Additional late Middle Cambrian trilobites from Karsha Formation (Haimanta Group) Zanskar region of Zanskar-Spiti basin, Northwest Himalaya

Additional latest Middle Cambrian polymerid trilobites are recorded from the Teta Member (Karsha Formation) along the Tangze-Yogma-Kuru section, Kurgiakh Valley, southeastern Zanskar, Northwest Himalaya. It includes record of a new genus Pianspis Saito & Sakakuru, 1936 of family Lisianiidae and others associated fauna comprises of Neoanomocarella asiatica (Lisianiidae), Parablackwelderia sp., (Damesellidae) and two undetermined pygidium and one unidentified cranidia. All these specimens are collected from the level representing the Lejopyge acantha Zone of Guzhangian Stage (Cambrian Series 3) of the Middle Cambrian. The fauna is mainly correlated with the South China, North Korea, Kazakhastan, Uzbekistan, Siberia and Australia.     

湘西王村剖面寒武系花桥组浊积岩特征及其大地构造意义

湘西王村地区的寒武系代表江南斜坡相沉积序列,传统的下寒武统划分为杷榔组、清虚洞组;中寒武统划分为敖溪组、花桥组（下部）;上寒武统划分为花桥组（上部）、追屯组。其中,花桥组中浊流沉积发育,可识别出7次规模较大的浊流活动期,第一期浊积岩包括69个浊积序列,分别归属于11个副层序;第二期浊积岩包括37个浊积序列,分别归属于7个副层序。每个副层序的下部表现为近源浊积岩,上部表现为远源浊积岩,所代表的海水深度变化由浅变深。第一期浊流活动发生在海平面上升时期,形成三级层序的海侵体系域;第二期浊流活动发生在海平面较低、斜坡较陡时期,形成三级层序的低水位体系域。江南斜坡带中—上寒武世浊积岩频繁出现反映台地边缘深大断裂周期性活动强,断裂两盘升降差异运动显著。     

The oldest bivalved arthropods from the early Cambrian of East Gondwana: Systematics,biostratigraphy and biogeography

The oldest bradoriid fauna from Australia, occurring in the lower Cambrian Ajax and Wirrapowie limestones of the Flinders Ranges, South Australia consists of eleven taxa, including one new genus and species, Quadricona madonnae gen. et sp. nov. and two new species, Liangshanella circumbolina sp. nov. and Zepaera jagoi sp. nov. In the Ajax Limestone, Liangshanella circumbolina sp. nov. occurs c. 20 m below the FAD of the zonal trilobite Abadiella huoi. This pre-trilobitic occurrence represents the oldest bivalved arthropod hitherto known from East Gondwana and suggests a lower Cambrian (Series 2, Stage 3) age for the assemblage. The recognition of distinct bradoriid assemblages associated with the Abadiella huoi (Atdabanian), Pararaia tatei, P. bunyerooensis and P. janeae (all Botoman) trilobite biozones in South Australia indicates great potential for future regional biostratigraphic correlation. Quantitative biogeographic analysis including new taxonomic data from the lower Cambrian of South Australia, highlights the strong endemism displayed by early Cambrian bradoriid communities and strengthens the close faunal affinities with South China and Antarctica.     

The oldest trilobites in Cambria: Early Cambrian trilobite faunas from the Llanberis Slates Formation,Gwynedd, North Wales

Historical published works on the fossil fauna of the Llanberis Slates Formation in Gwynedd, North Wales, concentrated on the endemic trilobite Pseudatops viola (Woodward, 1888), and placed the formation towards the top of Stage 3 (Series 2) of the Cambrian chronostratigraphy.The impression given was that the fauna is not particularly diverse and fossils are rare. However, recent collecting has produced a comprehensive fauna of sponges, hyolithids and diverse arthropods including over 250 trilobite specimens from two localities of the uppermost Green Slate horizon. Among them are examples of the endemic trilobite Pseudatops viola, sufficiently well preserved to enable a reappraisal of its status. The most abundant trilobite is a form of Strenuella (cf. strenua), but the collection includes previously unrecorded eodiscoid trilobites and specimens representing unrecorded and potentially new taxa. These additions result in the most comprehensive lower Cambrian trilobite assemblage yet described from Wales, and place it in the lower part of Stage 4, Series 2 in standard global terms.     

芙蓉统和排碧阶底界全球层型剖面的牙形刺生物地层

寒武系芙蓉统和排碧阶全球层型剖面——湖南花垣排碧剖面底界界线层的牙形刺生物地层,由下至上划分为Westergaardodina tetragonia带、Westergaardodina matsushitai带和Westergaardodina bicuspidate 3个牙形刺带,可与该界线层的三叶虫带进行很好对比;同时讨论了芙蓉统和排碧阶底界划在牙形刺Westergaardodina mat-sushitai带和Westergaardodina bicuspidata带之间的合理性;此外,还将该牙形刺分带与华北和东北地区寒武系牙形刺带进行了对比。     

A new early Cambrian bradoriid (Arthropoda) assemblage from the northern Flinders Ranges,South Australia

A new assemblage of early Cambrian bivalved arthropods (Bradoriida) is described from the Arrowie Syncline in the northern Flinders Ranges, South Australia. The well preserved, largely endemic fauna comprises a total of six taxa (including five new species): Jiucunella phaseloa sp. nov., Jixinlingella daimonikoa sp. nov., Mongolitubulus anthelios sp. nov., Neokunmingella moroensis sp. nov., Phasoia cf. spicata (Öpik, 1968), and Sinskolutella cuspidata sp. nov. This assemblage is derived from a carbonate sedimentary package representing a high energy, shallow water archaeocyath-Renalcis biohermal facies of Terreneuvian, Stage 2 age which transitions up-section to a more restricted, low energy, intra-shelf lagoonal environment that correlates with a Cambrian Series 2, Stage 3 age. The new taxon J. phaseloa sp. nov., has a first appearance datum (FAD) in shallow water biohermal facies of the Hideaway Well Member of the Wilkawillina Limestone at a level 47 m below the FAD of Pelagiella subangulata which is taken to approximate the base of Series 2, Stage 3 in South Australia. Along with Liangshanella circumbolina, this makes J. phaseloa sp. nov. amongst the oldest bivalved arthropods in South Australia and potentially greater Gondwana. The presence of 25 bradoriid taxa from the early Cambrian of South Australia suggests East Gondwana represents a major centre of origin for the Bradoriida.     

Biostratigraphy and ammonites of the Middle Oxfordian to lowermost Upper Kimmeridgian in northern Central Siberia

The Middle Oxfordian to lowermost Upper Kimmeridgian ammonite faunas from northern Central Siberia (Nordvik, Chernokhrebetnaya, and Levaya Boyarka sections) are discussed, giving the basis for distinguishing the ammonite zones based on cardioceratid ammonites of the genus Amoeboceras (Boreal zonation), and, within the Kimmeridgian Stage, faunas–for distinguishing zones based on the aulacostephanid ammonites (Subboreal zonation). The succession of Boreal ammonites is essentially the same as in other areas of the Arctic and NW Europe, but the Subboreal ammonites differ somewhat from those known from NW Europe and Greenland. The Siberian aulacostephanid zones—the Involuta Zone and the Evoluta Zone—are correlated with the Baylei Zone (without its lowermost portion), and the Cymodoce Zone/lowermost part of the Mutabilis Zone (the Askepta Subzone) from NW Europe. The uniform character of the Boreal ammonite faunas in the Arctic makes possible a discussion on their phylogeny during the Late Oxfordian and Kimmeridgian: the succession of particular groups of Amoeboceras species referred to successive subgenera is revealed by the occurrence of well differentiated assemblages of typical normal-sized macro and microconchs, intermittently marked by the occurrence of assemblages of paedomorphic “small-sized microconchs” appearing at some levels preceeding marked evolutionary modifications. Some comments on the paleontology of separate groups of ammonites are also given. These include a discussion on the occurrence of Middle Oxfordian ammonites of the genus Cardioceras in the Nordvik section in relation to the critical review of the paper of Rogov and Wierzbowski (2009) by Nikitenko et al. (2011). The discussion shows that the oldest deposits in the section belong to the Middle Oxfordian, which results in the necessity for some changes in the foraminiferal zonal scheme of Nikitenko et al. (2011). The ammonites of the Pictonia involuta group are distinguished as the new subgenus Mesezhnikovia Wierzbowski and Rogov.     

Paleobiogeography,paleoecology and paleoenvironmental significance of the Cambrian trilobites from the Zanskar region (Zanskar-Spiti-Kinnaur Basin), Northwest Himalaya

In present study the newly recorded latest Middle Cambrian trilobite fauna from the Cambrian succession of the Zanskar region of Zanskar-Spiti-Kinnaur Basin (Tethyan Himalaya) is analyzed critically to assess relationships with other Cambrian faunal elements of equatorial peri-Gondwanaland. The identification of genus Neoanomocarella, Parablackwelderia, Kunmingaspis, Fuchouia, Damesella and Dorypyge from the Cambrian of the Zanskar region and their comparison with those of South China and Australia is significant. It constitutes the basis for assessing the paleobiogeographic affinities during the Cambrian. The latest Middle Cambrian trilobite fauna from Zanskar shows proximity of Indian margin with that of southwest China “outboard” micro-continent. The recovery of analogous Middle Cambrian species i.e., Dorypyge perconvexlis, Fuchouia bulba, Fuchouia cf. oratolimba, Parablackwelderia sp. and Damesella sp. from the Zanskar region (Tethyan Himalaya) suggests a contiguous close proximity with south China and Australia during the latest Middle Cambrian, which supports the model of Meert and Van der Voo (1997) for assembly of Gondwanaland. The presence of Kunmingaspis in Zanskar and similar reports from northwestern Yunnan-Tibetan region, northern Henan, central and southeastern Hubei, north China, western Xinjiang and Yangtze platform reveal a close affinity between the Indian margin and the Yangtze platform during the Middle Cambrian. The trilobite fauna indicates the deeper shelf-shallow slope environment of deposition under fluctuating conditions of relative sea-level. The faunal elements of the Lejopyge acantha and Proagnostus bulbus zones indicate that the sea inundated the northern margin of Zanskar region during the latest Middle Cambrian time (Teta transgression) which is synchronous with globally recognized eustatic events during Lejopyge laevigata Zone.     

A new lower Cambrian shelly fossil biostratigraphy for South Australia

Definition of early Cambrian chronostratigraphic boundaries is problematic with many subdivisions still awaiting ratification. Integrated multi-proxy data from well-resolved regional-scale schemes are ultimately the key to resolving broader issues of global correlation within the Cambrian. In Australia, early Cambrian biostratigraphy has been based predominantly on trilobites. Phosphatic shelly fauna have great potential as biostratigraphic tools, especially in pre-trilobitic strata because they are widespread and readily preserved, but they have remained underutilised. Here we demonstrate their value in a new biostratigraphic scheme for the early Cambrian of South Australia using a diverse shelly fauna including tommotiids, brachiopods, molluscs and bradoriids.Biostratigraphic data are derived from ten measured stratigraphic sections across the Arrowie Basin, targeting Hawker Group carbonates including the Wilkawillina, Wirrapowie and Ajax limestones and the Mernmerna Formation. The stratigraphic ranges of shelly fossils are predictable and repeatable across the Arrowie Basin, allowing three discrete shelly biozones to be identified, spanning Terreneuvian, Stage 2 to Series 2, Stages 3–4. The Kulparina rostrata Zone (new) and part of the overlying Micrina etheridgei Zone (new) are pre-trilobitic (predominantly Terreneuvian). The Cambrian Series 2, Stage 3 Dailyatia odyssei Zone (new) features a very diverse shelly fauna and will be described in detail in a separate publication. These zones provide robust means to correlate Terreneuvian–Series 2 successions in neighbouring coeval basins in Australia, particularly the Stansbury Basin. Wider correlation is possible throughout East Gondwana, and especially with South China.     

贵州剑河八郎“清虚洞组”Olenoides Meek,1877的研究

贵州剑河八郎松山(九高山)寒武系"清虚洞组"产有多门类具软躯体的生物群——剑河生物群,其中三叶虫非常丰富,属于叉尾虫科(Dorypygidae)的拟油栉虫(Olenoides Meek,1877),是一个重要的三叶虫属。Olenoides全球分布,出现于寒武纪第二世(都匀期),灭绝于寒武纪第三世(古丈期晚期),是进行地层对比重要的三叶虫。松山(九高山)"清虚洞组"中的Olenoides包括2个种(含1个相似种):收缩拟油栉虫Olenoides contrictus(Chien,1961)及湖北拟油栉虫(相似种)O.cf.hubeiensis(Sun,1984),两者应为"清虚洞组"上覆地层凯里组产出的O.paraptus及O.hubeiensis的原始类型。Olenoides在寒武系黔东统都匀阶"清虚洞组"的发现,为其在研究区产出的最低层位。为Olenoides早期的形态、演化及时空分布的研究提供了重要的材料,也为寒武系都匀阶(第4阶)的生物地层研究提供了一些新的信息。     

An Ordovician age for the Muggort's Bay Lower Palaeozoic inlier,County Waterford,Ireland—the southernmost exposure of the Irish Caledonides

The most southerly exposed Lower Palaeozoic strata in Ireland occur on the southwest coast of County Waterford along a 2.5 km long coastal section at Muggort's Bay where they are surrounded by Devonian rocks. Five formations can be distinguished which, in ascending order, are: the Ballycurreen, Carrickbrean, Rathnameenagh, Moanbrack and Killinoorin formations. The total thickness of the succession is over 1800 m. No macrofossils are present, but the lithologies are largely fine‐grained turbidites and subordinate volcanic rocks which closely resemble the Ribband Group seen elsewhere in southeast Ireland and have previously therefore been classified with it. Palynological analysis was undertaken on 25 samples collected from Muggort's Bay, of which eight were productive. Diagnostic microfossils, comprising acritarchs, chitinozoans and scolecodonts, indicate an Early to Middle Ordovician age for both the Rathnameenagh and the Moanbrack formations. These ages confirm that the strata are part of the Ribband Group which elsewhere has been biostratigraphically dated as ranging from Mid‐Cambrian to Mid‐Ordovician. Reworked mid‐Middle Cambrian acritarchs occur in the Moanbrack Formation and reworked late Middle to early Late Cambrian acritarchs in the Rathnameenagh Formation. Despite generally poor preservation of the organic matter, some 20 acritarch species have been distinguished. Among these, three species belong to the herein revised genus Retisphaeridium for which an emended diagnosis is proposed together with two new combinations, Retisphaeridium capsulatum (Jankauskas, 1976 ) Vanguestaine nov. comb. and Retisphaeridium pusillum (Moczydlowska, 1998 ) Vanguestaine nov. comb. Copyright © 2005 John Wiley & Sons, Ltd.     

Reefs in the Early Paleozoic Taebaek Group, Korea: A Review

Various early Paleozoic (Cambrian Series 3–Middle Ordovician) reefs are found in the Taebaek Group, eastern Korea, located in the eastern margin of the Sino-Korean Block. They occur in every carbonate-dominant lithostratigraphic unit of the group, but their morphology and composition differ markedly. The Daegi Formation (middle Cambrian: Cambrian Series 3) contains siliceous sponge-Epiphyton reefs formed in a shallow subtidal environment, which is one of the earliest metazoan-bearing microbial reefs after the archaeocyath extinction. The Hwajeol Formation (upper Cambrian: Furongian) encloses sporadic dendrolites consisting of Angulocellularia, which developed in a relatively deep subtidal environment, representing a rare deeper water example. The onset of the Ordovician radiation resulted in the formation of microbialite–Archaeoscyphia–calathiid patch reefs in shallow subtidal deposits of the Lower Ordovician Dumugol Formation. Subsequent late Early Ordovician relative sea-level fall established extensive peritidal environments, forming microbial mats and stromatolites of the Lower–Middle Ordovician Makgol Formation. Ensuing Ordovician radiation resulted in one of the earliest metazoan skeletal reefs of the Middle Ordovician Duwibong Formation, constructed by stromatoporoid Cystostroma and bryozoan Nicholsonella, and developed around shallow shoals. These reefs reflect ongoing evolution and sea-level change during the early Paleozoic, and exemplify a rare glimpse of peri-Gondwanan records of reef evolution, which warrant detailed investigations and comparison with their counterparts in other regions.     

Peri-Gondwana Late Early–Middle Cambrian trilobites from the Kuhbanan Formation in Dahu section,Kerman Province,Iran

Outcropped of the Kuhbanan Formation at Dahu, near Zarand, about 63 km north of Kerman, Iran contains peri-Gondwana trilobites. In this study, 185 trilobite samples including six species and genera were identified and described from Dahu section. This trilobite’s assemblage including Redlichia noetlingi, Redlichia sp., Kermanella kuhbananensis, Kermanella lata lata, Kermanella lata minuta, Iranoleesia pisiformis, and Iranaspis sp. based on occurrence of the trilobite fauna a late Early to Middle Cambrian (Series 2–3) is suggested for this strata. These trilobite fauna help confirm conclusions from recent geological studies that place the Kerman Basin of Iran during the Cambrian.     

Cambrian stratigraphy of Kashmir and its boundary problems

The biostratigraphy of the Cambrian of Kashmir based on trilobite assemblage zones and trace fossils has been attempted. Faunal gaps occur in lower part of Lower Cambrian, upper Lower Cambrian, lower Middle Cambrian and upper part of Upper Cambrian. In the Lower Cambrian Cruziana—Rusophycus and Redlichia Zones, in the Middle Cambrian Solenopleura—Tonkinella, Tonkinella—Anomocare, Anomocare—Bailiella and Bolaspidella Zones, and in the Upper Cambrian Chuangia and Dikelocephalites Zones are recognized. The position of intrasystem boundaries is also discussed. While the various taxa have lithological preferences and are not necessarily found in a continuous sequence, an attempt has been made to interpret the ranges of different genera. The affinities of the fauna with that of other Cambrian basins of the world are discussed. It is concluded that the bulk of the fauna is of a cosmopolitan nature.     

The Yudomian of Siberia,Vendian and Ediacaran systems of the International stratigraphic scale

In Russia, the terminal Neoproterozoic formally includes the Vendian of western part of the East European platform and the concurrent Yudoma Group of Siberia. As is shown in this work, the designated subdivisions correspond in the stratotypes only to the upper, Yudomian Series of the Vendian. In the Siberian platform, the Ust-Yudoma and Aim horizons of the Yudomian are tightly interrelated. The lower of them, bearing remains of Ediacaran Fauna, represents the Ediacarian Stage, whereas the upper one containing small-shelled fossils (SSF) corresponds to the Nemakit-Daldynian Stage divided into the trisulcatus and antiqua superregional zones. In more complete sections of the platform periphery, sediments of these subdivisions conformably rest on siliciclastic succession that should be ranked as basal subdivision of the Yudomian. The succession is concurrent to the Laplandian Stage of the East European platform. According to geochronological dates obtained recently, the Yudomian Series spans interval of 600–540 Ma. In the East European platform, the Upper Vendian (Yudomian) begins with the Laplandian basal tillites of synonymous stage. In the west of the platform, tillites are dated at 600 Ma like the Upper Vendian base in Siberia. The next Ediacarian Stage of the East European platform is stratigraphic equivalent of the Redkino Horizon, while summary range of the Kotlin and Rovno horizons is concurrent to that of the Nemakit-Daldynian Stage. The Vendian of Russia is conformably overlain by the Tommotian Stage of the Lower Cambrian. Intense pre-Vendian events constrained distribution areas of the Lower Vendian sediments in Russia. The Lower Vendian deposits of the East European platform are most representative and well studied in the central Urals, where they are attributed to the Serebryanka Group. In Siberia, separate subdivisions representing the Lower Vendian are the Maastakh Formation of the Olenek Uplift, two lower members of the Ushakovka Formation in the Baikal region, and the Taseeva Group of the Yenisei Range. Chronological interval of the Lower Vendian corresponds to 650–600 Ma. The Marinoan Glaciation dated in Australia at 650–635 Ma is concurrent to basal part of the pre-Yudomian interval of the Vendian in Russia, whereas the Laplandian Tillite and Gaskiers Glaciation (600–580 Ma) correspond to onset of the Yudomian Epoch. The new Ediacaran System (Knoll et al., 2004) legalized in the International Neoproterozoic scale is close in range to the entire Vendian (635–544 Ma), although without basal beds (Marinoan Tillite) it deprives the terminal Neoproterozoic of its original sense. Inferiority of the system consists also in its indivisibility into stages. Hence, it is clear that the Vendian System subdivided in detail in Russia should be retained in the rank of terminal system of the Precambrian, one of the basic in general scale of the Neoproterozoic.     

台江阶——我国寒武系一个新的年代地层单位

台江阶是继湘西建立的 4个中 -上寒武统新阶之后在我国江南斜坡带黔东传统的中寒武统下部建立的又一个新阶 ,它的底界层型剖面位于贵州台江县革东镇八郎村 ,底界划在 Oryctocephalus indicus的首现位置 ,位于凯里组下部第 10层之底。     

A proposed GSSP for the base of the Middle Ordovician Series： the Huanghuachang section,Yichang, China

The Huanghuachang section near Yichang, southern China meets the requirements of Global Stratotype Section and Point (GSSP) for the base of the Middle Ordovician Series and the yet-to-be-named third stage of the Ordovician System (or lower stage of Middle Ordovician Series). The conodont succession at the section is complete across the Lower to Middle Ordovician series boundary and several excellent phylogenetic lineages of Baltoniodus, Trapezognathus, Periodon, and Microzarkodina are represented. The definition of the base of the Middle Ordovician is proposed to be the first appearance datum (FAD) of Baltoniodus? triangularis in the section. It is followed closely by the FAD of Microzarkodina flabellum, which is taken as a reasonable proxy for the boundary. This level approximates the boundary between the lower and upper intervals of the Azygograptus suecicus graptolite Biozone, and nearly coincides with the base of the Belonechitina henryi chitinozoan Biozone.The proposed GSSP for the base of the international Middle Ordovician Series is located in a roadside exposure at the base of Bed (SHod) 16, 10.57 m above the base of the Dawan Formation in the measured Huanghuachang section near Yichang City, southern China. The same faunal succession is also recorded from the Chenjiahe (formerly Daping) section, 5 km to the north of the Huanghuachang section. The proposed boundary horizon can be recognized and correlated globally with high precision in both relatively shallow-water carbonate facies as well as in deep-water graptolite facies.     

寒武系全球排碧阶及芙蓉统底界的标准层型剖面和点位

国际地球科学联合会2003年2月批准了由中美科学家组成的联合研究小组的建立全球排碧阶和芙蓉统底界标准层型剖面和点位的提案,以我国地名命名的“排碧阶”和“芙蓉统”两个全球年代地层单位及其底界的全球层型正式确立,这是寒武系内所确立的第一个“金钉子”。排碧阶和芙蓉统分别是寒武系的首个全球阶级标准单位和首个全球统级标准单位。排碧阶和芙蓉统的共同底界的全球层型剖面位于湖南花垣排碧附近的四新村,底界划在该剖面花桥组底界之上369.06m,与球接子三叶虫Glyptagnostus retuculatus的首现一致。芙蓉统的建立,取代了长期使用的非正式年代地层单位“上寒武统”。     

Bolorian and Kubergandian stages of the Permian in the Sanandaj-Sirjan zone of Iran

Sections and fusulinids of the Bolorian (presumably) and Kubergandian (lower part) stages in the Sanandaj-Sirjan tectonic zone are described. Two fusulinid assemblages are distinguished in a most complete section near Sirjan. The lower one is represented by Skinnerella, Paraleeina, and relatively primitive Misellina forms, whereas Armenina, Kubergandella, and Yangchienia species appear in the upper assemblage and suggest its early Kubergandian age. Accordingly, the lower assemblage is attributed to the Bolorian Stage, although it is lacking fusulinids typical of this stage except for the primitive Misellina morphotypes. Fusulinids from the Tange-Darchaleh section near Qomsheh (Shahreza) are typical of the Kubergandian Stage. The described three new species of the genus Skinnerella are close to morphotypes known from younger (Murgabian) deposits and represent their ancestral forms most likely.     

Stratigraphic significance of the Cambrian ichnofauna of the Zanskar region,Ladakh Himalaya,India

The Cambrian succession in the Zanskar Basin of Tethys Himalaya contains an abundant ichnofossils like in the other Tethyan Himalayan successions. The ichnofossils are stratigraphically important as they occur below the trilobite body fossils and are useful to define the basal part of the Cambrian. The ichnofossil assemblage reported from the Zanskar Basin of Ladakh Himalaya is significant to demarcate the Early Cambrian age due to lack of other faunal elements so far. The body fossils of trilobites recorded from the overlying beds indicates the earliest part of the Middle Cambrian age. Sixteen ichnogenera identified include: Bifungites, Cruziana, Diplichnites, Dimorphichnus, Isopodichnus, Lockeia, Merostomichnites; Monomorphichnus, Psammichnites, Palaeophycus, Planolites, Rhizocorallium, Skolithos, Taphrhelminthopsis, Teichichnus, Trepitichnus and trilobite scratch marks etc. The ichnogenera reported so far from this part of the Tethyan Himalayan region belongs mostly to the traces of arthropod origin. The ichnofauna ranges in age from Lower Cambrian to late part of the Middle Cambrian. The ichnofaunal assemblage can be assigned to repichnial, cubichinial, pascichnial, to fodinichnial behaviour. The distribution of ichnofossils in the studied sections shows that the ichnofossils are predominately less in occurrence in the sections were trilobites dominates and higher in the successions the abundance of ichnofossils decreases.