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At hatching the larvae of flatfish closely resemble the bilateral symmetric larvae of other teleosts, especially perciforms. Literature data show that transformation to asymmetric benthic juveniles normally occurs at body lengths between 10 and 25 mm. Unexpectedly, minimal size at its completion (including eye migration) can be 4.1 mm SL and maximal size is over 72 mm. In this paper we consider the functional requirements for a successful switch from a symmetric pelagic larva to a typical asymmetric juvenile benthic flatfish partly based on evidence from other teleosts. The unfavourable period of eye migration and transition to a benthic habitat requires some food reserves and rewiring and/or recalibration of vision and gravity-associated structures utilised previously by the still symmetric larvae for e.g. food detection. Binocular fixation of the prey probably occurs in that stage. Critical or sensitive periods occurring during development of fish larvae suggest that a completely functional symmetric stage of development must precede transformation. The normal size range in flatfish larvae at transformation seems to confirm our considerations. Recent data on temperature effects during development provide an explanation for metamorphosis at the minimal size. Some evidence for paedomorphic heterochrony in flatfish larvae is presented. 相似文献
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LIU Luofu HUO Hong GUO Yongqiang CHEN Lixin LI Shuangwen ZHAO Yande LI Yan WANG Ping CHEN Zhijun 《中国地球化学学报》2008,27(1):1-8
The results presented in this paper indicated that the carbazole-type compounds have high thermal stability and also show stability in oxidation and bio-degradation. This kind of compounds still has high concentrations and a complete distribution in the analyzed dry asphalt samples, showing that they are particularly useful in the study of hydrocarbon migration of the paleo-pools. The difference in the contents of nitrogen compounds in the Silurian dry asphalts from the Awati, Tabei and Tazhong areas is attributed to the difference in the extent of oxidation and (or) bio-degradation for the areas; the Awati and Tabei areas underwent relatively strong oxidation and bio-degradation. During the first stage of hydrocarbon pool formation in the Silurian system in the Tazhong and Tabei areas of the Tarim Basin (at the end of the Silurian period) and at the second stage in the Awati area (in Permian), the hydrocarbons experienced a long-distance migration. 相似文献
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Large-scale ancient landslides of the area of more than 5 km2 and volume exceeding 200 × 106 m3 are characteristic features of the valleys incised in the northern periphery of the Crimean Mountains (Ukraine). The largely
affected area is located in the outermost cuesta range of the Crimean Mountains which consists of rigid Sarmatian limestones
overlying weak Middle Miocene and Upper Palaeogene deposits. A giant landslide arose in the Alma water gap as a reflection
of several coincident preparatory factors such as suitable bedrock stratification, smectite-rich bedrock exposed to swelling
activity, presence of faults parallel to the valley trend, and river capture event which preceded the landslide event. The
occurrence of such ancient megaslides is particularly interesting in the area which is characterized by low precipitation
(<500 mm/year) and weak contemporary seismicity. It probably reflects a more dynamic environment in humid phases of the Holocene;
however, seismic triggering along the Mesozoic suture zone cannot be rejected. Compressional features such as gravitational
folds in the central and distal parts of the landslide, which probably correlate with the whole landslide genesis or its significant
reactivation, arose, according to the radiocarbon dating, during the Holocene climatic optimum in the Atlantic period. The
slope deformation has been relatively quiescent since that time, except minor historic reactivization which took place in
the frontal part of the landslide. We suppose that the studied landslide could be classified as a transitional type of slope
deformation with some signs of spreading and translational block slides. 相似文献
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葛店煤矿双庙扩大区位于豫、皖两省交界处。勘探区为一轴向北北东向的不对称向斜构造,地层倾角26°~30°,测区深部有岩浆岩侵入体,严重影响煤层赋存形态;地表条件复杂,村庄密集、河流较多,测区东侧塌陷区水塘大面积分布.三维地震勘探野外施工难度大。本区地震勘探观测系统设计为12线18炮制,为避免偏移归位及叠加次数降低等问题,在向斜轴部两侧、村子周边及河岸处适当加密炮点,并在塌陷区利用钢钎插置检波器以保证CDP点完整性。数据处理采取了叠前反褶积、叠前时间偏移等技术,数据处理效果显著。遵循多波组对比、多个数据体综合解释的原则,以纵向、横向和任意时间剖面相结合,充分利用三维可视化技术进行地震资料解释。本次勘探成果查明了主采煤层三2^2、二2煤层的构造形态及埋藏深度;解释落差5m以上断层34条,其中修改原有断层5条;另外新发现小于5m的异常断点22个。经巷道验证煤层底板标高误差0.43%~1.96%,巷道已揭露断层7条,与解释成果基本一致。 相似文献