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Patagonia has yielded the most comprehensive fossil record of Cretaceous theropods from Gondwana, consisting of 31 nominal species belonging to singleton taxa and six families: Abelisauridae, Noasauridae, Carcharodontosauridae, Megaraptoridae nov. fam., Alvarezsauridae, and Unenlagiidae. They provide anatomical information that allows improved interpretation of theropods discovered in other regions of Gondwana. Abelisauroids are the best represented theropods in Patagonia. They underwent an evolutionary radiation documented from the Early Cretaceous through to the latest Cretaceous, and are represented by the clades Abelisauridae and Noasauridae. Patagonian carcharodontosaurids are known from three taxa (Tyrannotitan, Giganotosaurus and Mapusaurus), as well as from isolated teeth, collected from Aptian to Cenomanian beds. These allosauroids constituted the top predators during the mid-Cretaceous, during which gigantic titanosaur sauropods were the largest herbivores. Megaraptorans have become better documented in recent years with the discovery of more complete remains. Megaraptor, Aerosteon and Orkoraptor have been described from Cretaceous beds from Argentina, and these taxa exhibit close relationships with the Aptian genera Australovenator, from Australia, and Fukuiraptor, from Japan. The Gondwanan megaraptorans are gathered into the new family Megaraptoridae, and the Asiatic Fukuiraptor is recovered as the immediate sister taxon of this clade. Although megaraptorans have been recently interpreted as members of Allosauroidea, we present evidence that they are deeply nested within Coelurosauria. Moreover, anatomical information supports Megaraptora as more closely related to the Asiamerican Tyrannosauridae than thought. Megaraptorans improve our knowledge about the scarcely documented basal radiation of Gondwanan coelurosaurs and tyrannosauroids as a whole. Information at hand indicates that South America was a cradle for the evolutionary radiation for different coelurosaurian lineages, including some basal forms (e.g., Bicentenaria, Aniksosaurus), megaraptorans, alvarezsaurids less derived than those of Laurasia, and unenlagiids, revealing that Gondwanan coelurosaurs played sharply differing ecological roles, and that they were taxonomically as diverse as in the northern continents. The unenlagiids represent an endemic South American clade that has been recently found to be more closely related to birds than to dromaeosaurid theropods. Analysis of the theropod fossil record from Gondwana shows the highest peak of origination index occurred during the Aptian–Albian and a less intense one in the Campanian time spans. Additionally, peaks of extinction index are recognized for the Cenomanian and Turonian–Coniacian time spans. In comparison, the Laurasian pattern differs from that of Gondwana in the presence of an older extinction event during the Aptian–Albian time-span and a high origination rate during the Cenomanian time-bin. Both Laurasian and Gondwanan theropod records show a peak of origination rates during the Campanian. 相似文献
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Based on representatives of the Late Cretaceous genus Gauthieria (Gauthieria radiata – Gauthieria spatulifera – Gauthieria princeps), ontogenetic trajectories within the family Phymosomatidae are described for the first time. Due to shared similarities in their ontogenetic development, an intimate evolutionary relationship must be assumed. This interpretation is most supported by analyses of the development in the ambulacral plating pattern (alternation of simple plates and compound plates), which is not commonly found among the Phymosomatoidae. This pattern, however, is present among all three species during development. The developmental trajectories of 8 further characters were included in this study (arrangement of the adapical pore pairs, number of pore pairs, pore pair numbers in ambital ambulacral plates, number of interambulacral plates, peristomal opening, apical opening diameter, test height, radial ornament of the areoles). The evolution in this lineage is characterised by several different heterochronic processes, which suggest a dissociated heterochronic evolution, indicating a developmental modularity. Additionally, the systematic treatment of G. princeps is discussed on account of the presented results. 相似文献
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准噶尔盆地不同区块烃源岩成熟度指标(主要是Ro值和Tm ax值)有较大差异,烃源岩成熟度也各自不同,但侏罗系烃源岩热成熟度总体处于未成熟-低成熟阶段,故生油能力都较低。 相似文献
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全新世中期以来黄土高原中部生物多样性研究 总被引:5,自引:0,他引:5
根据孢粉分析、古文献记载、生物地名考证以及对现今物种群多样性等方面的研究资料,结合气候变迁,论述了全新世以来,在自然和人为因素影响下,黄土高原中部地区景观和生物多样性的演变过程及其恢复和保护措施。 相似文献