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Small-sized abelisaurid (Theropoda: Ceratosauria) remains from the Upper Cretaceous of northwest Patagonia,Argentina
Institution:1. CONICET, Consejo Nacional de Investigaciones Científicas y Técnicas, Argentina;2. Museo Municipal “Ernesto Bachmann”, Dr. Natali S/N, Villa El Chocón, 8311, Neuquén, Argentina;3. Instituto de Investigación en Paleobiología y Geología, Universidad Nacional de Río Negro, Museo Carlos Ameghino, Belgrano 1700, Paraje Pichi Ruca (predio Marabunta), Cipolletti, 8300, Río Negro, Argentina;4. Laboratorio de Anatomía Comparada y Evolución de los Vertebrados, Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Av. Ángel Gallardo 470, C1405DJR, Buenos Aires, Argentina;1. State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, Beijing 100083, China;2. School of the Earth Sciences and Resources, China University of Geosciences, Beijing 100083, China;3. Department of Organismal Biology and Anatomy, University of Chicago, Chicago, IL 60637 USA;4. Yingliang Stone Nature History Museum, Nan''an, 362300, China;5. Department of Biological Sciences, University of Alberta, Edmonton, Alberta T6G 2E9 Canada;1. Canadian Museum of Nature, P.O Box 3443 STN ‘D’, Ottawa, Ontario K1P 6P4, Canada;2. Shanxi Museum of Geology, Binhexilu Zhongduan, Taiyuan, Shanxi 030024, PR China;3. Department of Biology, Carthage College, 2001 Alford Park Drive, Kenosha, WI 53140, USA;4. Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, P. O. Box 643, Beijing 100044, PR China;5. University of Chinese Academy of Sciences, Beijing, China;1. Área Paleontología, Museo Nacional de Historia Natural, Casilla 787, Santiago, Chile;2. Red Paleontológica U-Chile, Departamento de Biología, Facultad de Ciencias, Universidad de Chile, Las Palmeras, 3425, Santiago, Chile;3. Laboratorio y Museo de Dinosaurios, Facultad de Ciencias Exactas y Naturales, Universidad Nacional de Cuyo, Av. Padre Contreras 1300, Edificio ECT, Parque General San Martín 5500, Mendoza, Argentina;4. ICB-CONICET UNCUYO, Argentina;5. Department of Biology, University of the South, Sewanee, TN, USA;6. Marchant Pereira 1931, D.305 G, Santiago, Chile;7. Consultora Paleosuchus Ltda, Huelen 165C, Providencia, Santiago, Chile;1. Museu Nacional/Universidade Federal do Rio de Janeiro, Departamento de Geologia e Paleontologia, 20940-040, Rio de Janeiro, RJ, Brazil;2. Museu de Zoologia da Universidade de São Paulo, Laboratório de Paleontologia, Avenida Nazaré, 48, Ipiranga, 04263-000, São Paulo, SP, Brazil;1. Laboratoire de Géologie de l’École Normale Supérieure (UMR CNRS 8538), 24, rue Lhomond, 75231 Paris cedex 5, France;2. Muséum d’Histoire Naturelle, Hôtel Boyer d’Eguilles, 6, rue Espariat, 13100 Aix-en-Provence, France
Abstract:A new abelisaurid theropod from the Cenomanian of Candeleros Formation (Neuquén Province, NW Patagonia, Argentina) is described. It includes an isolated frontal, partial pelvis, two fused sacral centra, femoral shaft, rib fragments, partial metatarsal II and shaft of metatarsal III. A paleohistological analysis shows that MMCh-PV 69 was somatically mature at time of death. The prevalence of relatively ordered intrinsic fibers in the primary bone indicates a relatively lower growth rate of MMCh-PV 69 compared with other abelisauroids, such as Aucasaurus garridoi. Phylogenetic analysis found MMCh-PV 69 as a basal abelisaurid, sister group of the node formed by Majungasaurinae and Brachyrostra. We estimated the body mass of MMCh-PV 69 in 240 kg, which makes it one of the smallest abelisaurids recorded. This new form adds to the extensive theropod fauna of the Candeleros Formation, probably the most diverse association of meat-eating dinosaurs recorded in a Cretaceous Formation from South America.
Keywords:Abelisauridae  Patagonia  Cretaceous  Candeleros Formation  Paleohistology
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