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The turtles from the uppermost Jurassic and Early Cretaceous of Galve (Iberian Range,Spain): Anatomical,systematic, biostratigraphic and palaeobiogeographical implications
Affiliation:1. Departamento de Paleontología, Facultad de Ciencias Geológicas, Universidad Complutense de Madrid, C/José Antonio Novais, 2. 28040 Ciudad Universitaria, Madrid, Spain;2. Paläontologisches Institut und Museum, Universität Zürich, Karl-Schmid-Strasse 4, CH-8006 Zürich, Switzerland;3. Departamento de Estratigrafía y Paleontología, Facultad de Ciencia y Tecnología, Universidad del País Vasco, UPV/EHU, Apartado 644, 48080 Bilbao, Spain;1. Laboratoire de Géologie de Lyon: Terre, Planète, Environnement, UMR CNRS 5276 (CNRS, ENS, Université Lyon1), Ecole Normale Supérieure de Lyon, 69364, Lyon cedex 07, France;2. La Plage aux ptérosaures, Le Mas de Pégourdy, 46150, Crayssac, France;1. Grupo Recursos Minerales, Departamento de Ciencias de la Tierra, Universidad de Zaragoza, Pedro Cerbuna 12, 50.009 Zaragoza, Spain;2. Grupo Aragosaurus – IUCA, Departamento de Ciencias de la Tierra, Universidad de Zaragoza, Pedro Cerbuna 12, 50.009 Zaragoza, Spain;1. Conicet, Instituto de Investigación en Paleobiología y Geología, Sede Alto Valle-Valle Medio-Universidad Nacional de Río Negro, Av. Roca 1242, CP8332, Gral. Roca, Río Negro, Argentina;2. Conicet, Museo Carlos Ameghino, Belgrano 1700, Paraje Pichi Ruca (predio Marabunta), CP8300, Cipolletti, Río Negro, Argentina;1. Grupo de Biología Evolutiva, Facultad de Ciencias, UNED, Paseo de la Senda del Rey 9, 28040 Madrid, Spain;2. Fundación Conjunto Paleontológico de Teruel–Dinópolis (Museo Aragonés de Paleontología), Avda. Sagunto s/n, 44002 Teruel, Spain;1. Dinosaur Institute, Natural History Museum of Los Angeles County, 900 Exposition Blv, 90007 CA, Los Angeles, USA;2. Grupo de Biología Evolutiva. Facultad de Ciencias, UNED, Paseo de la Senda del Rey, 9, 28040 Madrid, Spain;3. Instituto Dom Luiz, Universidade de Lisboa, Campo Grande, 1749-016 Lisbon, Portugal
Abstract:The faunal associations of the uppermost Jurassic and Early Cretaceous Villar del Arzobispo, El Castellar, and Camarillas formations of the Spanish town of Galve (Maestrazgo Basin of the Iberian Range) are composed of more than fifty taxa belonging to Chondrichthyes, bony fishes, lissamphibians, mammals, and reptiles. Although the presence of turtles has been recognized in these three formations, being very abundant, the published information is limited. The detailed study of these turtles allows us to recognize at least nine taxa, several of them previously not identified in the Spanish record. These taxa correspond to stem Testudines (Solemydidae), members of Paracryptodira (Pleurosternidae), several representatives of Plesiochelyidae and taxa closely related with this clade, representatives of the stem group of Cryptodira (Xinjiangchelyidae) and members of crown Cryptodira. The chelonian fauna of Galve increases the systematic, anatomical, biostratigraphic, and paleobiogeographic knowledge on several clades, represented by taxa shared with other European regions, clades also present in Early Cretaceous sites of North America or Asia, as well as taxa exclusive of the Iberian record.
Keywords:Galve  Stem Testudines  Paracryptodira  Plesiochelyidae  Xinjiangchelyidae  Cryptodira
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