Modelling the effect of low soil temperatures on transpiration by Scots pine

For ecosystem modelling of the Boreal forest it is important to include processes associated with low soil temperature during spring‐early summer, as these affect the tree water uptake. The COUP model, a physically based SVAT model, was tested with 2 years of soil and snow physical measurements and sap flow measurements in a 70‐year‐old Scots pine stand in the boreal zone of northern Sweden. During the first year the extent and duration of soil frost was manipulated in the field. The model was successful in reproducing the timing of the soil warming after the snowmelt and frost thaw. A delayed soil warming, into the growing season, severely reduced the transpiration. We demonstrated the potential for considerable overestimation of transpiration by the model if the reduction of the trees' capacity to transpire due to low soil temperatures is not taken into account. We also demonstrated that the accumulated effect of aboveground conditions could be included when simulating the relationship between soil temperature and tree water uptake. This improved the estimated transpiration for the control plot and when soil warming was delayed into the growing season. The study illustrates the need of including antecedent conditions on root growth in the model in order to catch these effects on transpiration. The COUP model is a promising tool for predicting transpiration in high‐latitude stands. Copyright © 2006 John Wiley & Sons, Ltd.     

Inter-annual variability in larval supply to populations of three invertebrate taxa in the northern California Current

We investigated sources of inter-annual variability in larval supply to crab and sea urchin populations at Bodega Head and Point Reyes in northern California. During the spring and summer upwelling seasons of the years 1992 through 1997 we monitored the weekly settlement rates of nine species of crabs and two species of sea urchins. As observed in previous studies, daily values of alongshore windstress, temperature and salinity provided evidence for the poleward flow of relatively warm, low salinity water from south of Point Reyes, an apparent retention zone, during upwelling relaxation events. In years dominated by these events (1992, 1993, 1995 and 1996) we observed that alongshore windstress, temperature and salinity were coherent and temperature was significantly correlated with cancrid crab settlement. During these years the magnitude of cancrid crab settlement and the fraction of cancrid crabs relative to other crab species settling were high. Over four years of concurrent sampling there was consistently greater cancrid crab settlement at the Point Reyes site, within the retention zone, than at Bodega Head. Settlement of non-cancrid crabs (porcellanids, grapsids, pagurids and majids) was not as closely linked to intra-annual patterns of upwelling and relaxation, possibly due to the shorter seasonal availability of larvae allowing for the influence of fewer relaxation events. Settlement of this group among years was positively correlated with environmental indicators of strong seasonal upwelling; high salinity, Bakun upwelling index and low temperature. Sea urchin settlement events were observed in June and July of 1992, 1994 and 1997 during warming periods when salinity and temperature were increasing and alongshore windstress was low. Across the six years of the study, we found that cancrid crab larvae had a more even seasonal availability than larvae of non-cancrid species, which settled in greatest numbers during the early portion of the upwelling season. Sea urchins settled in greatest numbers during the later part of the upwelling season. Together these patterns demonstrate the taxon-specific way that inter-annual variability in larval supply is forced by the coincidence of larval availability with favorable physical transport mechanisms.     

Colonization, succession, and nutrition of macrobenthic assemblages in a restored wetland at Tijuana Estuary, California

Modes of colonization, the successional trajectory, and trophic recovery of a macrofaunal community were analyzed over 19 months in the Friendship marsh, a 20-acre restored wetland in Tijuana Estuary, California. Traditional techniques for quantifying macrofaunal communities were combined with emerging stable isotopic approaches for evaluation of trophic recovery, making comparisons with a nearby natural Spartina foliosa habitat. Life history-based predictions successfully identified major colonization modes, although most taxa employed a variety of tactics for colonizing the restored marsh. The presence of S. foliosa did not seem to affect macrofaunal colonization or succession at the scale of this study. However, soil organic matter content in the restored marsh was positively correlated with insect densities, and high initial salinities may have limited the success of early colonists. Total macrofaunal densities recovered to natural marsh levels after 14 months and diversity, measured as species richness and the Shannon index (H′), was comparable to the natural marsh by 19 months. Some compositional disparities between the natural and created communities persisted after 19 months, including lower percentages of surface-feeding polychaetes (Polydora spp.) and higher percentages of dipteran insects and turbellarians in the Friendship marsh. As surficial structural similarity of infaunal communities between the Friendship and natural habitat was achieved, isotopic analyses revealed a simultaneous trajectory towards recovery of trophic structure. Enriched δ¹³C signatures of benthic microalgae and infauna, observed in the restored marsh shortly after establishment compared to natural Spartina habitat, recovered after 19 months. However, the depletion in δ¹⁵N signatures of macrofauna in the Friendship marsh indicated consumption of microalgae, particularly nitrogen-fixing cyanobacteria, while macroalgae and Spartina made a larger contribution to macrofaunal diets in the natural habitat. Future successional studies must continue to develop and employ novel combinations of techniques for evaluating structural and functional recovery of disturbed and created habitats.     

Dispersal, Genetic Differentiation and Speciation in Estuarine Organisms

For some of their occupants, estuaries represent spatially discrete habitats, isolated from each other by barriers to dispersal or physiological tolerance. We present contrasting strategies for the retention or export of larvae from their estuary of origin, and consider the implications these have on population structure and divergence. Reported patterns of genetic differentiation and inferred gene flow in estuarine taxa (principally animals) are reviewed, and difficulties in the interpretation of existing genetic data discussed. Species concepts and models of speciation relevant to estuaries are outlined, and patterns of speciation of estuarine taxa reviewed. It is concluded that estuarine environments tend to restrict gene flow and impose distinct selective regimes, generating physiologically adapted populations divergent from their marine counterparts, and the potential for in situ speciation in complete or partial isolation. The resulting taxa may represent sibling or cryptic species groups of truly estuarine origin, rather than simply estuarine populations of marine eurytopes.     

Government involvement in New Zealand tourism: A public choice perspective

The theoretical model of public choice is applied to the New Zealand tourism situation in order to evaluate the role of central government in providing tourism-related goods, services and amenities. Public choice theory provides a number of rationales for government involvement in the provision of certain goods, as well as explanations where government roles exceed what would normally be expected using the theory. A framework is developed for assessing the role of government in tourism generally. This framework is then applied specifically to the New Zealand situation. Some preliminary conclusions are drawn about the utility of public choice in analysing the provision of tourism-related goods.     

Spectral Simulation and Conditioning to Local Continuity Trends in Geologic Modeling

Spectral simulation has gained application in building geologic models due to the advantage of better honoring the spatial continuity of petrophysical properties, such as reservoir porosity and shale volume. Distinct from sequential simulation methods, spectral simulation is a global algorithm in the sense that a global density spectrum is calculated once and the inverse Fourier transform is performed on the Fourier coefficient also only once to generate a simulation realization. The generated realizations honor the spatial continuity structure globally over the whole field instead of only within a search neighborhood, as with sequential simulation algorithms. However, the disadvantage of global spectral simulation is that it traditionally cannot account for the local information such as the local continuity trends, which are often observed in reservoirs and hence are important to be accounted for in geologic models. This disadvantage has limited wider application of spectral simulation in building geologic models. In this paper, we present ways of conditioning geologic models to the relevant local information. To account for the local continuity trends, we first scale different frequency components of the original model with local-amplitude spectrum ratios that are specific to the local trend. The sum of these scaled frequency components renders a new model that displays the desired local continuity trend. The implementation details of this new method are discussed and examples are provided to illustrate the algorithm.     

Preliminary investigation of a late Wisconsinan fauna from K1 cave, Queen Charlotte Islands (Haida Gwaii), Canada

Recent investigations of a limestone solution cave on the Queen Charlotte Islands (Haida Gwaii) have yielded skeletal remains of fauna including late Pleistocene and early Holocene bears, one specimen of which dates to ca. 14,400 ¹⁴C yr B.P. This new fossil evidence sheds light on early postglacial environmental conditions in this archipelago, with implications for the timing of early human migration into the Americas.     

Knickpoint recession rate and catchment area: the case of uplifted rivers in Eastern Scotland

Knickpoint behaviour is a key to understanding both the landscape responses to a base‐level fall and the corresponding sediment fluxes from rejuvenated catchments, and must be accommodated in numerical models of large‐scale landscape evolution. Knickpoint recession in streams draining to glacio‐isostatically uplifted shorelines in eastern Scotland is used to assess whether knickpoint recession is a function of discharge (here represented by its surrogate, catchment area). Knickpoints are identified using DS plots (log slope versus log downstream distance). A statistically significant power relationship is found between distance of headward recession and catchment area. Such knickpoint recession data may be used to determine the values of m and n in the stream power law, E = KA^mSⁿ. The data have too many uncertainties, however, to judge definitively whether they are consistent with m = n = 1 (bedrock erosion is proportional to stream power and KPs should be maintained and propagate headwards) or m = 0·3, n = 0·7 (bedrock incision is proportional to shear stress and KPs do not propagate but degrade in place by rotation or replacement). Nonetheless, the E Scotland m and n values point to the dominance of catchment area (discharge) in determining knickpoint retreat rates and are therefore more consistent with the stream power law formulation in which bedrock erosion is proportional to stream power. Copyright © 2005 John Wiley & Sons, Ltd.     

Hominins without fellow travellers? First appearances and inferred dispersals of Afro-Eurasian large-mammals in the Plio-Pleistocene

Discoveries of fossil Homo outside Africa predating 1.0 Ma have generated much discussion about hominin dispersal routes. However, tool-using bipeds were only one element of the inter-continental mammalian dispersals that occurred during the climatic changes of the Pliocene and Pleistocene. This paper will place hominin movements in the context of those of the wider mammalian fauna, which includes carnivores, bovids and non-human primates. The distribution of these different taxa suggests that species moved individually when the environmental conditions were right for them, rather than in multi-species waves of dispersal, and allows evaluation of the contextual evidence for the newly emerging ‘Out of Asia’ paradigm as well as the established ‘Out of Africa’ model.     

The exponential decline in saturated hydraulic conductivity with depth: a novel method for exploring its effect on water flow paths and transit time distribution

The strong vertical gradient in soil and subsoil saturated hydraulic conductivity is characteristic feature of the hydrology of catchments. Despite the potential importance of these strong gradients, they have proven difficult to model using robust physically based schemes. This has hampered the testing of hypotheses about the implications of such vertical gradients for subsurface flow paths, residence times and transit time distribution. Here we present a general semi‐analytical solution for the simulation of 2D steady‐state saturated‐unsaturated flow in hillslopes with saturated hydraulic conductivity that declines exponentially with depth. The grid‐free solution satisfies mass balance exactly over the entire saturated and unsaturated zones. The new method provides continuous solutions for head, flow and velocity in both saturated and unsaturated zones without any interpolation process as is common in discrete numerical schemes. This solution efficiently generates flow pathlines and transit time distributions in hillslopes with the assumption of depth‐varying saturated hydraulic conductivity. The model outputs reveal the pronounced effect that changing the strength of the exponential decline in saturated hydraulic conductivity has on the flow pathlines, residence time and transit time distribution. This new steady‐state model may be useful to others for posing hypotheses about how different depth functions for hydraulic conductivity influence catchment hydrological response. Copyright © 2016 John Wiley & Sons, Ltd.