Vertebrate tracks from the Paso Córdoba fossiliferous site (Anacleto and Allen formations,Upper Cretaceous), Northern Patagonia,Argentina: Preservational,environmental and palaeobiological implications

The Paso Córdoba fossiliferous site (Río Negro, Northern Patagonia) is one of the first Mesozoic fossiliferous localities studied in Argentina. There, turtle, crocodile and dinosaur remains as well as dinosaur and bird tracks have been recorded. Recently, a new locality with vertebrate tracks, the Cañadón del Desvío, has been discovered in Paso Córdoba. Six track-bearing layers were located in outcrops belonging to the Anacleto (lower to middle Campanian, Neuquén Group) and Allen (middle Campanian-lower Maastrichtian, Malargüe Group) formations. The Cañadón del Desvío locality reveals that vertebrate trace fossils are distributed in two distinct environments, floodplains of a meandering fluvial to shallow lacustrine system and a wet interdune deposit that is associated to an aeolian setting. Also, in the logged section several soft sediment deformation structures were found. In regard of this, a sedimentary facies analysis is provided in order to assess the palaeoenvironmental implications of this new record. The analysed tracks are preserved in cross-sections, on bedding-planes and as natural casts. When it is possible, the tracking surface, true tracks, undertracks and overtracks/natural casts have been identified and the track preservation and the formation history of the tracksite are discussed. Only two tracks preserve enough anatomical details to relate them with their trackmakers, in this case hadrosaurid dinosaurs. The stratigraphical, facial and palaeoenvironmental data of this study support the idea of a transitional passage between the Anacleto and Allen Formation in Paso Córdoba. The presence of hadrosaurid dinosaur tracks suggests that the upper part of the log, where this kind of tracks were found, likely belong to Allen Formation due to this dinosaurs appear in the Southern Hemisphere in this epoch. The sum of osteological and ichnological remains improve the Paso Córdoba palaeofaunistic knowledge. The presence of six different levels in which the trackmakers walked reflects the abundance of vertebrates in the transition between Anacleto and Allen formations.     

Sedimentological and ichnological signatures of changes in wave,river and tidal influence along a Neogene tropical deltaic shoreline

Analysis of Neogene cores from the Eastern Venezuela Basin along 65 km of a west–east trending shoreline allows characterization of the sedimentological and ichnological signatures of wave, river and tidal processes. The area displays deltas prograding northward from the Guyana Shield. Twenty‐three facies are defined and grouped into four categories (wave‐influenced, river‐influenced, tide‐influenced and basinal). Wave‐dominated deltaic deposits occur mostly in the Tácata Field. The delta plain was characterized by tide‐influenced distributary channels separated by interdistributary bays. Fluvial discharge in the delta front and prodelta was repeatedly interrupted by storm‐wave reworking and suspended sediment fallout. Delta‐front and prodelta deposits contain some ichnotaxa that typically do not occur in brackish water (for example, Chondrites and Phycosiphon). Amalgamated storm deposits are unburrowed or contain vertical Ophiomorpha. Lateral (especially on the updrift side) to the river mouths, waves caused nearly continuous accretion of the associated strandplains. These deposits are the most intensely bioturbated, and are dominated by the estenohaline echinoid‐generated ichnogenus Scolicia. River‐dominated deltaic deposits are present in the Santa Bárbara, Mulata, Carito and El Furrial Fields. Low‐sinuosity rivers characterized the alluvial plain, whereas the subaerial delta plain was occupied by higher‐sinuosity rivers. The subaqueous delta plain includes distributary channels and tide‐influenced interdistributary bays. Further seaward, successions are characterized by terminal distributary‐channel and distributary mouth‐bar deposits, as well as by delta‐front and prodelta deposits showing evidence of sediment gravity‐flow and fluid‐mud emplacement. Delta‐front and prodelta deposits are unbioturbated to sparsely bioturbated, suggesting extreme stress, mostly as a result of high fluvial discharge and generation of sediment gravity flows. Tidal influence is restricted to interdistributary bays, lagoons and some distributary channels. From an ichnological perspective, and in order of decreasing stress levels, four main depositional settings are identified: river‐dominated deltas, tide‐influenced delta plains, wave‐dominated deltas and wave‐dominated strandplain–offshore complexes.     

Linking invertebrate burrow distributions (neoichnology) to physicochemical stresses on a sandy tidal flat: implications for the rock record

Invertebrate burrow distributions (neoichnology) across a modern tidal flat are presented as an analogue for interpreting the ichnology of palaeo‐tidal‐flat successions. Burrow distributions are linked to physical and chemical (physicochemical) stresses to establish the main controls on the distribution of biosedimentary structures. Across the tidal flat, there is clear heterogeneity in both the diversity of traces and the intensity of burrowing. This heterogeneity reflects a myriad of physicochemical stresses, with the sedimentation rate dominating burrow distributions. Across all substrates, the total area occupied by organisms rarely exceeds 3% of the tidal‐flat surface, and is commonly <1%; this equates to a bioturbation index value of one. To reach bioturbation index values of two to six, sediments must be available to biogenic reworking and/or recolonization. With an increasing sedimentation rate, substrates are rapidly buried and re‐exposed, which limits the time when a substrate is available to colonization. For palaeo‐ichnological studies, this research presents several key results. (i) Burrow cross‐cutting relationships in tidal‐flat successions commonly reflect natural heterogeneities in the areal distribution of infaunal communities, rather than infaunal tiering. (ii) Ichnofabric analysis of palaeo‐tidal flats with a high sedimentation rate would yield fabrics that reflect heterogeneities in the areal distribution of infaunal communities rather than variability in the physicochemical stresses of the environment. (iii) The composite trace‐fossil assemblage of tidal flats cannot be attributed to a single ichnofacies, but instead comprises elements typical of multiple ichnofacies. (iv) The main controls on trace assemblages across tidal flats in fully marine settings are sedimentological and include the sedimentation rate and, to a lesser extent, grain size.     

Bird and dinosaur footprints in the Woodbine Formation (Cenomanian), Texas

The ichnofauna of the Woodbine Formation (Cenomanian), Denton County, Texas includes bird and dinosaur tracks. A new bird trackway,Magnoavipes loweiichnogen. et ichnosp. nov., appears to represent the largest bird tracks known from the Mesozoic. A theropod trackway,Fuscinapedis woodbinensisichnogen. et ichnosp. nov., is characterized by long digits of uniform width and pointed small claws. Six hadrosaurid trackways,Caririchnium protohadrosaurichnosichnosp. nov., are the oldest hadrosaurid tracks associated with skeletal elements. They include one isolated small footprint, a medium sized quadrupedal, and five large bipedal hadrosaurid trackways.     

Bioerosion on Late Pleistocene Marine Mollusks: A Paleoclimatological and Paleoecological Comparison of MIS 7 and MIS 5e with Modern Beaches (Río Negro, Argentina)

The Late Pleistocene littoral ridges of southern South America, especially those of the north of Argentinean Patagonia, contain remains of mollusk shells with bioerosion traces. Eleven sites from marine isotope stages (2 from MIS 7, 4 from MIS 5e) and five sites from modern beaches from northern Río Negro Province, with 40 taxa (17 bivalves and 23 gastropods) were analyzed, in the area between west of Baliza San Matías and Las Grutas (41°S). Three ichnogenera were identified in deposits of MIS 7: Entobia, Gastrochaenolites (Domichnia) and Oichnus (Praedichnia) with one ichnospecies (G. torpedo). Six ichnogenera were identified in deposits of MIS 5e: Entobia, Meandropolydora, Pinaceocladichnus, Iramena, Caulostrepsis and Oichnus with one ichnospecies (O. simplex). Seven ichnogenera were identified from modern beaches: Entobia, Meandropolydora, Iramena, Caulostrepsis, Pinaceocladichnus, Gastrochaenolites (Domichnia) and Oichnus (Praedicnia), with two ichnospecies (O. simplex and O. paraboloides). On this basis, it is inferred that Pleistocene benthic communities were constituted from cheilostome bryzoans and polychaete annelids, together with drilling bivalves, predatory gastropods and overall warm-water species of mollusks. Of these, Chama iudicai was found on Plicatula gibbosa from MIS 7 and MIS 5e. The record of warm temperature species suggests a higher sea surface temperature (SST) than the present one and/or the influence of the warm southward Brazilian Current affecting the waters of Golfo San Matías. On the modern beaches surveyed, there is an increase in polychaete annelids, cheilostome bryzoans, temperate-cold species and drilling bivalves, as well as sandy substrate species, which together with an increase of Gastrochaenolites and Caulostrepsis, suggest a colder climate due to the greater influence of the Malvinas Current during the Holocene.     

遗迹相及其古环境的意义

遗迹相是指一组具有特征的遗迹化石组合,在一段相当长的地质时期中不断的重视以及它们的环境意义。本文结合中国实例指出7个常见和通用的遗迹相及其沉积环境。浊积岩或复理石沉积中含有Nereites遗迹相,文内列举了目前已知的中国10个复理石产地,证明它们主要分布在两个板块构造单元的对接（消减）带。     

Using ichnological relationships to interpret heterolithic fabrics in fluvio-tidal settings

In fluvio-tidal settings, the sediment is dominantly derived from the river systems. However, the importance of landward tidal transport of sediment in tidally influenced sedimentary environments is difficult to assess, particularly in the rock record. This problem is addressed using two intervals within the Lower Cretaceous McMurray Formation, each representing a distinct inclined heterolithic stratification motif. The ichnological variation between the heterolithic intervals is analyzed to determine which lithosomes are associated with brackish-water (tidally influenced) colonization windows. From this, the relative fluvial influence responsible for the deposition of the fine and coarse members can be determined. Both of the inclined heterolithic stratification fabrics studied record the deposition of fluvio-tidal point bars wherein the heterolithic bedding represents variations in river discharge. The first fabric comprises inclined heterolithic stratification in which bioturbation only occurs in mudstone beds. This fabric indicates that deposition occurred in more proximal positions within a fluvio-tidal system (i.e. the outermost inner to middle estuary or distributary channels). In this example sand deposition is interpreted to represent high-energy, freshwater dune migration within a fluvial-dominated setting, whereas mud beds reflect brackish-water suspension deposition during times of low river discharge. The second fabric, which is interpreted to have developed in more distal depositional positions (i.e. the middle estuary or seaward of the turbidity maximum in deltas), consists of inclined heterolithic stratification with laminated mudstone and bioturbated sandstone. In these inclined heterolithic stratification successions the mudstone beds were deposited under the influence of freshwater and heightened sedimentation rates, whereas bioturbated sandstone was colonized under brackish-water conditions and in the presence of tidally facilitated sediment transport. In both examples, the bioturbated lithosomes are related to colonization windows that indicate the predominance at that time of marine or tidally influenced processes over fluvial processes.     

中国遗迹学研究十年进展

近十年来,中国遗迹学的研究取得许多新进展和新认识,重要成果概述如下:(1)在显生宙海相和过渡相沉积中新识别出84个遗迹属和211个遗迹种,新建28种遗迹组合及26种遗迹组构;在中生代和新生代陆相冲积扇、河流与湖泊沉积中识别出44个遗迹属和107个遗迹种,新建22种遗迹组合及10种遗迹组构;(2)论述了寒武纪底质革命对早...     

Reliability of digit length impression as a character of tetrapod ichnotaxobase: considerations from the Carboniferous–Permian ichnogenus Ichniotherium

Although the relative length of digit impressions is often adopted as a binding character in ichnotaxa discrimination and trackmaker identification, it cannot be considered unequivocally operative. The reliability of this character has been evaluated by means of principal component analysis on several well‐preserved footprints (both manus and pes) referred to as Ichniotherium sphaerodactylum from the Early Permian Tambach Formation (Thuringia, Germany). The analysis reveals a substantial variability of digit impression length (especially in the manus), relativizing the validity of digit IV length as a diagnostic character as practised in different ichnological studies. Regarding the use of such a character, results clearly suggest the need to preliminarily conduct explorative analysis on the whole available ichnological material in order to detect which digit is the most consistent from an ichnotaxonomical standpoint, thus avoiding an arbitrary selection. Such a preliminary analysis should provide a better control on ichnotaxonomical splitting/lumping, particularly in the case of allied footprints, and can also be considered a valuable tool, if associated to functionality analysis, in improving and refining trackmaker identification. Copyright © 2014 John Wiley & Sons, Ltd.     

Integrated sedimentology and ichnology of Late Jurassic fluvial point‐bars – facies architecture and colonization styles (Lourinhã Formation,Lusitanian Basin,western Portugal)

Integrated sedimentological and ichnological case studies of ancient meandering river systems have, for the most part, focused on the deposits of the fluvial–tidal transition zone; much less emphasis has been placed on the purely fluvial realm above the landward limit of tidal effects. This problem needs to be addressed so that in future the defining sedimentological and ichnological criteria assigned to ancient fluvial reaches are sufficiently well‐established to enable their separation from the down‐dip fluvial–tidal transition zone. Accordingly, a case study has been carried out on a well‐exposed meander belt deposit from the Late Jurassic Lourinhã Formation of the Lusitanian Basin, western Portugal (Praia Do Valmitão, Ribamar). Analysis indicates that the meander belt here comprised mixed‐load fluvial channels traversing a vegetated floodplain subject to a seasonal winter wet/summer dry palaeoclimate. This setting facilitated the development of both calcic palaeosols and shallow lakes on the adjacent floodplain. Critically, there is no evidence of the effects of tidal modulation on bedding structures, thereby establishing purely fluvial conditions. Heterolithic point‐bar deposits generated in this setting are bioturbated extensively by a trace fossil assemblage dominated by the meniscate trace Taenidium barretti , with Skolithos linearis , Planolites beverleyensis and Cylindricum isp. also recognized. A number of factors suggest that the Taenidium barretti producer in this case was a subaquatic organism living in an active fluvial channel setting (i.e. not colonizing subaerially exposed channel‐margin/floodplain deposits). Accordingly, there are some implications for current ichnofabric/ichnofacies models in the continental realm. Firstly, Taenidium ‐dominated ichnofabrics need not necessarily be confined to colonization beneath subaerially exposed surfaces; they might also be produced within submerged substrates. Secondly, there is scope to extend the range of the Scoyenia ichnofacies to include active fluvial channels and not simply those channels that were inactive or abandoned.