Vertebrate remains from the Upper Cretaceous (Santonian) Ajka Coal Formation,western Hungary

Vertebrate remains from the Upper Cretaceous (Santonian) Ajka Coal Formation (Bakony Mountains, western Hungary) are described. Macro- and microfossils collected from two boreholes and from isolated chunks of sediment/matrix dumped on spoil heaps of the Jókai Mine represent pycnodontiform and lepisosteiform fishes, bothremydid turtles, the mosasauroid Pannoniasaurus inexpectatus, the crocodyliforms cf. Theriosuchus, Iharkutosuchus makadii and cf. Allodaposuchus, as well as ankylosaurian and theropod dinosaurs. This unit was deposited in a swampy lacustrine environment, in contrast with the neighbouring and contemporaneous floodplain deposit of the vertebrate-bearing Csehbánya Formation at Iharkút. Despite significant environmental differences, the faunal composition of the Ajka Coal Formation assemblage completely overlaps with that of the Csehbánya Formation, suggesting the occurrence of the same semi-aquatic and terrestrial species in both settings. The ankylosaurian remains further strengthen the previous view that ankylosaurus preferred wetland habitats such as fluvial systems and coastal regions.