Coupling of life cycles of the copepods Calanus chilensis and Centropages brachiatus to upwelling induced variability in the central-southern region of Chile

A time series of zooplankton sampling carried out at Station 18 off Concepción (36°S, 73°W) from August 2002 to December 2003 allowed the study of annual life cycles of the copepods Calanus chilensis and Centropages brachiatus in association with environmental variability in the coastal upwelling zone. Changes in the abundance of eggs, nauplii, and copepodids were assessed from samples taken at a mean time interval of ca. 20 days. Upwelling variability in near-surface waters was reflected in seasonal changes in salinity, water column stratification, and oxycline depth, as well as a weak seasonal signal in sea surface temperature (1-2 °C). Both copepods exhibited similar life cycles, characterized by continuous reproduction throughout the year. Estimates of generation times, as a function of temperature, were 25-30 days for C. chilensis and 27-35 days for C. brachiatus, predicting about 12 and 10 generations a year, respectively. These estimates were consistent with reproduction pulses observed in the field. It was thus suggested that copepods may grow under non-limiting food conditions in this upwelling area. However, despite continuous reproduction, there were abrupt changes in population sizes along with the disappearance of early naupliar and copepodid stages taking place even during the upwelling season (spring/summer). These changes were attributed to sudden increases in mortality taking place in spring or early summer, after which the populations remained at low levels through the fall and winter. It is thus suggested that, in addition to variability in the physical environment, biological interactions modulating changes in copepod mortality should be considered for understanding copepod life cycles in highly productive upwelling systems.     

Toward a generic method for studying water renewal, with application to the epilimnion of Lake Tanganyika

We present a method, based on the concept of age and residence time, to study the water renewal in a semi-enclosed domain. We split the water of this domain into different water types. The initial water is the water initially present in the semi-enclosed domain. The renewing water is defined as the water entering the domain of interest. Several renewing water types may be considered depending on their origin. We present the equations for computing the age and the residence time of a certain water type. These timescales are of use to understand the rate at which the water renewal takes place. Computing these timescales can be achieved at an acceptable extra computer cost.The above-mentioned method is applied to study the renewal of epilimnion (i.e. the surface layer) water in Lake Tanganyika. We have built a finite element reduced-gravity model modified to take into account the water exchange between the epilimnion and the hypolimnion (i.e. the bottom layer), the water supply from precipitation and incoming rivers, and the water loss from evaporation and the only outgoing river. With our water renewal diagnoses, we show that the only significant process in the renewal of epilimnion water in Lake Tanganyika is the water exchange between the epilimnion and the hypolimnion, other phenomena being negligible.     

Genetic Relationships Among One Non-Endemic and Two Endemic Mediterranean Triplefin Blennies (Pisces, Blennioidei)

Abstract. Three triplefin blennies occur sympatrically in the Mediterranean Sea; Tripterygion tripteronotus and T. melanurus are endemic, whereas T. delaisi is also found in the Eastern Atlantic. Although very similar in morphology, ecology and behaviour, some striking differences exist among reproductive strategies. Several authors proposed hypotheses on the evolution of these species. In order to enhance insight into the genetic structure of this genus, an enzyme electrophoretic survey was performed. Samples of 40 to 45 specimens of each species were collected near Calvi (Corsica, France). Twenty-four enzyme loci were analysed, of which 17 appeared polymorphic. T. delaisi showed a high level of average observed heterozygosity (H_o) of 0.140, as opposed to very low levels of 0.009 and 0.021 in T. tripteronotus and T. melanurus , respectively. Nei's genetic distance was 0.21 between T. delaisi and T. tripteronotus , 0.35 between T. delaisi and T. melanurus and 0.45 between T. tripteronotus and T. melanurus . We hypothesise that divergence of these species started before the Pleistocene and that the endemic species survived the Pleistocene glaciations in refugia within the Mediterranean. The possibility that T. tripteronotus and T. delaisi diverged through sympatric speciation is discussed.     

青岛文昌鱼的生活史──年龄、生长和死亡研究

于１９８９年８月－１９９０年７月对青岛文昌鱼的生活史进行研究。结果表明：（１）青岛文昌鱼种群由４个齿组组成，它至少生产４ａ以上，甚至可达６ａ左右。（２）在Ⅰ、Ⅱ齿时生长较快，平均年生长１３－１５ｍｍ，Ⅲ龄后生长速度减慢，即使同一年龄组的文晶鱼，在不同季节其生长速度也不同。（３）在冬春采集的文昌鱼中Ⅲ龄组的峰值较小。（４）繁殖季节过后，老龄文昌鱼因衰老而死亡。     

A phase-averaged Boussinesq model with effective description of carrier wave group and associated long wave evolution

From the phase-resolving improved Boussinesq equations (Beji and Nadaoka, Ocean Engineering 23 (1996) 691), a phase-averaged Boussinesq model for water waves is derived by more effectively describing carrier wave groups and accompanying long wave evolution with less CPU time. Linear shoaling characteristics of carrier wave equations are investigated and found to agree exactly with the analytical expression obtained from the constancy of energy flux for the improved Boussinesq equations themselves, showing that the present model equations are the results of a consistent derivation procedure regarding energy considerations. Numerical simulations of the derived equations for the single wave group and narrow-banded random waves show the validity of the present model and its high performance, especially on the CPU time.     

An experimental method for determining the frequency-dependent added mass and added mass moment of inertia for a floating body in heave and pitch motions

Most of the existing relevant materials have been obtained from experiments, in which evaluating the added mass at the resonant frequency corresponding to the peak of a frequency-response curve obtained from the “forced” vibration analysis is the most popular technique. In this paper, a simple experimental method was presented where the “free” vibration responses instead of the “forced” ones were used to determine the values of m_ah and I_ap. The main part of the experimental system is composed of a floating body (model) and a spring–shaft shaker. The “free” vibration of this main part was induced by imposing on it an initial displacement (and/or an initial velocity), and from the time histories of displacements information such as the “damped” natural frequencies, damping ratios, sectional added mass coefficients (C_V and C_P) were obtained. Since the displacements of the spring–shaft shaker are “translational” and those of the floating body due to pitch motions are “angular”, a technique for the transformation between the associated parameters of the two components of the main part was presented.     

Surface Elevation Distribution of Sea Waves Based on the Maximum Entropy Principle

A probability density function of surface elevation is obtained through improvement of the method introduced by Cieslikiewicz who employed the maximum entropy principle to investigate the surface elevation distribution. The density function can be easily extended to higher order according to demand and is non-negative everywhere, satisfying the basic behavior of the probability. Moreover because the distribution is derived without any assumption about sea waves, it is found from comparison with several accepted distributions that the new form of distribution can be applied in a wider range of wave conditions. In addition, the density function can be used to fit some observed distributions of surface vertical acceleration although something remains unsolved.     

Hydrodynamic modeling of flushing time in a small estuary of North Bay, Florida, USA

Freshwater fraction method is popular for cost-effective estimations of estuarine flushing time in response to freshwater inputs. However, due to the spatial variations of salinity, it is usually expensive to directly estimate the long-term freshwater fraction in the estuary from field observations. This paper presents the application of the 3D hydrodynamic model to estimate the distributions of salinity and thus the freshwater fractions for flushing time estimation. For a case study in a small estuary of the North Bay in Florida, USA, the hydrodynamic model was calibrated and verified using available field observations. Freshwater fractions in the estuary were determined by integrating freshwater fractions in model grids for the calculation of flushing time. The flushing time in the North Bay is calculated by the volume of freshwater fraction divided by the freshwater inflow, which is about 2.2 days under averaged flow conditions. Based on model simulations for a time series of freshwater inputs over a 2-year period, a power regression equation has been derived from model simulations to correlate estuarine flushing time to freshwater inputs. For freshwater input varying from 12 m³/s to 50 m³/s, flushing time in this small estuary of North Bay changes from 3.7 days to 1.8 days. In supporting estuarine management, the model can be used to examine the effects of upstream freshwater withdraw on estuarine salinity and flushing time.     

近58a来影响和登陆浙江的热带气旋统计特征分析

本文利用1949-2006年热带气旋资料,对58a来影响和登陆浙江热带气旋的登陆位置、登陆时间的年际、月际和日变化、登陆后的移速及路径变化与降水中心分布的关系、登陆后维持和衰减情况等进行了统计分析.统计结果表明：影响和登陆浙江的热带气旋有较明显的年际、月际和日变化特征;登陆浙江的热带气旋陆上维持时间与登陆时的强度正相关,登陆后12h内热带气旋衰减较快;登陆后移向偏北分量大的或移速加快的热带气旋,主要降水区出现在路径右侧的可能性大.     

Correspondence between the distribution of hydrodynamic time parameters and the distribution of biological and chemical variables in a semi-enclosed coral reef lagoon

Hydrodynamic modeling can be used to spatially characterize water renewal rates in coastal ecosystems. Using a hydrodynamic model implemented over the semi-enclosed Southwest coral lagoon of New Caledonia, a recent study computed the flushing lag as the minimum time required for a particle coming from outside the lagoon (open ocean) to reach a specific station [Jouon, A., Douillet, P., Ouillon, S., Fraunié, P., 2006. Calculations of hydrodynamic time parameters in a semi-opened coastal zone using a 3D hydrodynamic model. Continental Shelf Research 26, 1395–1415]. Local e-flushing time was calculated as the time requested to reach a local grid mesh concentration of 1/e from the precedent step. Here we present an attempt to connect physical forcing to biogeochemical functioning of this coastal ecosystem. An array of stations, located in the lagoonal channel as well as in several bays under anthropogenic influence, was sampled during three cruises. We then tested the statistical relationships between the distribution of flushing indices and those of biological and chemical variables. Among the variables tested, silicate, chlorophyll a and bacterial biomass production present the highest correlations with flushing indices. Correlations are higher with local e-flushing times than with flushing lags or the sum of these two indices. In the bays, these variables often deviate from the relationships determined in the main lagoon channel. In the three bays receiving significant riverine inputs, silicate is well above the regression line, whereas data from the bay receiving almost insignificant freshwater inputs generally fit the lagoon channel regressions. Moreover, in the three bays receiving important urban and industrial effluents, chlorophyll a and bacterial production of biomass generally display values exceeding the lagoon channel regression trends whereas in the bay under moderate anthropogenic influence values follow the regressions obtained in the lagoon channel. The South West lagoon of New Caledonia can hence be viewed as a coastal mesotrophic ecosystem that is flushed by oligotrophic oceanic waters which subsequently replace the lagoonal waters with water considerably impoverished in resources for microbial growth. This flushing was high enough during the periods of study to influence the distribution of phytoplankton biomass, bacterial production of biomass and silicate concentrations in the lagoon channel as well as in some of the bay areas.