科尔沁沙质草地植物群落的放牧退化及其自然恢复过程

 为了了解不同放牧强度对我国北方沙质草地的影响及其封禁停牧后草地植被的自然恢复过程,我们于1992—2006年在科尔沁沙地开展了草地放牧和封育试验,分析研究了人类放牧活动对草地植被的影响,比较了不同程度退化草地恢复速度及其所受气候变化影响。研究结果表明:①在科尔沁沙地,随着放牧强度的增加,草地植被受放牧影响的程度会加重,但只有持续重牧才会导致植被的严重退化,而且持续重牧对植被盖度、高度和草地生产力的危害程度要大于对物种丰富度和多样性的危害程度;中度放牧可以导致植被盖度、高度、生物量的轻度下降,但对物种丰富度、多样性没有不良影响;对于退化草地而言,持续轻牧可促进植被的有效恢复;②封禁停牧以后,草地原受干扰程度不同,其恢复的相对速度也不同,表现为原草地退化越严重,其恢复速度相对越快,而且是植被高度、盖度、生物量的恢复速度要大于物种丰富度、多样性和频度的恢复速度;③草地植被的恢复演替明显受到气候变化的影响,暖湿气候有利于植被的恢复,暖干气候对于植被的恢复不利。     

Effects of grazing and climate change on species diversity in sandy grassland, Inner Mongolia, China

To understand the effects of animal grazing activities and climate change on sandy grassland vegetation in northern China, a field grazing and protected enclosure experiment was conducted from 1992 through 2006 in Horqin Sand Land, Inner Mongolia. The results showed that (1) the grazing was primary responsible for changes of the vegetation richness and diversity in the grazing grassland and that changing climate was the main reason for changes in the species richness and diversity in the grassland protected from grazing; (2) light and moderate grazing can promote restoration of the richness and the diversity in the degraded grassland, and heavy grazing could result in a decrease of the richness and diversity; (3) heavy grazing can result in significant decrease of the perennial diversity, and moderate and light grazing promotes increase of the perennial diversity; the grazing, whether heavy or moderate and light grazing, was beneficial to increase of the annual diversity; (4) heavy grazing was not beneficial to diversity of Graminean and Chenopodiaceae, and moderate and light grazing was favorable the diversity of Compositae and Chenopodiaceae; (5) the warm-humid climate was favorable to increase of the richness and the diversity, and the warm-drought climate could result in decease of the richness and the diversity; (6) increased precipitation was favorable to perennial diversity and the diversity of Graminean, Leguminosae, and Compositae, and decreased precipitation had few effects on the annual diversity and Chenopodiaceae diversity.     

过牧及封育对红砂荒漠植被演替的影响

红砂灌丛植被是我国西北荒漠地区的主要放牧地,并在生态保护中具重要作用。本试验以内蒙古阿拉善盟不同过牧强度下的退化(轻度、中度和重度)红砂(Reaumuria soongorica)+无芒隐子草(Cleistogenes songorica)荒漠草地为对象。首次研究报道了不同过牧梯度和围栏封育对红砂植被演替的影响。经过6a过牧(平均1.8只羊·hm^-2),植被总盖度已由12%降至8%,平均每年盖度绝对值下降0.7%。随草地过牧加重红砂灌丛在逐渐消亡,在中度和重度退化样地的盖度已分别较轻度样地(7.5%)下降了74%和84% (P<0.05);质量差或产量低的草本在增加;无芒隐子草和匍根骆驼蓬(Peganum nigellastrum)的增加可分别作为该类草地中度和重度过牧演替的指示种。短期封育效果十分显著,封育一年的各样地平均较放牧样地比,植被盖度绝对值提高了1.7%,产草量增加2188%,红砂种子产量是放牧样地的371倍。然而,优势种红砂的产草量和盖度在封育期的相对恢复速率皆随样地退化加重而显著降低(P<0.05),重度退化区盖度封育当年的恢复速率为0。所以,封育是保护和恢复我国红砂荒漠植被的有效措施,但封育宜尽早开始,否则,红砂将恢复缓慢或难以复存。     

放牧强度对科尔沁沙地沙质草地植被的影响

为了研究科尔沁沙地沙质草地在不同放牧强度下植被特征的变化规律,比较了围封、中度放牧、重度放牧3种条件下的植被盖度、群落组成、物种多样性、地上/地下生物量和凋落物量等植被特征的变化特征。结果表明:(1)围封、中度放牧和重度放牧沙质草地的优势植物均为一年生植物,但围封和中度放牧中多年生禾本科植物优势度高于重度放牧,重度放牧沙质草地一年生植物所占优势度最大;(2)放牧强度对沙质草地植被盖度、地上生物量及凋落物量有显著影响。随着放牧强度的增加,植被盖度、地上生物量和凋落物量均降低;(3)围封、中度放牧和重度放牧沙质草地物种丰富度及多样性差异均不显著;(4)放牧强度和土壤深度及其交互作用均能影响地下生物量。随着放牧强度的增加,总的地下根系生物量减小。而重度放牧草地中, 3个土壤深度间地下根系生物量差异不显著。放牧改变了沙质草地植物群落的组成、极大降低了植被盖度和生产力;尤其是重度放牧急剧增加了一年生植物的优势、且对浅层根分布的减少影响最大。     

不同强度放牧后自然恢复的沙质草地土壤性状特征

通过对科尔沁沙质草地不同强度放牧后自然恢复过程中土壤的物理、化学和生物学性状特征的研究,结果表明,土壤有机碳、全氮、全磷、有效氮和速效磷含量以及基础土壤呼吸活性在各处理表层土壤的特征为:中牧后恢复草地>轻牧后恢复草地>无牧恢复草地>重牧后恢复草地;重牧后的恢复草地表层土壤较其它处理有较高的土壤容重和pH值以及较低的过氧化氢酶、脲酶和碱性磷酸酶活性;沙质草地的土壤养分和生物学活性在剖面中的分布为上高下低,主要富集于0~ 25cm表层,放牧对土壤系统的影响主要表现在0~ 75cm土层。重度放牧使沙质草地超出了其承载能力和承受干扰的阈限,土壤性状恶化,恢复力降低;而适度的放牧后恢复有利于土壤化学和生物学性状的保持和提高。     

Effects of Grazing on the Grassland Vegetation Community Characteristics in Inner Mongolia

The continuous increase of livestock production in Inner Mongolia has caused severe degradation of the grassland ecosystems in recent years. Previous grazing experiments have shown a wide range of vegetation responses between the biome types on a global scale, but there is still a lack of sufficient studies to discern the relative responses of a given biome type. We conducted a meta-analysis of vegetation coverage (VC), plant density (PD), total biomass (TB), above-ground biomass (AGB), under-ground biomass (UGB) and Shannon-Weaver Index (SI) in different grassland types in Inner Mongolia obtained under conditions of different grazing intensities and durations. The results showed that grazing decreased VC, TB, AGB, UGB, and PD significantly. Compared to the global and national average values, the negative effects of grazing to steppe biomass in Inner Mongolia were higher than that on the global scale, while less pronounced than that in China. TB of the meadow steppe in Inner Mongolia increased by 40% under moderate grazing intensity and duration because of compensatory growth. SI of the desert and meadow steppe showed negative linear relationships with the grazing intensity in Inner Mongolia. The percentage changes in AGB, PD, and SI to grazing showed quadratic relationships with the mean annual temperature of the experimental year. With increasing mean annual precipitation, the negative effects of grazing on UGB and SI first decreased and then increased, with that of VC and grazing showing a cubic relationship.     

中国草畜平衡状态时空演变指示的草地生态保护格局

中国草原牧区作为重要生态安全屏障和草地畜牧业生产基地,其草畜平衡状态直接影响草地退化与恢复,进而影响草地生态系统服务能力的强弱。本文分析了2000—2015年主要草原牧区草地植被覆盖、牧草供给、草畜平衡状态的时空变化特征,深入探讨草地退化与恢复及载畜压力下草地生态系统保护与恢复空间格局。结果表明：过去16年主要草原牧区草地面积净减少约163万hm²,6.7%的草地出现植被覆盖退化,而5.4%的草地呈现植被覆盖明显恢复。天然草地牧草供给量以增加为主,年增率约0.3 kg/hm²,然而其载畜压力亦持续增加,不考虑补饲的载畜压力指数高达3.8,除内蒙古东北部、青藏高原中部仍有载畜潜力,其余多处于超载状态;考虑实际冷季补饲的载畜压力指数约3.1,内蒙古中东部有所缓解;假设冷季全额补饲则载畜压力指数减至1.9,内蒙古、青藏高原等区域明显缓解。叠加上述数据,本文针对自然保护地、牧区、半农半牧区和农区等不同区域的草地生态保护格局,提出了平衡草地生态保护与畜牧生产利用的不同发展策略。     

Changes in intensity of wind erosion at different stages of degradation development in grasslands of Inner Mongolia, China

Overgrazing by increasing numbers of livestock in the Horqin Sandy Land of Inner Mongolia, China, has led to extensive degradation of the region's sandy steppes. Degraded grasslands are generally classified into four main types: fixed (least degradation), semi-fixed (light degradation), semi-shifting (moderate degradation) and shifting (severe degradation) sandy lands, representing four stages of degradation development. An experiment was conducted in the Horqin Sandy Land to investigate changes in intensity of wind erosion at different stages of degradation development in sandy grasslands and determine the extent to which surface wind erosion was affected by surface-related soil and vegetation factors through their effects on surface roughness length and wind regimes. Daily wind erosion rate was monitored at four sites of degraded grassland over an erosive period from 1 April to 10 June in 2001. Soil and vegetation properties for these sites were also measured twice: one in mid-April prior to the establishment of vegetation and again in mid-June after the establishment of vegetation. Relationships between surface roughness length and soil and vegetation variables were examined at each of the two stages of vegetation development. This study shows striking differences in the intensity of surface wind erosion among sites. The daily wind erosion rate in the fixed sandy land was, on average, only about 1/5 of the rate in the semi-fixed sandy land, 1/14 of the rate in the semi-shifting sandy land and 1/47 of the rate in the shifting sandy land suggesting a much higher resistance of the fixed sandy land to wind erosion compared to other sites. Differences in rate of wind erosion between sites were attributed to between-site differences in soil and vegetation properties that exerted significant effects on wind regimes by altering surface roughness length. At the pre-establishment stage of vegetation, surface roughness length was determined by a combination of litter amount on the ground, soil surface hardness and soil moisture content, with litter amount explaining the greatest proportion of the variation. At the post-establishment stage of vegetation, the development of the surface roughness effects was mainly governed by vegetation characteristics (vegetation cover in particular), while the effects of soil surface hardness and soil moisture on surface roughness length are likely to be masked by vegetation effects. The findings suggest that better management practices of restoring vegetation in degraded grasslands are required to reduce soil erosion losses and achieve a sustainable livestock production in the Horqin Sandy Land, an ecologically fragile sandy land ecosystem.     

蒙古高原草地利用特征及其国别差异

内蒙古草原是蒙古高原草原带的南部组成部分,是我国北方重要的生态屏障,草原生态修复任务重、压力大。本文以蒙古高原典型草原乌珠穆沁—温都尔汗样带为例,以1988—2016年5期TM数据为信息源,获取2016年样带草地利用信息基础上,按国别和年限对样带草地利用特征进行分析,揭示草地主要干扰因素,探寻有效的修复途径和可借鉴的国际经验。研究发现：① 草地面积占样带95.05%,均匀覆盖于整个样带;其他用地类型占比均不大,但分布相对集中,其中耕地集中于样带西北部蒙古国段,工矿用地、沙地、盐碱地集中于样带东南部内蒙古段。② 1988—2016年样带耕地面积减幅最大,为35.71%,集中位于蒙古国段;草地、水域面积逐年减少,在中国境内、境外变化趋势相同,但内蒙古段草地减幅大于蒙古国段且多转为未利用地;工矿用地面积增幅最大,为初期的近367倍,集中分布于内蒙古段;盐碱地、沙地面积逐年增加,在中国境内、境外的变化幅度相当。③ 草地是高原主要土地利用类型,放牧是草地主要利用方式,开矿是除放牧以外中国草地的主要干扰因子,蒙古国则为垦殖,内蒙古的草地干扰远大于蒙古国。草地退化与干旱化是蒙古高原面临的主要生态环境问题,在中国境内尤为突出。     

The conservation patterns of grassland ecosystem in response to the forage-livestock balance in North China

Being a key ecological security barrier and production base for grassland animal husbandry in China,the balance between grassland forage supply and livestock-carrying pressure in North China directly affects grassland degradation and restoration,thereby impacting grassland ecosystem services.This paper analyzes the spatiotemporal variation in grassland vegetation coverage,forage supply,and the balance between grassland forage supply and livestock-carrying pressure from 2000 to 2015 in North China.We then discuss the spatial pattern of grassland ecological conservation under the impacts of grassland degradation and restoration,and livestock-carrying pressure.Over the last 16 years,the total grassland area in North China decreased by about 16,000 km2,with vegetation coverage degraded by 6.7％ of the grasslands but significantly restored by another 5.4％ of grasslands.The provisioning of forage by natural grassland mainly increased over time,with an annual growth rate of approximately 0.3 kg/ha,but livestock-carrying pressure also increased continuously.The livestock-carrying pressure index without any supplementary feeding reached as high as 3.8.Apart from the potential livestock-carrying capacity in northeastern Inner Mongolia and the central Tibetan Plateau,most regions in North China are currently overloaded.Considering the actual supplementary feeding during the cold season,the livestock-carrying pressure index is about 3.1,with the livestock-carrying pressure mitigated in central and eastern Inner Mongolia.Assuming full supplementary feeding in the cold season,livestock-carrying pressure index will fall to 1.9,with the livestock-carrying pressure alleviated significantly in Inner Mongolia and on the Tibetan Plateau.Finally,we propose different conservation and development strategies to balance grassland ecological conservation and animal husbandry production in different regions of protected areas,pastoral areas,farming-pastoral ecotone,and farming areas,according to the grassland ecological protection patterns.     

我国北方沙区退化植被的恢复机理

通过讨论我国北方沙区退化植被的恢复机理,认为气候驱动机制、植被自我修复机制、土壤环境变化和景观格局的驱动机制是我国北方沙区退化植被自然恢复的四个主要机制。其中,气候机制决定着植被恢复演替的方向和终点,是植被恢复的外在驱动机制;植被的自我修复机制决定着植物恢复的起始速度和演替速度,是植被恢复的内在驱动机制;而景观驱动作用只发生于斑块较小且呈镶嵌分布的退化植被区,对于面积很大且连片分布的流沙裸地其作用微弱;土壤条件的改善既是植被对土壤作用的结果,又反馈于植被的演替,它决定着群落演替的阶段性和群落类型,自始自终伴随着植被的演替过程。     

放牧对退化草地近地面辐射的影响

随着环境变化,草原地表状态也会改变,尤其是近地面能量收支过程变化更明显。为深入理解半干旱草原地表辐射能量平衡过程对放牧的响应特征,利用内蒙古正镶白旗典型退化草原2020年生长季（6—10月）的辐射通量观测资料,对比分析了禁牧、放牧草地太阳总辐射、地表反射辐射、大气长波辐射、地表长波辐射、净辐射以及地表反照率的日变化和生长季变化规律的差异。结果表明：在生长季,太阳总辐射随着时间推进逐月递减;禁牧区的地表反射辐射总体小于放牧区;各月大气长波辐射日变化幅度很小,处于130—370 W·m^-2;禁牧、放牧条件下地表长波辐射存在明显的季节变化规律,但是二者之间的差异甚微。内蒙古典型退化草原生长季近地面辐射通量有显著单峰型日变化特征。禁牧、放牧草地地表反照率都呈现“U”型日变化规律。生长季放牧草地的反照率明显高于禁牧草地。在禁牧区,辐射分量（地表反射辐射、大气长波辐射）和植被指数（归一化差异植被指数）对净辐射的影响是正向极显著的;而地表反照率和另一辐射分量（地表长波辐射）对净辐射有显著的负向作用。在放牧区,地表反射辐射和大气长波辐射对净辐射有极显著的正向作用;而地表反照率和地表长波辐射对净辐射的作用则是负向显著的。植被状况是影响内蒙古典型退化草原近地表辐射能量收支过程的首要因子。     

中国草原牧区和半牧区草畜平衡状况监测与评价

草原超载过牧是造成我国草原大面积退化、沙化的主要原因之一,草畜平衡监测、评价和管理是实现我国草原植被恢复和重建的关键,也是研究的热点和难点。本文采用遥感和地面调查相结合的方法从宏观上监测和评价了农业部认定的264个牧区和半牧区县的草畜平衡状况,主要结论如下:(1)2008年监测区平均草畜平衡指数为33.58%,总体处于超载状态;(2)120个牧区县草畜平衡指数为27.37%,144个半牧业县为42.07%,半牧区县是我国实行草畜平衡管理的重点和难点;(3)六大牧区中牧区县2008年超载程度排序为:甘肃>四川>新疆>青海>西藏>内蒙古,半牧区县超载程度排序为青海>西藏>内蒙古>新疆>四川>甘肃。本文所得结论可为我国草原资源管理和保护提供参考价值和借鉴意义。     

科尔沁沙地植被恢复过程中群落组成及多样性演变特征

流动沙丘、固定沙丘和沙质草地均是科尔沁沙地土地沙化的产物,植被恢复重建则是该区域流沙治理的主要措施,理解植被恢复过程中群落组成及植物多样性演变特征有利于沙地植被恢复和生态重建.本研究基于连续15年对科尔沁沙地流动沙丘、固定沙丘和沙质草地长期植物群落结构调查数据,分析了 3种沙地类型植物群落年际变化特征及不同生活型植物年...     

科尔沁沙地退化植被恢复过程中碳氮化学计量特征的变化

为研究沙地退化植被恢复过程中碳氮化学计量特征的变化,分别于2011年、2013年和2015年8月中旬对流动、半固定、固定沙丘和草地进行植被调查,并测定植被—土壤系统的碳氮化学计量特征。结果表明：（1） 随着沙地退化植被的恢复,地上植物、凋落物和根系的C、N含量及C/N比波动变化,土壤（0～10 cm）C和N含量及C/N比显著增加（P <0.05）。（2）沙地4种生境上的地上植物、凋落物、根系、土壤的C、N含量及C/N比年际间变化显著（P <0.05）。（3） 沙地退化植被恢复过程中地上植物、凋落物、土壤的C/N比与物种丰富度呈正线性关系（P <0.01）。沙地退化植被恢复过程中优势植物演替导致植被—土壤系统中C、N化学计量特征的变化,植被恢复过程中植物的氮素利用效率也在逐渐增强,而封育时间的增加能够促进沙地土壤中N的积累。     

1981-2010年气候变化和人类活动对内蒙古地区植被动态影响的定量研究

基于GIMMS NDVI以及MODIS NDVI数据,分析内蒙古地区1981-2010年的植被变化趋势,并结合气候、社会经济数据,以旗县为单位定量分析气候变化和人类活动对植被变化的影响,结果表明：①1981-2010年间,内蒙古地区植被变化具有典型的空间异质性,其中植被显著增加区域主要集中在西南部的阿拉善盟、鄂尔多斯市以及东部通辽市等地区,显著减少区域主要集中在北部的锡林郭勒盟以及东北部的呼伦贝尔市的部分地区;②对于植被显著增加区域,人类活动作用的影响面积最大,其次为气候因素,气候与人类活动的耦合作用也对植被增加有一定显著影响;内蒙古西部降雨量的增加、围封禁牧政策的实施以及农作物播种面积的增加为驱动植被增加的主要因素;③对于植被显著减少区域,人类活动的作用略大于气候因素;内蒙古中东部地区降雨减少以及近10年来部分旗县风速的增加是导致植被显著减少的重要气候因素;虽然人工造林、农作物播种面积会增加局部植被盖度,但在县域尺度不足以抵消干旱对植被生长的不利影响,反而会导致区域植被退化。     

晚第四纪科尔沁沙地干湿变化的黄土记录

利用光释光（OSL）定年技术对科尔沁沙地南缘的典型黄土剖面进行了测年,结合两个剖面的磁化率、粒度、有机质含量等气候替代性指标,对沙地古气候及环境变化过程进行了分析。结果表明,晚第四纪科尔沁沙地的古气候经历了万年-千年时间尺度的多次干湿变化,特别是在146.96 ka BP前后、141.98 ka BP左右和107.06 ka BP前后发生过重大转变;相应地,沙地地表植被也经历了干冷多风条件下的干草原、荒漠草原和温暖湿润或温凉稍湿气候条件下的森林草原、疏林草原的多次变化。科尔沁沙地古气候的干湿变化可能是晚第四纪东亚冬、夏季风强弱变化的反映。     

羊啃食动力学模拟与内蒙古西部气候变化适应

将定性分析、民主参与式评估和定量数值模拟方法结合起来,对阿拉善气候变化适应问题进行了简要评估并重点分析了土地退化、放牧和草原管理策略之间的关系。阿拉善主要的适应政策是禁牧和生态移民。禁牧后荒漠草原植被得到恢复,但移民增加了绿洲压力,过度开垦又造成周边土地沙化;过度抽取地下水致使水位下降并伴随咸水入侵和地表盐碱化。禁牧的另一个副作用是造成草原鼠害加重及火灾风险增加,生物多样性也呈现先升后降趋势。一些灌木因缺乏啃食出现退化迹象等,说明一定程度的啃食有利于维持草原生态平衡。因此,动物是草原生态系统不可或缺的成员之一。我们用一个包含动物啃食等的森林生态动力模式模拟了啃食动力学,结果表明,有限的放牧有利于草场更新并能充分利用草原资源;过度放牧导致草场退化并伴随羊群死亡。因此,需要将圈养和限制性放牧结合起来,制定可持续性适应政策。     

青藏高原退化高寒草甸生长季承载力

近年来,在气候变化和人类活动的共同影响下,青藏高原部分地区草场退化严重。在青藏高原北麓河流域多年冻土区选取中度退化及未退化高寒草甸,用热红外辐射器模拟气候变暖,研究草地退化和气候变化对生长季草场数量和营养承载力（可消化蛋白承载力和代谢能承载力）的影响。结果表明：（1）退化后,莎草科植物在群落中的重要值降低,而禾本科和杂草类植物重要值逐渐增加;（2）退化对植被地上生物量无显著影响,但退化伴随着气候变暖使地上生物量在6月和9月分别显著降低了（87.17±6.93） g·m^-2和（38.89±2.23） g·m^-2;（3）退化在6月和9月使牧草粗蛋白含量分别显著降低了29.15%和33.74%,但使牧草中酸性洗涤纤维含量分别显著增加了11.68%和15.34%,牧草品质下降明显;（4）退化伴随着气候变暖使数量承载力和代谢能承载力在6月分别降低了（2.63±0.21）、（6.94±0.55）羊单位·hm^-2,在9月分别降低了（1.17±0.07）、（3.1±0.17）羊单位·hm^-2。研究区草场代谢能承载力>可消化蛋白承载力>数量承载力。研究区生长季牧草营养供给充足,适宜承载力为数量承载力     

Fundamental Strategic Shift Required by the Expensive Ecological Policy in Chinese Grasslands

Grasslands have critically important ecological and economic values while most of them have been suffering from various degrees of degradation in China due to overgrazing. The “Forage-Livestock Balance” (FLB) policy has been implemented for more than a decade, aims to balance the relationship between forage productivity and grazing consumption of grasslands by livestock. According to the review of statistical data and literatures on policy evaluation, FLB-dominated subsidies for grassland ecological conservation policies are ineffective on grassland restoration, livestock reduction in some overloaded areas and improvement on herdsmen livelihood. To deal with the dilemma, we suggest a fundamental shift of strategy from controlling livestock numbers to maintaining and improving grassland health (MIGH) based on ecological theories, and promote the sustainable development of grassland in China. The results show that, FLB policy failed to obtain expected benefits mainly because it interfered with the herders' autonomous use of contracted grasslands along with the defects of its underlying theory and methodologies. Implementing reward and punishment based on ecosystem health will not only motivate herders to manage their grassland autonomously, but also be more scientific and feasible than FLB.