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The first water measurers from the Lower Cretaceous amber of Spain (Heteroptera,Hydrometridae, Heterocleptinae)
Institution:1. Departament d''Estratigrafia, Paleontologia i Geociències Marines and Institut de Recerca de la Biodiversitat (IRBio), Facultat de Geologia, Universitat de Barcelona, E-08028, Barcelona, Spain;2. Departamento de Zoología y Antropología Física, Facultad de Biología, Universidad Complutense, E-28040, Madrid, Spain;3. Institut de Systématique, Évolution, Biodiversité, ISYEB, UMR 7205, CNRS, MNHN, UPMC, EPHE, Muséum national d''Histoire naturelle, Entomologie, Sorbonne Universités, F-75005, Paris, France;1. Institute of Energy and Environment, University of São Paulo, Av. Prof. Luciano Gualberto 1289, 05508-010, São Paulo, SP, Brazil;2. Institute of Geosciences, University of São Paulo, Rua do Lago 562, 05508-080, São Paulo, SP, Brazil;3. CNPq, Brazil;4. School of Arts, Sciences and Humanities, University of São Paulo, Av. Arlindo Bettio 1000, 03828-000, São Paulo, SP, Brazil;5. Centre of Technology and Geosciences, Federal University of Pernambuco, Av. Acadêmico Hélio Ramos s/n, 50740-530, Recife, PE, Brazil;1. CNRS UMR 7205, Muséum National d''Histoire Naturelle, CP 50, Entomologie, 45 rue Buffon, F-75005 Paris, France;2. Adel Halabi Building, Baeth Street, Jaramana, Damascus Country Side, Syria;3. Lebanese University, Faculty of Sciences II, Department of Biology, Fanar – Matn, P. O. Box 26110217, Lebanon;1. Institut für Geowissenschaften und Meteorologie, Rheinische Friedrich-Wilhelms-Universität Bonn, Bonn, 53115, Germany;2. Department of Entomology, National Museum, Praha, 14800, Czech Republic;1. Hunan Province Key Laboratory of Economic Crops Genetic Improvement and Integrated Utilization, Hunan University of Science and Technology, Xiangtan 411201, China;2. Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China;3. Department of Entomology, China Agricultural University, Beijing 100193, China;4. College of Life Sciences and Academy for Multidisciplinary Studies, Capital Normal University, Beijing 100048, China;1. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, Jiangsu 210008, China;2. Lebanese University, Faculty of Science II, Fanar, Natural Sciences Department, Fanar – Matn, PO Box 26110217, Lebanon;3. Muséum National d''Histoire Naturelle, Institut de Systématique, Evolution, Biodiversité, ISYEB, UMR 7205 CNRS UPMC EPHE, CP50, Sorbonne Universités, 45 rue Buffon, F-75005 Paris, France
Abstract:Two fossils belonging to a new genus and species of water measurer (Gerromorpha, Hydrometridae), Alavametra popovi Sánchez-García and Nel gen. n., sp. n., are described as first definitive record of the family in Lower Cretaceous (upper Albian) amber from the Utrillas Group (Peñacerrada I site) in Spain. Although several parts of the specimens are obscured due to preservation, a sufficient number of taxonomical characters are visible to consider adequate placement within Heterocleptinae, including the very long posterior pair of cephalic trichobothria inserted on tubercles and the preapical articulation between the first and second antennal segments. The new fossil taxon is included into a cladistic analysis with extinct and extant hydrometrids, and it results putatively basal among the subfamily Heterocleptinae, suggesting that this clade was already present 105 Ma ago.
Keywords:Gerromorpha  Taxonomy  Phylogeny  Albian  Mesozoic  Spain
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