Ornithomimidae (Dinosauria: Theropoda) from the Bissekty Formation (Upper Cretaceous: Turonian) of Uzbekistan |
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| Affiliation: | 1. Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, MRC 121, P.O. Box 37012, Washington, DC 20013-7012, USA;2. Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, 199034 Saint Petersburg, Russia;3. Department of Sedimentary Geology, Geological Faculty, Saint Petersburg State University, 16 Liniya VO 29, 199178 Saint Petersburg, Russia;1. Departament d''Estratigrafia, Paleontologia i Geociències Marines and Institut de Recerca de la Biodiversitat (IRBio), Facultat de Geologia, Universitat de Barcelona, E-08028, Barcelona, Spain;2. Departamento de Zoología y Antropología Física, Facultad de Biología, Universidad Complutense, E-28040, Madrid, Spain;3. Institut de Systématique, Évolution, Biodiversité, ISYEB, UMR 7205, CNRS, MNHN, UPMC, EPHE, Muséum national d''Histoire naturelle, Entomologie, Sorbonne Universités, F-75005, Paris, France;1. Prehistoric Museum, Utah State University Eastern, Price, UT 84501, USA;2. University of Colorado Museum, Boulder, CO 80309, USA;3. Oklahoma Museum of Natural History, Norman, OK 73072, USA;1. Instituto de Investigaciones en Paleobiología y Geología, Universidad Nacional de Río Negro, Avenida Roca 1242, General Roca, 8332, Río Negro, Argentina;2. Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Avenida Rivadavia 1917, C1033AAJ, Ciudad Autónoma de Buenos Aires, Argentina;3. Laboratório de Paleontologia de Vertebrados, Departamento de Paleontologia e Estratigrafia, Instituto de Geociências, Universidade Federal do Rio Grande do Sul (UFRGS), Ave. Bento Gonçalves 9500, Agronomia, 91540–000, Porto Alegre, RS, Brazil;4. Department of Anatomical Sciences and Neurobiology, University of Louisville, 511 Floyd Street (40202), Louisville, USA;5. Sección Paleontología de Vertebrados, Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Avenida Ángel Gallardo 490, C1405DJR, Buenos Aires, Argentina;1. Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, 199034 Saint Petersburg, Russia;2. Department of Sedimentary Geology, Geological Faculty, Saint Petersburg State University, 16 Liniya VO 29, 199178 Saint Petersburg, Russia;3. Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, MRC 121, P.O. Box 37012, Washington, DC 20013-7012, USA;1. Unidad de Paleontología, Facultad de Ciencias, Universidad Autónoma de Madrid, Cantoblanco, 28049 Madrid, Spain;2. Grupo de Biología Evolutiva, Facultad de Ciencias, Universidad Nacional de Educación a Distancia, Paseo Senda del Rey 9, 28040 Madrid, Spain |
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| Abstract: | The stratigraphically oldest remains of ornithomimid theropod dinosaurs are known from the Cenomanian Khodzhakul Formation and the Turonian Bissekty Formation of Uzbekistan. The ornithomimid from the Bissekty Formation is documented by more than 800 isolated bones that represent much of the skeleton. It shows at least three unambiguous synapomorphies of Ornithomimidae: length of anterior cervical centra three to five times greater than transverse width; low and rounded fibular crest of tibia; metatarsal III pinched between metatarsals II and IV; and proximal end of metatarsal III not visible in anterior view. Phylogenetic analysis, based on a dataset with 568 morphological characters and including all known ornithomimosaurian taxa, places the ornithomimid from the Bissekty Formation near the base of the ornithomimid radiation, between Archaeornithomimus asiaticus and Sinornithomimus dongi. |
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| Keywords: | Dinosauria Theropoda Ornithomimidae Cretaceous Uzbekistan |
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