Bivalves and oncoids as palaeoenvironmental indicators at Late Jurassic and Early Cretaceous dinosaur sites from Spain

Bivalves from the Late Jurassic and Early Cretaceous stratigraphic section at Las Zabacheras (Galve Sub-basin Teruel, northern Spain), are reviewed from both systematic and palaeoautoecological perspectives. For this study the Villar del Arzobispo Formation, well known for important dinosaur occurrences, was sampled from the boundary with the underlying Higueruelas Formation (Late Jurassic), to the first levels of the overlying El Castellar Formation (Early Cretaceous). The bivalve taxa have Late Jurassic affinity, pointing to a possible Jurassic-Cretaceous boundary towards the top of the Villar del Arzobispo Formation. We have sampled oncoids, whose nuclei are bivalves, through the section to study environment change in this lithostratigraphical formation. Geochemical trace elements and δ¹³C and δ¹⁸O stable isotope analysis of the oncoids enable us to determine the conditions in which the microbialites were formed and provide further palaeoenvironmental data from the deposits containing the bivalves. Bivalve taxa change from the lower part of the Villar del Arzobispo Formation, where Ceratomya excentrica and Unicardium cf. subregulare are characteristic of marine conditions, becoming more continental towards the top of the formation with the presence of Unionoidean bivalves, and in the “Wealden” facies of the El Castellar Formation, where Teruella gautieri, has been found. Bivalves and oncoids allow us to recognize continental conditions where the first dinosaur of Spain, the sauropod Aragosaurus ischiatus was found, in an open water system, where there was limited evaporation, and with enough energy to produce well oxygenated water.