First Record of Fossil Mesocupes from China （Coleoptera： Archostemata： Cupedidae）

Three new fossil species of the genus Mesocupes of fossil cupedids,M.angustilabialis sp. nov.,M.latilabialis sp.nov.and M.collaris sp.nov.,are described from the Jiulongshan Formation of Inner Mongolia,China.These new species are the first three records of fossil Mesocupes in China as well as the oldest ones in the world now.This finding also extends the geographical distribution of this genus from Central to East Asia.In addition,based on the ratio of the length of the last to the penultimate abdominal ventrites of new beetles distinctly lower than that of the species from the Karabastau Formation,the age of Daohugou fossil-bearing beds might be older than that of Karatau assemblage and consequently of Middle Jurassic.     

A New Genus and Species of Damsel-Dragonfly (Odonata: Isophlebioidea: Campterophlebiidae) in the Middle Jurassic of Inner Mongolia, China

The campterophlebiid new genus and species Ctenogampsophlebia reni is described from the Middle Jurassic of Inner Mongolia, China. It shows close similarities with the Lower to Middle Jurassic genera Gampsophlebia, and Petrophlebia, with closed and short subdiscoidal cells, confirming the attribution of these two other genera to the Campterophlebiidae.     

A New Species of Liadotaulius (Insecta: Trichoptera) from the Middle Jurassic of Daohugou,Inner Mongolia

Diverse caddis flies are recently collected from the Middle Jurassic Daohugou fauna in Ningcheng County, Inner Mongolia, northern China, and different types of caddis cases have been reported in the same locality. A new species of caddisfly, Liadotaulius daohugouensis sp. nov. is described and illustrated; the establishment is based on a well-preserved isolated forewing collected from the Middle Jurassic Jiulongshan Formation of Daohugou. The family status of Liadotaulius is discussed on the basis of new material and is referred to the family Philopotamidae. The Daohugou fauna continues to provide new data for studying the early evolution of Trichoptera.     

First boganiine beetle in mid-Cretaceous amber from northern Myanmar (Coleoptera: Boganiidae)

Cretoboganium gei gen. et sp. nov., a new amber inclusion of the cucujoid family Boganiidae is described and figured based on a well-preserved adult from the mid-Cretaceous Burmese amber (Hukawng Valley, northern Myanmar), some 99 million years ago. Based on the presence of a pair of pronotal callosities, Cretoboganium can be firmly placed in the extant subfamily Boganiinae, a small group currently comprising two small austral genera. Our discovery represents the first fossil record for Boganiinae. It also demonstrates another example that an apparently austral group may have its sister group occurred in today’s northern hemisphere. Together with the other fossil boganiid known from the Middle Jurassic of China, the finding suggests that Boganiidae is an ancient and relict group. Moreover, the present biogeographic distribution of Boganiinae is indicative of an earlier origin of this subfamily, which likely originated before the breakup of the Gondwanan supercontinent.     

Weiwoboidae fam.nov.of 'Higher' Fulgoroidea (Hemiptera:Fulgoromorpha) from the Eocene Deposits of Yunnan,China

<正>A new family of so-called‘higher' planthoppers,Weiwoboidae fam.nov.,from the Lower Eocene of Yunnan is described.A new monotypic genus,Weiwoboa gen.nov.,with Weiwoboa meridiana sp.nov.is also described and illustrated.The characters of tegmen venation of the new family,its evolution,and supposed relationships are briefly discussed.     

A New Family of Moths from the Middle Jurassic (Insecta: Lepidoptera)

<正>Three lepidopteran species,from the Middle Jurassic Daohugou beds(inner Mongolia,China),are described in a new family,Mesokristenseniidae,and new genus,Mesokristensenia,which could represent the sister group of the Micropterigidae.Mesokristensenia differs from all extant Lepidoptera,but one genus(Agathiphaga, Agathiphagidae),in retaining four median veins in the forewing,a plesiomorphy also present in many Trichoptera. Evidence for placing Mesokristensenia in the Lepidoptera includes four traits,notably a previously unrecorded autapomorphy of this insect order:beyond stem M1+2,vein M1 is bent and connected to cross-vein r-m(in both wing pairs).Among 24 characters taken into account to assess the systematic position of Mesokristensenia,12 are considered informative for a cladistic analysis involving this fossil taxon and the four suborders recognized in present-day Lepidoptera(Zeugloptera,Aglossata,Heterobathmiina,and Glossata).     

Jurassic integrative stratigraphy and timescale of China

The Jurassic stratigraphy in China is dominated by continental sediments. Marine facies and marine-terrigenous facies sediment have developed locally in the Qinghai-Tibet area, southern South China, and northeast China. The division of terrestrial Jurassic strata has been argued, and the conclusions of biostratigraphy and isotope chronology have been inconsistent.During the Jurassic period, the North China Plate, South China Plate, and Tarim Plate were spliced and formed the prototype of ancient China. The Yanshan Movement has had a profound influence on the eastern and northern regions of China and has formed an important regional unconformity. The Triassic-Jurassic boundary(201.3 Ma) is located roughly between the Haojiagou Formation and the Badaowan Formation in the Junggar Basin, and between the Xujiahe Formation and the Ziliujing Formation in the Sichuan Basin. The early Early Jurassic sediments generally were lacking in the eastern and central regions north of the ancient Dabie Mountains, suggesting that a clear uplift occurred in the eastern part of China during the Late Triassic period when it formed vast mountains and plateaus. A series of molasse-volcanic rock-coal strata developed in the northern margin of North China Craton in the Early Jurassic and are found in the Xingshikou Formation, Nandailing Formation, and Yaopo Formation in the West Beijing Basin. The geological age and markers of the boundary between the Yongfeng Stage and Liuhuanggou Stage are unclear. About 170 Ma ago, the Yanshan Movement began to affect China. The structural system of China changed from the near east-west Tethys or the Ancient Asia Ocean tectonic domain to the north-north-east Pacific tectonic domain since 170–135 Ma. A set of syngenetic conglomerate at the bottom of the Haifanggou or Longmen Fms. represented another set of molasse-volcanic rock-coal strata formed in the Yanliao region during the Middle Jurassic Yanshan Movement(Curtain A1). The bottom of the conglomerate is approximately equivalent to the boundary of the Shihezi Stage and Liuhuanggou Stage. The bottom of the Manas Stage creates a regional unconformity in northern China(about 161 Ma, Volcanic Curtain of the Yanshan Movement, Curtain A2). The Jurassic Yanshan Movement is likely related to the southward subduction of the Siberian Plate to the closure of the Mongolia-Okhotsk Ocean. A large-scale volcanic activity occurred in the Tiaojishan period around 161–153 Ma. Note that 153 Ma is the age of the bottom Tuchengzi Formation, and the bottom boundary of the Fifth Stage of the Jurassic terrestrial stage in China should have occurred earlier than this. This activity was marked by a warming event at the top of the Toutunhe Formation, and the change in the biological assembly is estimated to be 155 Ma. The terrestrial Jurassic-Cretaceous boundary(ca. 145.0 Ma) in the Yanliao region should be located in the upper part of Member 1 of the Tuchengzi Formation, the Ordos Basin in the upper part of the Anding Formation, the Junggar Basin in the upper part of the Qigu Formation, and the Sichuan Basin in the upper part of the Suining Formation The general characteristics of terrestrial Jurassic of China changed from the warm and humid coal-forming environment of the Early-Middle Jurassic to the hot, dry, red layers in the Late Jurassic. With the origin and development of the Coniopteris-Phoenicopsis flora, the Yanliao biota was developed and spread widely in the area north of the ancient Kunlun Mountains, ancient Qinling Mountains, and ancient Dabie Mountain ranges in the Middle Jurassic, and reached its great prosperity in the Early Late Jurassic and gradually declined and disappeared and moved southward with the arrival of a dry and hot climate.     

A New Elaterid from the Middle Jurassic Daohugou Biota (Coleoptera: Elateridae: Protagrypninae)

A new genus and species of fossil elaterid namely Clavelater ningchengensis gen. et sp. nov. assigned to the subfamily Protagrypninae based on one specimen from the Middle Jurassic Jiulongshan Formation near Daohugou Village, Ningcheng County, Inner Mongolia, northeastern China is described and illustrated. This specimen displays a pair of large antennae that easily differs from those of most Mesozoic elaterids. There are around 36 species within 20 genera referred to elateroids described from the Early Jurassic to Miocene of China, with another three species within three genera reviewed, but the taxonomic assignment of at least seven species within three genera is doubted. The systematic position of fossil elateriform beetles reported from China is reviewed and the evolution of Mesozoic elateroids briefly discussed.