Faunal dating of the late Cenozoic of southern Africa,with special reference to the Carnivora

Since chronometric data comparable to that available from the late Cenozoic succession of East Africa have not, and probably cannot be obtained in southern Africa, faunal dating methods retain their traditional significance in the latter area. Five successive late Cenozoic mammal ages, the Namibian, Langebaanian, Makapanian, Cornelian, and Florisian have been proposed as a framework for discussions relating to the chronology of southern African mammalian faunas. The Namibian fauna is poorly known, but is evidently of Miocene age. It is not discussed in this paper. The Langebaanian fauna is well known only from the prolific occurrence at the type site of Langebaanweg and is Pliocene in age. Makapanian faunas are best represented at the Transvaal Australopithecine sites and probably overlap the Plio-Pleistocene boundary. The Cornelian fauna is not as well known as others, the largest assemblage having been recorded from Elandsfontein, although this assemblage is one which is unfortunately temporally heterogeneous. Florisian faunas have been recovered from a relatively large number of localities, including several for which there are radiometric dates. The recorded southern African fossil Carnivora are listed and the local evolution of this group is discussed in terms of the mammal age chronology. It is concluded that secure faunal dating of individual fossil occurrences is enhanced by an appreciation of the nature of changes undergone in evolutionary lineages, while an uncritical knowledge of recoreded taxa is less useful.