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Ptismidae fam. nov. (Coleoptera,Staphyliniformia) from the Lower Cretaceous Lebanese amber
Affiliation:1. Zoological Institute of the Russian Academy of Sciences, Universitetskaya emb., 1, St. Petersburg, 199034, Russia;2. CNRS UMR 7205, Muséum National d''Histoire Naturelle, Sorbonne Universités, CP 50, Entomologie, 45, rue Buffon, F-75005, Paris, France;3. Department of Invertebrate Zoology, St. Petersburg State University, Universitetskaya emb.,7/9, St. Petersburg, 199034, Russia;4. Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing, 210008, PR China;5. Lebanese University, Faculty of Sciences II, Department of Natural Sciences, Fanar, El-Metn, PO Box. 26110217, Lebanon;6. 6th Linia, 49-34, St. Petersburg, 199004, Russia;1. Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Vladivostok, 960022, Russia;2. Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, ul. Bogdana Khmel''nitskogo 15, Kiev, 01601, Ukraine;1. Museum of Natural History, University of Wrocław, Sienkiewicza 21, 50-335, Wrocław, Poland;2. Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Bogdana Khmel''nitskogo 15, Kiev, 01601, Ukraine;3. Paleontological Institute, Russian Academy of Sciences, 123, Profsoyuznaya ul, Moscow, Russia;1. Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China;2. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China;1. Department of Integrative Biology, Oregon State University, Corvallis, OR 97331, USA;2. 629 Euclid Avenue, Berkeley, CA 94708, USA;1. Museum of Natural History, University of Wrocław, Sienkiewicza 21, 50-335, Wrocław, Poland;2. Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Bogdana Khmel''nitskogo 15, Kiev, 01601, Ukraine;1. Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China;2. State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China
Abstract:Ptisma zasukhae gen et sp. nov. is described from the Lower Cretaceous Lebanese amber and attributed to a new family Ptismidae fam. nov. with unclear position within the infraorder Staphylinoformia, hypothetically in the superfamily Staphylinoidea. This species is characterized by the rather small body size (0.59 mm) and structural similarities with different staphylinoid groups. The particular development of the legs of the new taxa shows some similarities with those of small and medium-sized jumping beetles from the superfamilies Scirtoidea, Buprestoidea, Tenebrionoidea, Chrysomeloidea and Curculionoidea. However, jumping beetles of these groups have strong femora and tibiae with large spurs and thick apical setae only on the posterior legs while P. zasukhae sp. nov. demonstrates such features on both mid and posterior legs. Another particularity of P. zasukhae sp. nov. is the unusual large femoral plate of metacoxae extending behind the suture between ventrites 2 and 3. Strong development of these structures are rare among Coleoptera and occurs in different taxonomic groups without close relationship. The studies were carried out with the usage of usual conventional light microscopy and confocal laser scanning microscopy.
Keywords:Coleoptera  Staphylinoidea  New family  New genus and species  Lebanese amber  Lower Cretaceous
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