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A new trionychid turtle from the Early Cretaceous of Heilongjiang Province,Northeastern China
Institution:1. Key Laboratory of Evolutionary Systematics of Vertebrate, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Science, Beijing 100044, China;2. Palaeontological Research and Education Centre, Mahasarakham University, Kantarawichai, Mahasarakham 44150, Thailand;3. Geological Museum of Heilongjiang, Harbin 150090, China;1. Muséum National d’Histoire Naturelle, Département Histoire de la Terre : Sorbonne Universités–CR2P, UMR 7207 CNRS-MNHN-UPMC Paris 6, CP 38, 57, rue Cuvier, 75231 Paris Cedex 05, France;2. Estratigrafía y Paleontología, Facultad de Ciencia y Tecnología, Universidad del País Vasco UPV/EHU. Sarrienea s/n, 48940 Leioa, Spain;3. Museu Geològic del Seminari de Barcelona, Diputació 231, 08007 Barcelona – Geolab-Vic, Spain;1. Programa de Pós-graduação em Análise de Bacias e Faixas Móveis, Universidade do Estado do Rio de Janeiro (UERJ), Faculdade de Geologia, Rua São Francisco Xavier, 524, Maracanã, 20550-013, Rio de Janeiro, RJ, Brazil;2. Universidade Federal do Triângulo Mineiro (UFTM), Instituto de Ciências Exatas, Naturais e Educação (ICENE), Av. Randolfo Borges Jr. 1400, Univerdecidade, 38064-200, Uberaba, MG, Brazil;3. Universidade Federal do Triângulo Mineiro (UFTM), Complexo Cultural e Científico de Peirópolis (CCCP), Centro de Pesquisas Paleontológicas “Llewellyn Ivor Price” (CPPLIP), Uberaba, MG, Brazil;4. Museu de Paleontologia “Prof. Antonio Celso de Arruda Campos”, Centro de Artes, Praça do Centenário, 15910-000, Monte Alto, SP, Brazil;5. Museu de Paleontologia “Pedro Candolo”, Estação Cultura, Praça Farmacêutico Bruno Garisto, 15890-000, Uchoa, SP, Brazil;1. Queensland Museum Geosciences Program, 122 Gerler Road, Hendra, Queensland 4011, Australia;2. School of Geosciences Program, Monash University, Victoria 3800, Australia;3. 8 Manna Way, Silverdale, New South Wales 2752, Australia;1. Museo Geologico e Paleontologico ‘Giovanni Capellini’, Alma Mater Studiorum, Università di Bologna, Via Zamboni 67, 40126 Bologna, Italy;2. Dipartimento di Scienze Biologiche, Geologiche e Ambientali, Alma Mater Studiorum, Università di Bologna, Via Zamboni 67, 40126 Bologna, Italy;3. Dipartimento di Scienze della Terra, Università degli Studi di Napoli ‘Federico II’, Largo San Marcellino 10, 80138 Napoli, Italy;4. Grup de Recerca de Faunes del Mesozoic, Institut Català de Paleontologia “Miquel Crusafont”, c/ Escola Industrial, 23 E-08201 Sabadell, Spain;1. Natural History Museum of Geneva, Department of Geology and Palaeontology, Route de Malagnou 1, CP 6434, CH-1211 Geneva 6, Switzerland;2. IPHEP, UMR CNRS 7262, Université de Poitiers, UFR SFA, 6, rue M. Brunet, TSA 51106, 86073 Poitiers Cedex 9, France;3. Palaios Association, 86300 Valdivienne, France
Abstract:A new soft-shelled turtle (“Trionyxjixiensis sp. nov.) from the Lower Cretaceous Chengzihe Formation, Jixi city, Heilongjiang Province, China is described on the basis of a nearly complete carapace. The new species is diagnosed by the absence of suprascapular fontanelles and absence of a preneural; eight neurals, tetragonal fifth neural; and eight pairs of costals, with the large eighth costals meeting after the eighth neural. Due to the incompleteness of the specimen and confused classification of the genera of trionychids, the new species cannot be included in any genus of Trionychinae and is temporarily assigned to “Trionyx” (sensu lato). “Trionyxjixiensis is one of the earliest trionychids, its discovery indicates that the family was already diversified during the Early Cretaceous in Asia.
Keywords:Trionychidae  Trionychinae  Early Cretaceous  Chengzihe Formation  China
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