Genetic Diversity in Marine Molluscs: A Test of the Niche-Width Variation Hypothesis

Abstract. A test of the niche width variation hypothesis was conducted in the three Mediterranean littoral species of the family Trochidae: Monodonta turbinata, M. turbiformis and Gibbula richardi. Allozymic variation encoded by 26 gene loci was analyzed, and the levels of polymorphism (P), heterozygosity (H) and allele diversity per locus (A) were estimated and appear to support the niche-width variation hypothesis. The broad-niche species, M. turbiformis and G. richardi are more polymorphic and heterozygous than the relatively narrow-niche M. turbinata (P = 0.571; 0.731 and 0.417; H = 0.108; 0.116 and 0.078; A = 2.025; 2.489 and 1.965, respectively).     

Species Abundance Distribution Patterns of a Toona ciliata Community in Xingdoushan Nature Reserve

With the goal of model fitting species abundance distribution patterns of the tree, shrub and herb layers of the natural Toona ciliata community in Xingdoushan Nature Reserve, Enshi Autonomous Prefecture, Hubei Province, we used the data collected from the field survey and employed different ecological niche models. The models tested were the broken stick model (BSM), the overlapping niche model (ONM) and the niche preemption model (NPM), as well as three statistic models, the log-series distribution model (LSD), the log-normal distribution model (LND) and the Weibull distribution model (WDM). To determine the fitted model most suitable to each layer, the fitting effects were judged by criteria of the lowest value of Akaike Information Criterion (AIC), Chi-square and the K-S values with no significant difference (P>0.05) between the theoretical predictions and observed species abundance distribution values. The result showed: (1) The fitting suitability and goodness of fit of the tree, shrub and herb layers by using the three ecological niche models were ranked as: NPM>BSM>ONM. Of the three statistical models, by accepting the fitting results of the three layers, WDM was the best fitting model, followed by LND. By rejecting the fitting tests of the herb layer, LSD had the worst fitting effect. The goodness of the statistical models was ranked as: WDM>LND>LSD. In general, the statistical models had better fitting results than the ecological models. (2) T. ciliata was the dominant species of the tree layer. The species richness and diversity of the herb layer were much higher than those of either the tree layer or the shrub layer. The species richness and diversity of the shrub layer were slightly higher than those of the tree layer. The community evenness accorded to the following order: herb>shrub>tree. Considering the fitting results of the different layers, different ecological niche models or statistical models with optimal goodness of fit and ecological significance can be given priority to in studying the species abundance distribution patterns of T. ciliata communities.     

开封市的城市生态位变化分析

城市生态位作为生态学重要理论之一,用于分析城市生态不同功能模块间竞争、演化等现象。通过以开封市不同功能模块为研究对象,在各模块污染指数和适宜度指数的基础上,运用生态位理论和方法,探讨改革开放以来,特别是近十年来开封市不同功能模块生态位格局变化规律,并对变化的原因进行了分析。研究结果表明：(1) 1994~2003年开封市自然生态环境系统各模块生态位呈上升趋势的有大气环境模块、城市绿化模块,而呈下降趋势的有水环境模块、声环境模块。(2) 1994~2003年开封市经济系统各模块生态位呈上升趋势的是第三产业模块,呈下降趋势的是第二产业模块,而变化不大的是第一产业模块。(3) 1994~2003年开封市社会系统各模块适宜度生态位呈上升趋势的有社会基础设施模块、社会资源配置模块、社会生产与保障模块,而呈下降趋势的只有人口模块。(4) 国家政策体制的改革是改革开放以来特别是近十年来开封市各功能模块生态位发生变化的最直接和最根本的原因。该研究不但对城市不同功能模块的宏观规划、经济发展和环境保护有具体的指导意义,而且对拓展城市生态学理论、研究复合生态系统的内在作用机制也同样具有重大的理论和现实意义。     

A case study on Landscape Component Niche based on Landscape Pattern Indices: Yanchi,Ningxia Province,China

Yanchi County is located in the agro-pastoral ecotone and belongs to the ecologically fragile area of Northwest China.It is important to study the evolution of landscape pattern to curb its environmental degradation.In order to intuitively show how the landscape pattern of the study area changes over time,Landsat Thematic Mappers(TM)and Landsat Operational Land Imager(OLI)data of 1991,2000,2010 and 2017 were used.This paper attempts to apply niche theories and methods into landscape ecology,and constructs a niche model of landscape components by using"n-dimentional hypervolume niche theory"and landscape pattern indices.By evaluating the spatial and temporal evolution of niche from the perspective of two-dimensional space to reflect the changes of landscape pattern in the study area over the past 26 years,new theories and methods were introduced for the characterization of landscape pattern.The results indicate that:1)The larger the attribute and dominance value of landscape components,the higher the ecological niche and the stronger the control effect on the overall landscape.2)The ecological niche of each landscape component was significantly different,just as its control effect on the overall landscape.3)The dynamic change of the ecological niche of each landscape component was different,with grassland,unused land and arable land always in a high dominant position,although the ecological niche of construction land and water area was always low.In general,the introduction of niche theory into the landscape ecology provided a new method to study the changes in regional landscape pattern.     

甘肃酒泉荒漠戈壁灌木群落优势物种生态位特征

了解物种利用资源和占据生态空间的能力,对维持完善和科学保育荒漠戈壁植物群落的多样性具有重要意义。在综合反映各生态因子作用的群落类型和海拔梯度组合而成的两条资源轴上,测度分析了甘肃酒泉荒漠戈壁灌木群落主要优势种的生态位特征。结果表明：(1)在群落类型和海拔梯度两条资源轴上,红砂（Reaumuria songarica）、泡泡刺（Nitraria sphaerocarpa）和合头草（Sympegma regelii）的重要值和生态位宽度均较大,说明这些物种适应能力强,能够较好地利用环境资源,分布范围大,作为荒漠戈壁灌木群落中的广域种具有重要的生态地位和作用。(2)荒漠戈壁优势物种间的生态位重叠值多数较小,在群落类型和海拔梯度资源轴上生态位重叠值小于0.5的分别占总种对的62.63%和77.89%。生态位宽度大的物种之间一般生态位重叠值较高,物种利用资源能力强且存在竞争关系;然而,生态位宽度较小的物种与其他物种之间的生态位重叠程度较低,不同物种在环境资源的需求上产生互补,可以和谐共存;生态位宽度小的物种之间生态位重叠值仍较高,物种分布呈斑块现象;因此,生态位重叠与生态位宽度之间无显著相关性。(3)荒漠戈壁优势物种间总体表现为不显著的正关联,表明该植被群落结构及其物种之间处于稳定共存的状态。     

中国西部生态脆弱地区景观生态位的时空差异——以陕北延安地区为例

In this study, we attempt to put forward a conception of landscape ecological niche, enlightened by international scholars on extending the ecological niche theory from spatial niche to functional niche. That is helpful for comprehensively appraising landscape spatial patterns and ecological functions, also, presents a new method for analyzing landscape features from multidimensional aspects. The practice process is demonstrated by taking Yan’an region in northwestern China as a case. Firstly, the indices system including spatial attribute and functional attribute is established for assessing landscape ecological niche. Additionally, two-dimensional figures are drawn for comparing the spatio-temporal features of landscape ecological niche in 1987 and 2000 among the 13 administrative counties. The results show that from 1987 to 2000, towards Yan’an region, spatial attribute value of landscape ecological niche changes from 1.000 to 1.178 with an obvious increment, and functional attribute value changes from 0.989 to 1.069 with a little increment, both of which enhance the regional landscape ecological niche. Towards each county, spatial attribute value of landscape ecological niche increases to different extent while functional attribute value changes dissimilarly with an increment or a decrement.     

地形位指数模型改进及其在植被覆盖评价中的应用

地形作为影响植被覆盖的重要因素,对植被恢复评价以及生态修复规划意义重大。地形位作为测度地形因子综合效应的指标,具有单一地形因子难以比拟的优势。然而,现有地形位指数算法以整个研究区为参照,未考虑空间异质性,难以反映植被生长的局部环境特征。基于地理学第三定律对地理环境相似的强调,融合“空间位置邻近”和“环境特征相似”,对传统地形位模型进行改进：(1)引入局部窗口算法,以突出局部地形特征;(2)在不同生境因子组合下测算地形位指数,以排除环境差异对地形位作用的干扰。案例研究表明,新模型有效提升了地形位指数对植被覆盖水平的解释力度;当考虑生境因子组合,且采用最优窗口时,局部窗口地形位指数与植被覆盖度之间的相关性最高。此外,通过观察不同生境组合下的地形位指数发现,在高热少水阴坡和高热少水阳坡条件下,植被覆盖度与局部窗口地形位指数具有更高相关性;可见,越是水资源相对匮乏区域,植被覆盖对局部窗口地形位的响应越敏感。本文有望为植被恢复评价与规划提供新的指标。     

Spatio-temporal differentiation of landscape ecological niche in western ecological frangible region: A case study of Yan’an region in northwestern China

In this study, we attempt to put forward a conception of landscape ecological niche, enlightened by international scholars on extending the ecological niche theory from spatial niche to functional niche. That is helpful for comprehensively appraising landscape spatial patterns and ecological functions, also, presents a new method for analyzing landscape fea-tures from multidimensional aspects. The practice process is demonstrated by taking Yan’an region in northwestern China as a case. Firstly, the indices system including spatial attribute and functional attribute is established for assessing landscape ecological niche. Additionally, two-dimensional figures are drawn for comparing the spatio-temporal features of landscape ecological niche in 1987 and 2000 among the 13 administrative counties. The results show that from 1987 to 2000, towards Yan’an region, spatial attribute value of landscape ecological niche changes from 1.000 to 1.178 with an obvious increment, and functional attribute value changes from 0.989 to 1.069 with a little increment, both of which enhance the regional landscape ecological niche. Towards each county, spatial attribute value of landscape eco-logical niche increases to different extent while functional attribute value changes dissimilarly with an increment or a decrement.     

Spatio-temporal differentiation of landscape ecological niche in western ecological frangible region： A case study of Yan＇an region in northwestern China

In this study, we attempt to put forward a conception of landscape ecological niche, enlightened by international scholars on extending the ecological niche theory from spatial niche to functional niche. That is helpful for comprehensively appraising landscape spatial patterns and ecological functions, also, presents a new method for analyzing landscape features from multidimensional aspects. The practice process is demonstrated by taking Yan＇an region in northwestern China as a case. Firstly, the indices system including spatial attribute and functional attribute is established for assessing landscape ecological niche. Additionally, two-dimensional figures are drawn for comparing the spatio-temporal features of landscape ecological niche in 1987 and 2000 among the 13 administrative counties. The results show that from 1987 to 2000, towards Yan＇an region, spatial attribute value of landscape ecological niche changes from 1.000 to 1.178 with an obvious increment, and functional attribute value changes from 0.989 to 1.069 with a little increment, both of which enhance the regional landscape ecological niche. Towards each county, spatial attribute value of landscape ecological niche increases to different extent while functional attribute value changes dissimilarly with an increment or a decrement.