Standardization of CPUE for Chilean jack mackerel (Trachurus murphyi) from Chinese trawl fleets in the high seas of the Southeast Pacific Ocean

The generalized linear model (GLM) and generalized additive model (GAM) were applied to the standardization of catch per unit effort (CPUE) for Chilean jack mackerel from Chinese factory trawl fishing fleets in the Southeast Pacific Ocean from 2001 to 2010 by removing the operational, environmental, spatial and temporal impacts. A total of 9 factors were selected to build the GLM and GAM, i.e., Year, Month, Vessel, La Niña and El Niño events (ELE), Latitude, Longitude, Sea surface temperature (SST), SST anomaly (SSTA), Nino3.4 index and an interaction term between Longitude and Latitude. The first 5 factors were significant components in the GLM, which in combination explained 27.34% of the total variance in nominal CPUE. In the stepwise GAM, all factors explained 30.78% of the total variance, with Month, Year and Vessel as the main factors influencing CPUE. The higher CPUE occurred during the period April to July at a SST range of 12–15°C and a SSTA range of 0.2–1.0°C. The CPUE was significantly higher in normal years compared with that in La Niña and El Niño years. The abundance of Chilean jack mackerel declined during 2001 and 2010, with an increase in 2007. This work provided the relative abundance index of Chilean jack mackerel for stock assessment by standardizing catch and effort data of Chinese trawl fisheries and examined the influence of temporal, spatial, environmental and fisheries operational factors on Chilean jack mackerel CPUE.     

Standardizing CPUE of <Emphasis Type="Italic">Ommastrephes bartramii</Emphasis> for Chinese squid-jigging fishery in Northwest Pacific Ocean

Generalized linear models (GLM) and generalized additive models (GAM) were used to standardize catch per unit fishing effort (CPUE) of Ommastrephes bartramii for Chinese squid-jigging fishery in Northwest Pacific Ocean. Three groups of variables were considered in the standardization: spatial variables (longitude and latitude), temporal variables (year and month) and environmental variables, including sea surface temperature (SST), sea surface salinity (SSS) and sea level height (SLH). CPUE was treated as the dependent variable and its error distribution was assumed to be log-normal in each model. The model selections of GLM and GAM were based on the finite sample-corrected Akaike information criterion (AICC) and pseudo-coefficient (Pcf) combined P-value, respectively. Both GAM and GLM analysis showed that the month was the most important variable affecting CPUE and could explain 21.3% of variability in CPUE while other variables only explained 8.66%. The interaction of spatial and temporal variables weakly influenced the CPUE. Moreover, spatio-temporal factors may be more important in influencing the CPUE of this squid than environmental variables. The standardized and nominal CPUEs were similar and had the same trends in spatio-temporal distribution, but the standardized CPUE values tended to be smaller than the nominal CPUE. The CPUE tended to have much higher monthly variation than annual variations and their values increased with month. The CPUE became higher with increasing latitude-high CPUE usually occurred in 145°E–148°E and 149°E–162°E. The CPUE was higher when SST was 14–21°C and the SLH from −22 cm to −18 cm. In this study, GAM tended to be more suitable than GLM in analysis of CPUE.     

Evaluating the impact of spatio-temporal scale on CPUE standardization

This study focused on the quantitative evaluation of the impact of the spatio-temporal scale used in data collection and grouping on the standardization of CPUE(catch per unit effort).We used the Chinese squid-jigging fishery in the northwestern Pacific Ocean as an example to evaluate 24 scenarios at different spatio-temporal scales,with a combination of four levels of temporal scale(weekly,biweekly,monthly,and bimonthly)and six levels of spatial scale(longitude×latitude:0.5°×0.5°,0.5°×1°,0.5°×2°,1°×0.5°,1°×1°,and 1°×2°).We applied generalized additive models and generalized linear models to analyze the24 scenarios for CPUE standardization,and then the differences in the standardized CPUE among these scenarios were quantified.This study shows that combinations of different spatial and temporal scales could have different impacts on the standardization of CPUE.However,at a fine temporal scale(weekly)different spatial scales yielded similar results for standardized CPUE.The choice of spatio-temporal scale used in data collection and analysis may create added uncertainty in fisheries stock assessment and management.To identify a cost-effective spatio-temporal scale for data collection,we recommend a similar study be undertaken to facilitate the design of effective monitoring programs.     

Modeling a habitat suitability index for the eastern fall cohort of Ommastrephes bartramii in the central North Pacific Ocean

The eastern fall cohort of the neon flying squid, Ommastrephes bartramii, has been commercially exploited by the Chinese squid jigging fleet in the central North Pacific Ocean since the late 1990s. To understand and identify their optimal habitat, we have developed a habitat suitability index (HSI) model using two potential important environmental variables - sea surface temperature (SST) and sea surface height anomaly (SSHA) - and fishery data from the main fishing ground (165°-180°E) during June and July of 1999-2003. A geometric mean model (GMM), minimum model (MM) and arithmetic weighted model (AWM) with different weights were compared and the best HSI model was selected using Akaike’s information criterion (AIC). The performance of the developed HSI model was evaluated using fishery data for 2004. This study suggests that the highest catch per unit effort (CPUE) and fishing effort are closely related to SST and SSHA. The best SST- and SSHA-based suitability index (SI) regression models were SISST-based = 0.7SIeffort-SST + 0.3 SICPUE-SST, and SISSHA-based = 0.5SIeffort-SSHA + 0.5SICPUE-SSHA, respectively, showing that fishing effort is more important than CPUE in the estimation of SI. The best HSI model was the AWM, defined as HSI=0.3SISST-based+ 0.7SISSHA-based, indicating that SSHA is more important than SST in estimating the HSI of squid. In 2004, monthly HSI values greater than 0.6 coincided with the distribution of productive fishing ground and high CPUE in June and July, suggesting that the models perform well. The proposed model provides an important tool in our efforts to develop forecasting capacity of squid spatial dynamics.     

水温变动对2009年西北太平洋柔鱼产量下降的影响

分布在西北太平洋的柔鱼是我国远洋鱿钓渔业的重要捕捞对象,近些年来其产量一直处在稳定的水平。然而,2009年8～10月旺汛期间在传统作业渔场(150°E～165°E、38°E～46°E)柔鱼产量出现大幅度下降,其日产量仅为正常年份的一半。为此,根据2007～2009年8～10月我国在西北太平洋鱿钓生产数据,以及产卵场表层水温,探讨2009年柔鱼产量下降及渔场变动的原因。研究表明,其产量出现下降的原因可能有2个:(1)柔鱼产卵场(20°N～30°N,130°E～170°E)黑潮大弯曲的发生,使得21℃等温线向南偏移,使得柔鱼资源补充量受到影响,从而使得渔汛期间柔鱼产量的下降;(2)旺汛期间(8～9月)传统作业渔场(42°N～46°N,150°E～165°E)的100m水层有一个明显冷水南下,分布位置为154°E～156°E,将传统作业渔场(150E～165°E)一分为二,向南的前锋(水温低于5℃)到达42°N,明显不同于正常年份,使得作业渔场的范围明显缩小,不适合柔鱼的集群,导致产量出现大幅下降。     

基于频度统计和神经网络的北太平洋柔鱼渔场预报模型比较

利用2004~2010年北太平洋鱿钓船队生产数据和海洋环境数据,以海表温度(SST)1℃、海面高度(SSH)为1 cm、叶绿素a浓度(CHL-a)为0.1 mg/m3的间距,分析作业产量、CPUE与SST、SSH、CHL-a的关系,得到柔鱼渔场适宜环境因子范围,并将生产数据和环境数据匹配组成样本集,建立北太平洋柔鱼空间分布BP神经网络模型;利用2011年环境数据预报柔鱼渔场,并与2011年实际生产数据进行对比。结果表明,6~10月各月实际作业位置落入基于频度统计方法预报渔场的概率达90%以上;而BP模型预报的平均精度为79.2%,最低精度为52.5%。基于多环境因子的频度统计柔鱼渔场预报模型优于神经网络模型。     

A comparison of fishery biology of jumbo flying squid, Dosidicus gigas outside three Exclusive Economic Zones in the Eastern Pacific Ocean

Although many studies on the fishery biology of jumbo flying squid, Dosidicus gigas, have been conducted in the coastal areas within Exclusive Economic Zones (EEZs) of various countries due to its commercial and ecological importance, limited biological information is available from waters outside these EEZs. In this paper, we examined D. gigas fishery biology from waters outside Chilean, Peruvian and Costa Rican EEZs, based on the fishery data collected by Chinese jigging vessels during 2006 to 2010. The dominant mantle lengths of D. gigas were 350–450 mm, 250–400 mm and 250–350 mm outside Chilean, Peruvian and Costa Rican EEZs, respectively. Size structure analysis show that a medium-sized group existed mostly in the waters outside the Chilean and Peruvian EEZs, whereas a small-sized group occurred mainly in the waters outside the Costa Rican EEZ. The longevity of the squid outside the Costa Rican EEZ was less than 10 months, while most of those outside Chilean and Peruvian EEZs were about 1–1.5 years and very few large individuals were 1.5–2 years old. A higher percentage of mature individuals existed outside Costa Rican EEZ implying the region as a potential spawning ground, while lower proportions of mature squid outside the Peruvian and Chilean EEZs indicated that spawning may be occurring outside our study area. Spatial differences in sizes at maturity of the squid are thought to be result from different environmental factors especially different temperature and nutrition among the three areas. Stomach-content analysis showed that cannibalism was important in the diet of D. gigas. Stress generated by jigging may increase the incidence of cannibalism.     

Analyses of Trawling Track and Fishing Activity Based on the Data of Vessel Monitoring System (VMS)：A Case Study of the Single Otter Trawl Vessels in the Zhoushan Fishing Ground

The original purpose of Vessel Monitoring System(VMS) is for enforcement and control of vessel sailing. With the application of VMS in fishing vessels, more and more population dynamic studies have used VMS data to improve the accuracy of fisheries stock assessment. In this paper, we simulated the trawl trajectory under different time intervals using the cubic Hermite spline(c Hs) interpolation method based on the VMS data of 8 single otter trawl vessels(totally 36000 data items) fishing in Zhoushan fishing ground from September 2012 to December 2012, and selected the appropriate time interval. We then determined vessels’ activities(fishing or non-fishing) by comparing VMS speed data with the corresponding speeds from logbooks. The results showed that the error of simulated trajectory greatly increased with the increase of time intervals of VMS data when they were longer than 30 minutes. Comparing the speeds from VMS with those from the corresponding logbooks, we found that the vessels’ speeds were between 2.5 kn and 5.0 kn in fishing. The c Hs interpolation method is a new choice for improving the accuracy of estimation of sailing trajectory, and the VMS can be used to determine the vessels’ activities with the analysis of their trajectories and speeds. Therefore, when the fishery information is limited, VMS can be one of the important data sources for fisheries stock assessment, and more attention should be paid to its construction and application to fisheries stock assessment and management.     

A comparative study of spatially clustered distribution of jumbo flying squid (<Emphasis Type="Italic">Dosidicus gigas</Emphasis>) offshore Peru

We examined spatially clustered distribution of jumbo flying squid (Dosidicus gigas) in the offshore waters of Peru bounded by 78°–86°W and 8°–20°S under 0.5°×0.5° fishing grid. The study is based on the catch-per-unit-effort (CPUE) and fishing effort from Chinese mainland squid jigging fleet in 2003–2004 and 2006–2013. The data for all years as well as the eight years (excluding El Niño events) were studied to examine the effect of climate variation on the spatial distribution of D. gigas. Five spatial clusters reflecting the spatial distribution were computed using K-means and Getis-Ord Gi* for a detailed comparative study. Our results showed that clusters identified by the two methods were quite different in terms of their spatial patterns, and K-means was not as accurate as Getis-Ord Gi*, as inferred from the agreement degree and receiver operating characteristic. There were more areas of hot and cold spots in years without the impact of El Niño, suggesting that such large-scale climate variations could reduce the clustering level of D. gigas. The catches also showed that warm El Niño conditions and high water temperature were less favorable for D. gigas offshore Peru. The results suggested that the use of K-means is preferable if the aim is to discover the spatial distribution of each sub-region (cluster) of the study area, while Getis-Ord Gi* is preferable if the aim is to identify statistically significant hot spots that may indicate the central fishing ground.     

Estimating time-based instantaneous total mortality rate based on the age-structured abundance index

The instantaneous total mortality rate(Z) of a fish population is one of the important parameters in fisheries stock assessment. The estimation of Z is crucial to fish population dynamics analysis,abundance and catch forecast,and fisheries management. A catch curve-based method for estimating time-based Z and its change trend from catch per unit effort(CPUE) data of multiple cohorts is developed. Unlike the traditional catch-curve method,the method developed here does not need the assumption of constant Z throughout the time,but the Z values in n continuous years are assumed constant,and then the Z values in different n continuous years are estimated using the age-based CPUE data within these years. The results of the simulation analyses show that the trends of the estimated time-based Z are consistent with the trends of the true Z,and the estimated rates of change from this approach are close to the true change rates(the relative differences between the change rates of the estimated Z and the true Z are smaller than 10%). Variations of both Z and recruitment can affect the estimates of Z value and the trend of Z. The most appropriate value of n can be different given the effects of different factors. Therefore,the appropriate value of n for different fisheries should be determined through a simulation analysis as we demonstrated in this study. Further analyses suggested that selectivity and age estimation are also two factors that can affect the estimated Z values if there is error in either of them,but the estimated change rates of Z are still close to the true change rates. We also applied this approach to the Atlantic cod(G adus morhua) fishery of eastern Newfoundland and Labrador from 1983 to 1997,and obtained reasonable estimates of time-based Z.     

基于底拖网调查数据的渔业资源质量状况评价体系构建——以北部湾为例

【目的】构建渔业资源质量状况评价体系,评估渔业管理成效,服务海洋渔业可持续发展。【方法】以北部湾底拖网调查数据为基础,结合渔业统计数据,构建渔业资源质量状况评价指标体系。该体系分3个层次,下设7项指标。通过专家经验法和平均赋值法对指标进行赋值,得到各指标权重,建立质量状况评价等级指数,并利用该等级指数对北部湾渔业资源进行评价。【结果与结论】1992―2017年北部湾渔业资源评价指标中有6项呈下降趋势;质量状况评价等级指数波动在0.44~0.56之间。自20世纪90年代以来,北部湾渔业资源一直处于临界状态。该评价结果与北部湾渔业资源利用的实际状况基本相符,证明该方法具有合理性和可操作性,可为其他海区渔业资源质量状况评价提供借鉴。     

2004年北太平洋柔鱼钓产量分析及作业渔场与表温的关系

根据2004年5~11月我国鱿钓船在北太平洋生产数据,结合表温资料,按经纬度1°×1°的格式,利用Marineexplorer 4.0软件作图进行柔鱼钓产量及渔场与表温的关系分析。结果表明,5~7月在160°E以东海域作业,产量较低;8~10月在150°~160°E海域作业,为生产作业的产量高峰期,占总产量的62.5%;11月在150°E以西海域作业,产量也较低。在150°E以西海域CPUE最高,150°~160°E中部海域次之,160°E以东海域最低。作业渔场的适宜表温呈现出季节性变化。各月适宜表温分别为:5月12~14℃;6月15~16℃;7月14~16℃;8月18~19℃;9月16~17℃;10月15~16℃;11月12~13℃。     

Age, maturation, and population structure of the Humboldt squid Dosidicus gigas off the Peruvian Exclusive Economic Zones

Age, maturation and population structure of the Humboldt squid Dosidicus gigas were studied based on random sampling of the Chinese jigging fishery off the Peruvian Exclusive Economic Zones (EEZ) during 2008–2010. Estimated ages ranged from 144 to 633 days, confirming that the squid is a short-lived species with longevity no longer than 2 years. Occurrence of mature females and hatching in each month indicated that Humboldt squid spawned year-round. Back-calculated hatching dates for the samples were from January 22^nd, 2008 to April 22^nd, 2010 with a peak between January and March. Two size-based and two hatching date-based populations could be defined from mantle length (ML) at maturity and back-calculated hatching dates, respectively. Females matured at a larger size than males, and there was a significant difference in ML at maturity between the two hatching groups (P <0.05). The waters adjacent to 11°S off the Peruvian EEZ may be a potential spawning ground. This study shows the complexity of the population structure and large variability in key life history parameters in the Humboldt squid off the Peruvian EEZ, which should be considered in the assessment and management of this important resource.     

2006年北太平洋柔鱼作业渔场时空变化及其与表温的关系

<正>柔鱼(Ommastrephes bartramii)广泛分布在北太平洋,20世纪70年代初首先由日本鱿钓船开发,我国大陆于1993年开始利用该资源,1994年进行较大规模地商业性生产。目前北太平洋鱿钓渔业已成为我国远洋渔业的支柱[1]。据估计,历史上北太平洋柔     

智利外海南极褶柔鱼的形态学及部分生物学特征

描述了智利外海南极褶柔鱼Todarodes filippovae的形态特征,并参考前人研究资料,确定了其分类地位,同时对其作了初步分析。鉴定认为:该种系柔鱼科、褶柔鱼亚科、褶柔鱼属的南极褶柔鱼;其最北分布可到27°57′S,比原先文献报道的35°S偏北;渔获物的调查期间的胴长为157～366mm,体重为65～960g,性腺成熟度以Ⅰ、Ⅱ期为主,占总样本的71%。同时,还建立了胴长与体重、胴长与角质额喙长的关系式。     

西北太平洋秋刀鱼渔场分布及与海水表层温度的关系分析

根据2003~2004年西北太平洋秋刀鱼资源调查结果,对西北太平洋秋刀鱼渔场分布及其与海水表层温度(SST)的关系进行分析。结果表明,7~9月西北太平洋秋刀鱼渔场主要集中在40.5°N~44.5°N、151.5°E~158°E,SST为10℃~19℃,捕捞群体以中大型个体为主;各月最高产量及最大CPUE时的SST各不相同,渔场的形成和丰度与亲潮和黑潮的势力强弱及其分布密切相关。经K-S检验,结果表明,各月SST与产量及样本平均体长、平均体重的差异均不显著。这些渔场可作为我国远洋鱿钓渔业的兼作渔场。     

基于渔船AIS数据的南海北部海洋渔业捕捞强度空间特征挖掘

高精度渔业捕捞强度数据是开展捕捞限额管理的前提与关键,也是海洋渔业资源可持续发展的重要保障.因此,本文以挖掘海洋渔业捕捞强度空间特征为出发点,选用2018年2、4、9和11月典型季节的中国籍6364艘渔船1.8亿条高时空粒度AIS数据.运用专家知识经验、空间统计及数据挖掘分析方法,以广西南岸北部湾渔场、广东沿岸和环海南...     

Relationship between trawl selectivity and fish body size in a simulated population

Trawl is a main fishing gear in Chinese fishery,capturing large fish and letting small ones at large.However,long-term use of trawl would result in changes of phenotypic traits of the fish stocks,such as smaller size-at-age and earlier age-at-maturation.In this study,we simulated a fish population with size characteristics of trawl fishing and the population produces one generation of offspring and lives for one year,used trawl to exploit the simulated fish population,and captured individuals by body size.We evaluated the impact of the changes on selectivity parameters,such as selective range and the length at 50% retention.Under fishing pressure,we specified the selectivity parameters,and determined that smaller selection rates and greater length at 50% retention were associated with an increased tendency towards miniaturization.     

西北太平洋柔鱼资源与海洋环境的GIS空间分析

本文根据1995～2001年的西北太平洋地区(35°N～45°N,140°E～170°W)巴特柔鱼资源调查与生产的实际情况对柔鱼渔获量进行了研究,并利用同期遥感反演的海洋表层温度数据(SST)和近表层叶绿素a数据(Chlorophylla),拓展了GIS的空间分析功能,定量地研究了我国远洋柔鱼产量与水温、叶绿素等海洋要素场之间的关系,揭示西北太平洋柔鱼中心渔场的环境特征,以期为我国西北太平洋海区的鱿鱼生产服务。     

Natural mortality estimation and rational exploitation of purpleback flying squid Sthenoteuthis oualaniensis in the southern South China Sea

Based on the biological data of purpleback flying squid(Sthenoteuthis oualaniensis)collected by light falling-net in the southern South China Sea(SCS) during September to October 2012 and March to April 2013,growth and mortality of 'Medium' and 'Dwarf' forms of squid are derived using the Powell-Wetherall,ELEFAN methods and length-converted catch curves(FiSAT package).Given a lack of commercial exploitation,we assume total mortality to be due entirely to natural mortality.We estimate these squid have fast growth,with growth coefficients(k) ranging from 1.42 to 2.39,and high natural mortality(M),with estimates ranging from 1.61 to 2.92.To sustainably exploit these squid stocks,yield per recruitment based on growth and natural mortality was determined using the Beverton-Holt dynamic pool model.We demonstrate squid stocks could sustain high fishing mortality and low ages at first capture,with an optimal fishing mortality 3.0,with the optimal age at first capture increased to 0.4-0.6 years when fishing mortality approached optimal levels.On the basis of our analyses and estimates of stock biomass,we believe considerable potential exists to expand the squid fishery into the open SCS,relieving fishing pressure on coastal waters.