Middle Pleistocene bivalves of the ?znik lake basin (Eastern Marmara, NW Turkey) and a new paleobiogeographical approach

?znik Lake is a tectonically originated basin mainly controlled by the E–W trending middle strand of the North Anatolian Fault (NAF) system. Pleistocene sediments occurring in front of the faults are well exposed both in the northern and in the southern shorelines of the basin. In this study, two endemic brackish water bivalve species, Didacna subpyramidata Pravoslavkev 1939 and Didacna nov. sp. were found in the oldest terrace of the northern Pleistocene sequence. Having characterized morphology, these species serve as stratigraphic indicators in the regional Pleistocene stratigraphy of the Ponto-Caspian region, and thus are well correlated to the assemblages of the early Khazarian subhorizon (Middle Pleistocene). Hence, these data demonstrate that the early Khazarian brackish water sea covered the study area. Additionally, a model for the formation of the basin is proposed: the ?znik lake basin was a gulf of the former Marmara Sea in the early Khazarian, connecting the Marmara to the Black Sea and the Caspian Sea. The subsequent regional prograding uplifts, main dextral strike-slip fault and many normal faults of the NAF Zone cut off the marine connections to the basin, leading to its present location and topographic level.