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Earliest mordellid-like beetles from the Jurassic of Kazakhstan and China (Coleoptera: Tenebrionoidea)
Authors:Tong Bao  Xuesong Zhang  Katarzyna S. Walczyńska  Bo Wang  Jes Rust
Affiliation:1. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Centre for Excellence in Life and Palaeoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China;2. Institut für Geowissenschaften und Meteorologie, Rheinische Friedrich-Wilhelms-Universität Bonn, Nussalle 8, 53115 Bonn, Germany;3. Beijing Museum of Natural History, 126 Tian Qiao Nan Street, Dongcheng District, Beijing, 100050, China;4. Institute of Oceanography, University of Gdansk, Al. Pilsudskiego 46, 81-378 Gdynia, Poland;5. Shandong Provincial Key Laboratory of Depositional Mineralization & Sedimentary Minerals, Shandong University of Science and Technology, Qingdao, Shandong 266590, China
Abstract:Within this study the subfamily Praemordellinae is reviewed and the holotype of Praemordella martynovi Scegoleva-Barovskaja, 1929 is re-described. The genera Cretanaspis Huang and Yang, 1999, Mirimordella Liu et al., 2007, Bellimordella Liu et al., 2008 and Wuhua Wang and Zhang, 2011 is attributed to Praemordellinae. One new species, Wuhua peregrina sp. nov., is erected based on two well-preserved specimens from the Middle Jurassic Daohugou Biota (Inner Mongolia, China). The morphological characters of Praemordellinae are given and a key to genera of Praemordellinae is presented. Praemordellinae is closely related to extant Mordellidae in having humpbacked body, strongly deflexed head, filiform antennae and pectinate tarsal claws, but it is differing in that the pygidium absent and hind femora not well developed. It is also similar to Ripiphoridae in having a convex body and simple tarsi, but differs Ripiphoridae in having filiform antennae. Praemordellinae is probably a stem group including the ancestor of Mordellidae and Ripiphoridae. Also, within this paper, the fossil record of Mordellidae is summarized.
Keywords:Coleoptera  Tenebrionoidea  Praemordellinae  Jurassic  Palaeoecology
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