Abstract: | New evidence reveals that the first arthropods colonized dry land over 450 million years ago, more than 20 million years before the appearance of vascular plants. Traditionally, it was thought that animals only emerged on to land after plants had become established, thus providing a plentiful new food source. The earliest known terrestrial trace fossils occur in volcanogenic sediments from the English Lake District. The trace fossils are thought to have been made on land by a small millipede-like creature. These rocks contain no other fossils. |