neritic paleobathymetry is inferred for the deposition of this part of the Upper Bhuban Formation. The foraminiferal assemblage suggests late Early Miocene to early Middle Miocene, (∼16 Ma) equivalent to planktic foraminiferal zones N8-N9 for the Upper Bhuban Formation.     

Correlation of Mississippian (Upper Viséan) foraminiferan,conodont, miospore and ammonoid zonal schemes,and correlation with the Asbian–Brigantian boundary in northwest Ireland

The microbiota of the upper Viséan (Asbian–Brigantian) rocks in the Lough Allen Basin in northwest Ireland is analysed. The Middle Mississippian sequence studied extends from the upper part of the Dartry Limestone/Bricklieve Limestone formations of the Tyrone Group to the Carraun Shale Formation of the Leitrim Group. The rocks have been traditionally dated by ammonoid faunas representing the B_2a to P_2c subzones. The Meenymore Formation (base of the Leitrim Group) also contains conodont faunas of the informal partial‐range Mestognathus bipluti zone. The upper Brigantian Lochriea nodosa Conodont Zone was recognized by previous authors in the middle of the Carraun Shale Formation (Ardvarney Limestone Member), where it coincides with upper Brigantian ammonoids of the Lusitanoceras granosus Subzone (P_2a). Foraminifera and algae in the top of the Dartry Limestone Formation are assigned to the upper Cf6γ Foraminifera Subzone (highest Asbian), whereas those in the Meenymore Formation belong to the lower Cf6δ Foraminifera Subzone (lower Brigantian). The Dartry Limestone Formation–Meenymore Formation boundary is thus correlated with the Asbian–Brigantian boundary in northwest Ireland. For the first time, based on new data, a correlation between the ammonoid, miospore, foraminiferan and conodont zonal schemes is demonstrated. The foraminiferans and algae, conodonts and ammonoids are compared with those from other basins in Ireland, northern England, and the German Rhenish Massif. Historically, the Asbian–Brigantian boundary has been correlated with several levels within the P_1a Ammonoid Subzone. However, the new integrated biostratigraphical data indicate that the Asbian–Brigantian boundary in northwest Ireland is probably located within the B_2a Ammonoid Subzone and the NM Miospore Zone, but the scarcity of ammonoids in the Tyrone Group precludes an accurate placement of that boundary within this subzone. Copyright © 2006 John Wiley & Sons, Ltd.     

Marine connections in North America during the late Maastrichtian: palaeogeographic and palaeobiogeographic significance ofJeletzkytes nebrascensisZone cephalopod fauna from the Elk Butte Member of the Pierre Shale, SE South Dakota and NE Nebraska

The Elk Butte Member of the Pierre Shale of southeast South Dakota and northeast Nebraska yields a late Maastrichtian cephalopod fauna of nautiloids, belemnites and ammonites of theFeletzkytes nebrascensisZone, best known from the near-shore facies of the Fox Hills Formation. ThenebrascensisZone is the highest distinct marine assemblage that can be recognised in the Western Interior, although ammonites occur as rarities high in the Lance Formation in Wyoming. Elements of the fauna occur in the Gulf Coast and Atlantic Seaboard, and extend into the highest Maastrichtian nannofossil Subzone CC26b, ofMicula prinsii, in Texas. These occurrences point to the existence of a southerly marine route for migration into and out of the northern Interior during the late late Maastrichtian. An analysis of Maastrichtian ammonite occurrences in West Greenland reveals no evidence for a marine link to the western Interior at this time, but rather indicates an open marine link to the North Atlantic region.The presence of upper upper Maastrichtian Pierre Shale in southeast South Dakota and northeast Nebraska, deposited in water depths that are conservatively estimated at 100-200 m, suggests that marine conditions (evidence for which has been removed by post-Cretceous erosion) may have extended well to the north of the shoreline position indicated in recent palaeogeographic reconstructions.     

广西隆或马平组的(竹蜓)类分带

<正> 广西隆林县隆或的马平组灰岩,与下伏黄龙组为整合接触,与上覆栖霞组由于第四纪坡积覆盖关系不清,厚215m。宜山马平组厚364m(赵金科,1947),德胜马平组厚481m(盛金章、侯祐堂等,1959),隆或马平组的厚度较小,但(竹蜓)类在地层上的垂直演化阶段仍是完整的,规律清楚。自下而上可分为四个(竹蜓)类化石带。 1.Montiparus带,2.Triticites带,3.Pseudoschwagerina带,4.Pseudofusulina带。各带特征详述如下: 1.Montiparus带 为最低的(竹蜓)类化石带,厚22m。以产旋壁蜂巢层发育较差的(竹蜓)     

Lower Aptian ammonite biostratigraphy and potential for further studies of OAE 1a in Bulgaria

Sediments of Early Aptian age in Bulgaria can be assigned to four different facies: platform carbonates (Urgonian complex), shallow-water siliciclastics, hemipelagic and flyschoid siliciclastics. The taxonomic analysis of the ammonite faunas of 18 sections from these four different facies resulted in a revision of the existing ammonite zonation scheme so far applied in Bulgaria and adjoining areas. A new biostratigraphic scheme, which bridges the western and eastern Tethys, is thereby proposed for the Lower Aptian of Bulgaria.The Upper Barremian Martelites sarasini Zone is characterized in its upper part by the Pseudocrioceras waagenoides Subzone in the shallow-water sections and by a horizon with Turkmeniceras turkmenicum in the deep-water settings. The Upper Barremian/Lower Aptian boundary is fixed by the first appearance of Paradeshayesites oglanlensis. For the Lower Aptian the following ammonite zones were established (from bottom to top): The Paradeshayesites oglanlensis Zone, the Deshayesites forbesi Zone (= formerly Paradeshayesites weissi Zone) including the Roloboceras hambrovi Subzone in the upper part, the Deshayesites deshayesi Zone including the Paradeshayesites grandis Subzone in the upper part and the Dufrenoyia furcata Zone. The Lower–Middle Aptian boundary has been defined by the appearance of species belonging to the genera Epicheloniceras and Colombiceras.The Lower Aptian ammonite faunas of Bulgaria, allow an interregional correlation with other areas of the Tethyan Realm. The presence of Turkmeniceras in the Upper Barremian enables a correlation with the Transcaspian region, whereas Roloboceras, Koeneniceras and Volgoceratoides found in the middle part of the Lower Aptian are more typical representatives of the ammonite faunas in northern Europe (England, Germany, Volga region).The analysis of the ammonite successions in combination with sedimentological observations enable us to conclude that the marls and marly limestones of the Lower Aptian studied here also cover the interval of the Oceanic Anoxic Event 1a. An interval of thin-laminated clays, rich in organic matter, was identified in the upper part of the D. forbesi Zone (Roloboceras hambrovi Subzone). This interval is characterized by a total lack of benthic faunas.     

Calpionellid biostratigraphy of the Upper Tithonian–Upper Valanginian interval in Western Sicily (Italy)

A revision of the calpionellid bio-chronostratigraphy in Western Sicily (Italy) was carried out, in order to update the taxonomy of this group, through the identification of some genera (Borziella, Longicollaria, Chitinoidella, Dobeniella, Sturiella, Borzaiella and Praecalpionellites) and species (Remaniella catalanoi Pop, Remaniella duranddelgai Pop, Remaniella colomi Pop, Remaniella borzai Pop, Remaniella filipescui Pop) recorded for the first time in the area. The studied sections, belonging to the Trapanese and Saccense Domains (Western Sicily), include at the base the upper portion of the Rosso Ammonitico Unit (Upper Tithonian), followed by the Lattimusa Formation (Lower Cretaceous). The quantitative and biostratigraphic analysis of the calpionellid assemblages allowed us to identify 13 assemblages and to define some important bioevents for the Upper Tithonian–Valangianian interval. The recorded calpionellid bioevents allowed us to recognize Zone/Subzone schema, which have been correlated with the bio-chronostratigraphy previously proposed for the Western Tethys. Editorial handling: M. Chiari (Guest) & J.-P. Billon-Bruyat     

Report on the 4th International Meeting of the IUGS Lower Cretaceous Ammonite Working Group, the “Kilian Group” (Dijon, France, 30th August 2010)

The 4th Kilian Group meeting (Dijon, France, 30th August 2010) focused on the Aptian and Albian Stages. For the Aptian, a two-fold division of the stage was adopted for the Mediterranean area with a boundary between the Dufrenoyia furcata and Epicheloniceras martini Zones. The main changes to the zonal scheme concern the Lower Aptian with: the introduction of a Deshayesites luppovi Subzone in the upper part of the Deshayesites oglanlensis Zone; the replacement of Deshayesites weissi by Deshayesites forbesi as new index-species of the second interval zone; the introduction of a Roloboceras hambrovi Subzone in the upper part of the D. forbesi Zone; and the subdivision of the D. furcata Zone into the D. furcata and Dufrenoyia dufrenoyi Subzones. For the Albian, the upper part of the Douvilleiceras mammillatum Zone (Lower Albian) is now characterized by a Lyelliceras pseudolyelli Subzone. The main amendments concern the Upper Albian. The base of this substage is defined by the base of the Dipoloceras cristatum Zone. Above it, the Upper Albian zonal scheme comprises in stratigraphic order the Mortoniceras pricei, Mortoniceras inflatum, Mortoniceras fallax, Mortoniceras rostratum, Mortoniceras perinflatum and Arrhaphoceras briacensis Zones.     

Newly Discovered Fish Faunas from the Early Triassic, Karoo Basin, South Africa, and their Correlative Implications

Recent collecting in exposures of the lowermost Burgersdorp Formation (Beaufort Group), of the Karoo Basin of South Africa, has revealed a previously unknown fish fauna from the Early Triassic (Scythian), lowermost Cynognathus Assemblage Zone (CAZ), which forms an important component of the total vertebrate assemblage. The newly discovered fish material includes lungfish, saurichthyids, and a large microfauna that includes numerous isolated chondrichthyan teeth, two fin spine fragments, and actinopterygian scales and teeth. The latest fish finds, together with the lowermost Cynognathus Assemblage Zone vertebrate faunas, make this Karoo Basin Assemblage Zone one of the most diverse Early Triassic faunal assemblages, comparable in faunal diversity to those from the Czatkowice Formation (Poland) and the Arcadia Formation (Australia). The presence of the lungfish Ptychoceratodus phillipsi in the early Middle Triassic Cynognathus Assemblage Zone (Subzone B), and in the underlying latest Early Triassic Cynognathus Assemblage Zone (Subzone A), indicates that these lungfish could serve as range index fossils within the CAZ, and thus are potentially useful biostratigraphic markers across the Early-Middle Triassic boundary. Furthermore the ‘new’ fish fauna provides a vital marine realm link in particular with the faunas of Madagascar and Australia, that is unavailable using the tetrapod faunal elements of the lower CAZ.     

Middle Westphalian plant fossils from the West Cumberland Coalfield,UK

Twenty-six form-species of plant macrofossils occur in the Coal Measures exposed in the Howgill Head Quarry, Whitehaven, Cumbria, UK. They belong to the Neuropteris semireticulata Subzone in the lower part of the Paripteris linguaefolia Zone, indicating the uppermost Duckmantian. They confirm that the sandstones exposed in this part of the West Cumberland Coalfield are part of the Whitehaven Sandstone Formation, and not a stratigraphically lower unit as previously suggested. The Howgill head fossils represent the only well documented example of a N. semireticulata Subzone assemblage in Britain, although comparable assemblages are known from coeval strata in northern France, Belgium, northern Germany and Poland.     

Ammonite biostratigraphy of the Albian in the Kirchrode II borehole, Hannover, Germany

The ammonite biostratigraphy of the 279.35 m of sediments of mid-Late Albian–Early Albian age traversed by the Kirchrode II (1/94) boring is described. The borehole was drilled in the Hermann-Löns Park, Kirchrode (Hannover), northwest Germany, in the central region of the Lower Saxony sedimentary basin. The core commenced within the Kirchrode Mergel Member of the Gault Formation in sediments of Callihoplites auritus Subzone age and showed a Late Albian ammonite zonal succession similar to that previously described by Wiedmann and Owen from the lower part of the nearby Kirchrode I (1/91) core, with which it is correlated. The thick underlying clay sediments of the Minimus Ton Member (Middle Albian–late Early Albian) provided a relatively sparse ammonite fauna. In the Middle Albian part of the sediment succession, several hiatuses are present and only sediments of the lower Euhoplites loricatus Zone (Anahoplites intermedius Subzone) and the Hoplites dentatus Zone (Hoplites spathi Subzone) have been identified. This is followed downward by a thick sedimentary succession through the upper part of the Early Albian Douvilleiceras mammillatum Superzone (Otohoplites auritiformis Zone). Earlier mammillatum and perhaps latest Leymeriella tardefurcata Zone portions of the core straddling the Minimus Ton/Schwicheldt Ton boundary, did not yield ammonites. The underlying sediments at the top of the Schwicheldt Ton Member, consist of dark clays and mudstones with a good representation of the Leymeriella (Neoleymeriella) regularis Subzone and the uppermost part of the Leymeriella acuticostata Subzone (Leymeriella tardefurcata Zone). Of particular importance is the succession through the sediments of the L. (N.) regularis Subzone, hitherto poorly known in north Germany. A brief comparison and correlation is made with other surface and borehole sections in northern Germany and elsewhere. The Boreal and more cosmopolitan Tethyan elements of the fauna are indicated and discussed. An appendix of ammonites obtained from the Mittellandkanal section at Misburg of latest Albian, Arraphoceras (Praeschloenbachia) briacensis Subzone age, completes the study.     

柴达木盆地北缘晚石炭世地层

本文对柴达木盆地北缘晚石炭世地层进行了划分、对比,分2个组,下部为克鲁克组,上部称扎布萨朵秀组。克鲁克组自下而上分为4个段。第2段建立(竹蜓)类Eostaffella su-bsolana-Pseudowedekindellina prolixa组合带;腕足类1—2段为Productus concinus-Choristitesyanghukouensis组合带;3段(竹蜓)类为Profusulinella-Pseudostaffella qinghaiensis组合带,4段(竹蜓)类为Fusulina-     

Late Albian (Early Cretaceous) ammonites from the Provincial Formation of central Cuba

A late Albian ammonite assemblage from the Provincial Formation of Villa Clara Province, Cuba is described. The Provincial Formation is a lithostratigraphic unit of Albian-Cenomanian age extensively exposed in central Cuba and formed within a part of the Caribbean Tethys that was volcanic during the Cretaceous. The formation is mainly composed of calcareous, terrigenous marine, and volcano-sedimentary deposits characterized by a series of micritic limestones intercalated with marls, sandstones, calcareous conglomerates, ash, and tuffaceous material. A rich assemblage of ammonites recovered from the calcareous biomicrites and marls is of late Albian (Stoliczkaia dispar Zone, Mortoniceras rostratum Subzone) age. The ammonite fauna shows a strong Tethyan affinity, and only a single hoplitid ammonite species was recorded. Although scarce, the first Cuban report of this and other boreal ammonite species now allows precise correlations to be made between Cuba and Albian sediments elsewhere in the world.     

新疆昆仑山金陵山一带晚石炭世纺锤虫

金陵山一带发现的晚石炭世纺锤虫类计有8属15种,其中新种3个,未定种3个,可称为Sphaeroschwagerina-Pseudofusulina动物群,与新疆柯坪地区康克林组Pseudoschwagerina带的下两个亚带对比。     

New Data of Macrofossils in the Ediacaran Wenghui Biota from Guizhou, South China

The abundance and diversity of macrofossils in the Ediacaran have attracted much attention. The upper Doushantuo macrobiotas in South China, including the Miaohe biota from Hubei and the Wenghui biota from Guizhou, are mainly preserved as the carbonaceous compressions and dominated by macroscopic algae and metazoans. Here, we describe 10 genera and 10 species (including 6 new genera and species) of macrofossils from the Wenghui biota. At present, the Wenghui biota yields macrofossils in more than 31 genera and 33 species, excluding those given no image and established on a few unascertained specimens. Based on the occurrence and distribution of macrofossils in both Miaohe and Wenghui areas, the middle-late Ediacaran (back shales of the upper Doushantuo Formation) in South China can be subdivided into two assemblage biozones in biostratigraphy: (i) the Protoconites–Linbotulitaenia–Eoandromeda–Anomalophton assemblage biozone characterized by abundant and diverse macrofossils and by the ranges of Protoconites, Linbotulitaenia and Anomalophton; and (ii) the Baculiphyca–Gesinella–Cucullus–Beltanelliformis assemblage biozone characterized by the acme of the longer macrofossils Baculiphyca and Gesinella, and by few shorter and discoidal macrofossils.     

上扬子区下志留统四射珊瑚组合带与层序地层序列的关系

摘 要　 根据动物群演替和层序地层的研究‚厘定上扬子区下志留统 （Llandovery） 四射珊瑚 组合带序列：（1） 香树园组 Dinophyllum-Rhabdocyclus 组合带‚时限为 Aeronian 期;（2） 雷家 屯组下部 Stauria-Briantelasma 组合带; （3） 雷家屯组上部 Maikottia-Kodonophyllum 组合带‚ 它们对应于 Telychian 早期;（4） 宁强组 Gyalophylloides -Idiophyllum 组合带‚相当于 Telychian 晚期。上扬子区下志留统可以划分为5个三级层序‚四射珊瑚组合带分别位于三级层序的高 水位体系域内。     

西藏喜马拉雅瑞替阶和赫塘阶菊石组合及其生物年代学对比

基于西藏聂拉木拉弄拉地区的格米格剖面新近逐层采集的菊石及缝合线研究,已识别出晚三叠世瑞替阶Choristoceras marshi组合,过渡带Neophyllites-Choristoceras组合,下赫塘阶Psiloceras calliphyllum组合和中赫塘阶Discamphiceras pleuronotum组合。作为东特提斯唯一的三叠系-侏罗系界线地层剖面,格米格菊石组合序列提供了特提斯范围内三叠/侏罗系界线菊石组合(或菊石带)以及地层对比的基础:Choristoceras marshi组合可以和欧洲,南美的秘鲁和智利的MARSHI菊石带,北美的内瓦达,不列颠哥伦比亚(加拿大)等地CRICKMAYI菊石带比较。Neophyllites-Choristoceras组合可和北美的内瓦达侏罗系底部TILMANNI菊石带Neophyllites和Choristoceras共生动物群比较。Psiloceras calliphyllum组合能够和北阿尔卑斯地区CALLIPHYLLUM菊石带比较, 证明北阿尔卑斯地区三叠系和侏罗系之间存在地层缺失。