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Phillip S. Toms 《第四纪科学杂志》2003,18(8):815-816
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Properties of the light saturation curve of photosynthesis and ribulose-1,5-bisphosphate carboxylase (RuBPC) activity are shown to change qualitatively in a natural population of marine phytoplankton during a spring bloom. Evidence is presented to show that these changes constitute photoadapative responses to increasing irradiance. As irradiance increased during the bloom, both the level of light-saturated photosynthesis (Pm) and the initial slope of the light saturation curve (α = photosynthetic efficiency) increased whether those parameters were normalized to chlorophyll a concentration (Pmb, αb) or to cell numbers (Pmc, αc). The magnitudes of these changes were such that Ik (= Pm/α, the photoadaptation parameter) did not change, but Im, the light intensity at which photosynthesis becomes saturated, increased. RuBPC activity, both chlorophyll a (RuBPCb) and cell number normalized (RuBPCc), also increased during the bloom. We suggest that these adaptations were achieved by simultaneously increasing the number of photosynthetic units, proportionately decreasing the photosynthetic unit size, and increasing both the concentrations of the enzymes of the dark reactions and possibly also of photosynthetic electron transport components.We also observed diminished levels of photoinhibition in the high light adapted cells late in the bloom and have suggested that this was a consequence of the same suite of physiological changes.In situ carbon fixation per cell increased during the bloom whereas no change occurred in this parameter when normalized to chlorophyll a concentration. Although these photoadaptive responses thus permitted carbon to be fixed in situ more rapidly per cell, at a constant efficiency with respect to investment of energy in the photosynthetic apparatus, they did not result in a change in growth rate. Based on consideratios of the role of time scale in physiological adaptation, however, it is suggested that the observed alterations in photosynthesis with increasing irradiance might permit a cell to more rapidly fill an energy quota for division, possibly an advantage in a mixing environment in which energy is patchily distributed, both spatially and temporalyy.Phosphoenolpyruvate carboxylase activity when normalized to chlorophyll a (PEPCb) did not change during the bloom while chlorophyll a normalized dark carbon fixation decreased sharply and was quantitatively small compared to PEPCb. On this basis and considering that RuBPCb increased during the bloom, it is suggested that, although PEPC may be involved in dark carbon fixation, its most important quantitative role is probably an indirect one in light dependent photosynthesis.We have also considered the relevance of laboratory results on photoadaptation to interpretations of field studies and have suggested that batch culture studies must be treated with caution but that turbidistat and semi-continuous methods provide reasonable simulations of natural conditions. 相似文献
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In this paper, we formulate a finite-element procedure for approximating the coupled fluid and mechanics in Biot’s consolidation
model of poroelasticity. We approximate the flow variables by a mixed finite-element space and the displacement by a family
of discontinuous Galerkin methods. Theoretical convergence error estimates are derived and, in particular, are shown to be
independent of the constrained specific storage coefficient, c
o
. This suggests that our proposed algorithm is a potentially effective way to combat locking, or the nonphysical pressure
oscillations, which sometimes arise in numerical algorithms for poroelasticity. 相似文献
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Surface accumulations of foam and flotsam as well as sharp salinity, density, turbidity gradients and regions of acoustic scatter were characteristic of ebb-tidal fronts in Charleston Harbor, South Carolina. Surface convergence velocities at these fronts averaged 0.06 m s?1 into the front at an angle of 30° to 60° with respect to the frontal axis, indicating along-front transport during the ebb. These fronts are tidally-induced, forming on the late flood and ebb along the interfaces of water masses. Horizontal and vertical measurements of density revealed that the upper harbor fronts form along the margin of a freshwater lens produced by riverine input. The hypothesis that these frontal zones have higher densities of phytoplankton and zooplankton than adjacent water masses was tested using chlorophylla measurements and net collections. The fronts did not demonstrate any significant accumulations of phytoplankton or zooplankton during the ebb tide. The results of this study suggest that the physical characteristics of ebb-tidal estuarine fronts in Charleston Harbor are periodic in nature and may indirectly affect plankton transport in this coastal plain estuary. 相似文献
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Heidi M. Cullen Alexey Kaplan Phillip A. Arkin Peter B. deMenocal 《Climatic change》2002,55(3):315-338
Interannual to decadal variations in Middle Eastern temperature, precipitationand streamflow reflect the far-field influence of the North Atlantic Oscillation (NAO), a dominant mode of Atlantic sector climate variability. Using a new sea surface temperature (SST) based index of the NAO and availablestreamflow data from five Middle Eastern rivers, we show that the first principal component of December through March streamflow variability reflects changes in the NAO. However, Middle East rivers have two primary flooding periods.The first is rainfall-driven runoff from December through March, regulated on interannual to decadal timescales by the NAO as reflected in local precipitation and temperature.The second period, from April through June, reflects spring snowmelt and contributes in excess of 50% of annual runoff.This period, known locally as the khamsin, displays no significantNAO connections and a less direct relationship with local climatic factors, suggesting that streamflow variability during this period reflects land-cover change, possibly related to agriculture and hydropower generation, and snowmelt. 相似文献
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Seagrass meadows are often cited as important nursery areas for newly settled red drum even though many estuaries, such as Galveston Bay, Texas, support large numbers of red drum and have limited seagrass cover, suggesting the use of alternate nursery areas. We examined patterns of habitat use for newly settled red drum at six sampling areas in Galveston Bay; two areas had seagrass beds and four areas had no seagrass. We measured densities in different habitat types using epibenthic sleds and enclosure samplers. Peak recruitment of young red drum to the estuary occurred during September through December. Highest densities of new settlers were found in seagrass meadows (primarilyHalodule wrightii), but when seagrass was absent, the highest densities of red drum occurred along theSpartina alterniflora marsh edge interface. Densities were relatively low on nonvegetated bottom away from the marsh edge. We also examined density patterns in other habitat types at selected sampling areas and found no red drum within marsh vegetation away from the marsh edge interface (5 and 10 m into the marsh interior). Oyster reefCrassostrea virginica was sampled using lift nets, and we found no red drum using this habitat, although adjacent seagrass and marsh interface habitats were used. Even though red drum densities in marsh edge were low relative to seagrass, the large areal extent of marshes in the bay complex probably makes marsh edge the most important nursery habitat for red drum in Galveston Bay. 相似文献
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Since their post-war inception, Sydney's metropolitan plans have tended to be overtaken by the social, economic and environmental conditions they have had to confront. The depth and scope of Sydney's recent urban transformation threatens again to overtake metropolitan planning capacity creating, in the context of competitive globalisation, a potentially significant market disadvantage for the city, not to mention poor urban development outcomes. This paper reviews Sydney's post-war metropolitan planning strategies, examining the social and economic contexts and the policy paradigms in which they have been framed, in order to draw out the lessons to be learned from their successes and failures. We argue that future success in planning urban development will rely on richly informed and fine-grained understanding of the complex spatial outcomes of Sydney's ever-deeper global integration. Only such fine-grained understanding can empower metropolitan planning to be responsive to the evolving challenges of managing development in the contemporary urban context. 相似文献