Two new species of mid-Cretaceous webspinners in amber from northern Myanmar (Embiodea: Clothodidae,Oligotomidae) |
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| Institution: | 1. Division of Entomology, Natural History Museum, and Department of Ecology & Evolutionary Biology, 1501 Crestline Drive – Suite 140, University of Kansas, Lawrence, KS 66045, USA;2. Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024-5192, USA;3. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, People''s Republic of China;1. Institute of Cellular and Organismic Biology, Academia Sinica, 128 Academia Road, Section 2, Nankang, Taipei 11529, Taiwan;2. Sorbonne Universités, UPMC Université de Paris 06, CNRS, Biologie Intégrative des Organismes Marins (BIOM), Observatoire Océanologique, F-66650 Banyuls-sur-Mer, France;3. Marine Biology Research Division, Scripps Institution of Oceanography, University of California at San Diego, La Jolla, CA 92093, USA;1. School of Biomedical Sciences, Faculty of Medicine, The Chinese University of Hong Kong, Shatin, N.T., Hong Kong, China;2. Department of Neurology, The Third Affiliated Hospital of Sun Yat-Sen University, Guangzhou, Guangdong, China;3. Department of Anatomy and Embryology, School of Basic Medical Sciences, Peking University, Beijing, China;1. Department of Radiation Oncology, Osaka Medical Center for Cancer and Cardiovascular Diseases, Osaka, Japan;2. Department of Radiology, National Cerebral and Cardiovascular Center, Osaka, Japan;1. School of Chemical Engineering, Northwest University, Xi’an 710069, China;2. Xi’an Modern Chemistry Research Institute, Xi’an 710065, China |
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| Abstract: | Two new genera and species of Cretaceous webspinners (Embiodea) are described and figured, both preserved in amber from northern Myanmar. Atmetoclothoda orthotenes Engel and Huang, gen. et sp. nov., is the first fossil representative of the putatively primitive family Clothodidae, and is segregated into its own subfamily, Atmetoclothodinae Engel and Huang, subfam. nov., owing to its primitive retention of a distinct mentum, a quadrate submentum with straight borders, a postocciptal suture that meets the hypostomal sulcus at the posterior tentorial pit, and subgenae that do not meet medially (thus a ventral bridge or gula is lacking), completely symmetrical terminalia, and unsegmented cerci that apically bear a small inner lamellar projection. Litoclostes delicatus Engel and Huang, gen. et sp. nov., is the first definitive fossil of Oligotomidae (not including a subfossil species in Pleistocene copal and which is likely synonymous with an extant taxon), and differs from modern genera in a combination of head, tarsal, and terminalic morphology. The new fossil species double the known Cretaceous fossils and add a further two families to the Mesozoic record for the order. |
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| Keywords: | Cenomanian Clothodidae Embioptera Oligotomidae Polyneoptera Taxonomy |
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