Over the last 10 years a great interest in spineless cactus pear was shown in the drier areas in terms of both fresh fruit and fodder production. However, there is a lack of knowledge on quantitative data on root dynamics of these plants needed to fully understand its potential under water limiting conditions. This study aimed at quantifying the effects of water stress on the growth of tap roots, side roots and rain roots of the species Opuntia ficus-indica (L.) Miller (cultivar Morado—green cladode) and O. robusta Wendl. (cultivar Monterey—blue cladode). One-year-old cladodes were planted in root boxes and pots (2002/2003 season) that were kept in the greenhouse at day/night temperatures of 25–30 °C/15–18 °C. Placing the cladodes flat on the soil, more areoles came in contact with the soil and therefore more roots developed in both species with an average of only 3.4% areole complexes not rooting. Each areole complex formed on average 3 roots. The highest daily tap root growth was 42 and 36 mm for O. ficus-indica and O. robusta, respectively. Tap root growth increased in the morning with water stress for both species but decreased in the afternoon. Side root growth increased with water stress, with that of O. robusta more per tap root than O. ficus-indica. O. robusta showed a finer root system than O. ficus-indica. The side roots grew as much as 8 and 5 mm per day for O. ficus-indica and O. robusta, respectively. Whitish rain roots developed on the established roots within the first hour after rewetting the soil and grew for only 3 days. Rain roots grew up to 7 and 5 mm within a day for O. ficus-indica and O. robusta, respectively. Considering all studied aspects of their roots systems, O. robusta appears to be better adapted to drought (less sensitive to water stress) than O. ficus-indica. 相似文献
Soil crust lichens can be the dominant vegetation in arid lands, yet their importance as habitat to secondary producers is relatively unknown. This study examines the distribution of arthropod communities in the northern Namib Desert to evaluate whether a lichen-rich area is more or less productive than adjacent habitats in terms of the consumers each supports. Arthropods are diverse and highly endemic in the Namib Desert and lichens dominate this desert's extensive gravel plains. We sampled lichen-rich, dwarf shrub, and unvegetated sites and found distinct arthropod assemblages in the lichen-dominated sites, including species unique to lichen sites. Arthropod assemblages in two of the lichen sites were similar to those found in the dwarf shrub site. In a canonical correspondence analysis, crustose lichens and overall lichen cover were key in driving the variance in arthropod assemblages within the lichen sites. Furthermore, lichen morphotypes, overall lichen cover and species richness, were significantly correlated with the representation of arthropod subgroups and arthropod species richness. These findings provide evidence that lichen-dominated soil crusts in the Namib Desert are important supporters of secondary production, warranting more in-depth studies into the ecology and conservation of this lichen-rich habitat in warm deserts. 相似文献
Seed passage through the gut of vertebrates is critical for endozoochorus seed dispersal because it may influence seed viability and germination capacity. Foxes are important fruit-eaters and presumed seed dispersers in the Chaco ecoregion. We experimentally assessed the seed passage time (SPT) and its effect on seed viability and germination in two fox species (Pseudalopex gymnocercus and Cerdocyon thous) that occur in the Chaco. Passage time was measured using both artificial seeds of three sizes and wild seeds of four native fleshy-fruit species commonly consumed by foxes (Ziziphus mistol, Acacia aroma, Celtis tala, and Syagrus romanzoffiana). The average SPT was 6.8 h for the Pampa fox and 7.7 h for the crab-eating fox. Passage time did not differ either among seeds of different sizes or between fox species. Passage through digestive tract of both fox species did not affect seed survival, whereas germination rate was slightly improved. Our data suggest that both fox species are legitimate seed dispersers of the four species studied, having a positive effect on both germination rate and dispersal distance from the parent plant. 相似文献
Bivalve, brachiopod and cirripede faunas from the latest Jurassic and Early Cretaceous Barents Sea boreholes 7320/3-U-l and 7425/9-U-l are systematically described and illustrated. Microfossils have also been studied and the cores arc dated on the basis of the fossil recovery. The bivalve Buchia whose zonal sequence has been used for correlation of boreal marine sections is the most important biostratigraphic marker group in the condensed Boreal Berriasian-Hauterivian intervals of these cores. A new species of cirripede Zeugmatolepas? borealis Collins sp. nov. and dinoflagellate cyst Muderongia aequicornus Århus sp. nov. are described. The Late Jurassic fine-grained elastics of core 7320/3-U-l are overlain by about 3 m of grey dolomitic limestone of Valanginian and Hauterivian age. The lowermost part of 7425/9-U-l is represented by a latest Volgian-earliest Berriasian fossiliferous greyish green marl. It is followed by a reddish brown fossilifcrous claystone of Berriasian and perhaps partly Valanginian age. Core 7425/9-U-l also contains a mainly Valanginian greyish green marly limestone which changes into a dark grey to black limestone of Early Barremian age in its upper part. The sedimentological change from condensation to dark grey clay deposition took place in the middle Barremian H. rude-fissicostatum ammonite Zone in 7425/9-U-l and probably slightly earlier in 7320/3-U-l. This was commenced at about the same time as deposition of the inaccurately and only indirectly dated fluvio-deltaic Festningen Sandstone Member on Spitsbergen. The dark claystone may thus be a distal equivalent to this sandstone unit. 相似文献
The Nakuru-Elmenteita basin in the Central Kenya Rift, contains two shallow, alkaline lakes, Lake Nakuru (1770 m above sea level) and Lake Elmenteita (1786 m). Ancient shorelines and lake sediments at 1940 m suggest that these two lakes formed a single large and deep lake as a result of a wetter climate during the early Holocene. Here, we used a hydrological model to compare the precipitation–evaporation balance during the early Holocene to today. Assuming that the Nakuru-Elmenteita basin was hydrologically closed, as it is today, the most likely climate scenario includes a 45% increase in mean-annual precipitation, a 0.5°C decrease in air temperature, and an increase of 9% in cloud coverage from the modern values. Compared to the modeling results from other East African lake basins, this dramatic increase in precipitation seems to be unrealistic. Therefore, we propose a significant flow of water from the early Holocene Lake Naivasha in the south towards the Nakuru-Elmenteita basin to compensate the extremely negative hydrological budget of this basin. Since we did not find any field evidence for a surface connection, as often proposed during the last 70 years, the hydrological deficit of the Nakuru-Elmenteita basin could have also been compensated by a subsurface water exchange. 相似文献
The use of spontaneous potential (SP) anomalies is well known in the geophysical literatures because of its effectiveness and significance in solving many complex problems in mineral exploration. The inverse problem of self-potential data interpretation is generally ill-posed and nonlinear. Methods based on derivative analysis usually fail to reach the optimal solution (global minimum) and trapped in a local minimum. A new simple heuristic solution to SP anomalies due to 2D inclined sheet of infinite horizontal length is investigated in this study to solve these problems. This method is based on utilizing whale optimization algorithm (WOA) as an effective heuristic solution to the inverse problem of self-potential field due to a 2D inclined sheet. In this context, the WOA was applied first to synthetic example, where the effect of the random noise was examined and the method revealed good results using proper MATLAB code. The technique was then applied on several real field profiles from different localities aiming to determine the parameters of mineralized zones or the associated shear zones. The inversion parameters revealed that WOA detected accurately the unknown parameters and showed a good validation when compared with the published inversion methods.
In the northwestern sector of the Zagros foreland basin, axial fluvial systems initially delivered fine-grained sediments from northwestern source regions into a contiguous basin, and later transverse fluvial systems delivered coarse-grained sediments from northeastern sources into a structurally partitioned basin by fold-thrust deformation. Here we integrate sedimentologic, stratigraphic, palaeomagnetic and geochronologic data from the northwestern Zagros foreland basin to define the Neogene history of deposition and sediment routing in response to progressive advance of the Zagros fold-thrust belt. This study constrains the depositional environments, timing of deposition and provenance of nonmarine clastic deposits of the Injana (Upper Fars), Mukdadiya (Lower Bakhtiari) and Bai-Hasan (Upper Bakhtiari) Formations in the Kurdistan region of Iraq. Sediments of the Injana Formation (~12.4–7.75 Ma) were transported axially (orogen-parallel) from northwest to southeast by meandering and low-sinuosity channel belt system. In contrast, during deposition of the Mukdadiya Formation (~7.75–5 Ma), sediments were delivered transversely (orogen-perpendicular) from northeast to southwest by braided and low-sinuosity channel belt system in distributive fluvial megafans. By ~5 Ma, the northwestern Zagros foreland basin became partitioned by growth of the Mountain Front Flexure and considerable gravel was introduced in localized alluvial fans derived from growing topographic highs. Foredeep accumulation rates during deposition of the Injana, Mukdadiya and Bai-Hasan Formations averaged 350, 400 and 600 m/Myr respectively, suggesting accelerated accommodation generation in a rapidly subsiding basin governed by flexural subsidence. Detrital zircon U-Pb age spectra show that in addition to sources of Mesozoic-Cenozoic cover strata, the Injana Formation was derived chiefly from Palaeozoic-Precambrian (including Carboniferous and latest Neoproterozoic) strata in an axial position to the northwest, likely from the Bitlis-Puturge Massif and broader Eastern Anatolia. In contrast, the Mukdadiya and Bai-Hasan Formations yield distinctive Palaeogene U-Pb age peaks, particularly in the southeastern sector of the study region, consistent with transverse delivery from the arc-related terranes of the Walash and Naopurdan volcano-sedimentary groups (Gaveh-Rud domain?) and Urumieh-Dokhtar magmatic arc to the northeast. These temporal and spatial variations in stratigraphic framework, depositional environments, sediment routing and compositional provenance reveal a major drainage reorganization during Neogene shortening in the Zagros fold-thrust belt. Whereas axial fluvial systems initially dominated the foreland basin during early orogenesis in the Kurdistan region of Iraq, transverse fluvial systems were subsequently established and delivered major sediment volumes to the foreland as a consequence of the abrupt deformation advance and associated topographic growth in the Zagros. 相似文献