Bayeritheca hughesii gen. et sp. nov., a new Eucommiidites -bearing pollen organ from the Cenomanian of Bohemia

A new pollen organ, Bayeritheca hughesii gen. et sp. nov. with in situ Eucommiidites pollen grains, is described from the Cenomanian of Bohemia in Central Europe. It is based on a single lignified specimen which is the largest Eucommiidites -producing pollen structure described so far. It shows whorls of microsporangiate units attached to a massive main axis forming a cone-like structure. Each microsporangiate unit forms distally an angular head bearing trichomes and centrally arranged mucronate tubercles. The head is attached to a short stalk. Proximal sides of heads display radially arranged synangia. Each synangium contains an unknown number of pollen sacs with numerous Eucommiidites pollen grains.Bayeritheca is the only genus of Erdtmanithecales in which synangia instead of sporangia are found. Bayeritheca differs from Eucommiiditheca Friis & Pedersen in the whorled arrangement of microsporangiate units, which are densely spaced forming a cone-like structure. It differs from Erdtmanitheca Pedersen, Crane & Friis in the elongate shape of the whole organ, slender main axis, shorter pollen sacs, and different Eucommiidites pollen.