Cheats in a cooperative behaviour? Behavioural differences and breakdown of cooperative behaviour in aggregating,intertidal littorinids (Mollusca)

Cooperative behaviours result in the evolution of cheats: individuals that benefit from the behaviour without sharing the costs required to generate the behaviour. Normally the proportion of cheats is small, as large numbers of cheats will result in the breakdown of the behaviour. Using empirical and simulation‐derived results we demonstrate a cooperative behaviour (aggregation between two species of intertidal snails that provides a benefit by reducing desiccation stress) that shows many characteristics similar to those of a cooperative system with cheats present. In this system, the high rocky shore littorinid Echinolittorina malaccana forages for longer after high water than Echinolittorina radiata, which stops foraging and begins to form aggregations earlier. Nevertheless, E. malaccana, the ‘cheat’ in this system, still occupies an equal proportion of the most beneficial places in aggregations. Computer simulations demonstrate that up to 65% of individual snails can show the behaviour of E. malaccana before the breakdown of this aggregation behaviour begins to occur through aggregations becoming smaller, and hence less effective against desiccation. The high proportion of ‘cheats’ possible in this cooperative behaviour implies that different selective pressures may act on individuals of different species in multi‐species cooperative behaviours to those acting on individuals engaging in single species cooperative behaviours. Social symbiosis appears to be occurring between the species, but it appears that both mutualistic and parasitic symbioses are occurring.     

Submillimetre observations of luminous z>4 radio-quiet quasars and the contribution of AGN to the submm source population

We present polarization observations of the gravitational lens system B1422+231 made at 8.4 GHz using the VLBA and the 100-m telescope at Effelsberg. All four images of the quasar show structure on the milliarcsec scale. The three bright images show tangential stretching as expected from lens models. Some basic properties of gravitational lensing are exhibited by this system. The surface brightness of images A and B are the same and the parity reversal expected in image B is revealed, for the first time, by polarization observations. There is a large differential Faraday rotation between images A and B.     

The geomorphology of Sugar Loaf Mound: Prehistoric cemeteries and the formation of loess cones in the lower Wabash Valley

Burial mounds found on bluff crests in the Midwest United States overlap in scale with conical shaped Peoria Loess hills along the valley margins of the lower Wabash River. Coring of Sugar Loaf Mound, a feature thought to be an artificial mound for some 200 years, indicates that it is a 9 m eolian feature that contains prehistoric human remains. Analysis indicates that a sharp-edged till ridge resulted in a zone of eolian sand and silt accumulation forming both a climbing and a perched dune. A streamline body resulted. Grain-size analysis indicates a bimodal population, suggesting a dust suspension from an outwash source and a local saltation sand source, with a time transgressive shift in dominance. Three other loess cones of comparable scale, shape, lithostratigraphy, and landscape position that were also used as late prehistoric cemeteries indicate a similar pattern of both formation and prehistoric mortuary use in the lower Wabash valley. © 1998 John Wiley & Sons, Inc.     

Prehistoric settlement and landscape change on alluvial fans in the upper Mississippi river valley

Geoarchaeological studies of alluvial fans in the northern Sny Bottom (Upper Mississippi River valley) focused on distal fan lobe processes, the effects of small-scale landscape change on prehistoric locational decisions, and the resulting structure of the archaeological record. Aerial imagery together with coring and trenching of paleochannels on distal lobes found that frequent channel avulsion (1) produced abrupt landform/habitat changes and (2) shifted the primary loci of sediment deposition on fans through time. Analysis of wood charcoal from prehistoric occupations indicates that vegetation succession during post-avulsion overbank aggradation along a new channel was characterized by a shift from Fraxinus-dominated to mesophytic forest. Results also indicate that use of fans by Early Woodland (ca. 2550–2100 B.P.) and other foragers centered on portions along active channels. Shifting depositional loci on fans led to surfaces and depositional units of varying age, and hence a complex stratigraphic record of Holocene occupations.     

A fresh framework for the ecology of arid Australia

A revised set of propositions about ecology in arid Australia is presented, based on research literature since publication of Stafford Smith and Morton (1990). Fourteen propositions distil our argument that most features of the Australian deserts are explicable in terms of two dominant physical and climatic elements: rainfall variability, leading to extended droughts and occasional flooding rains; and widespread nutrient poverty. Different landscapes within the arid zone show these features to varying degrees, and so it is important to think about different places separately when considering our propositions. Plant life-histories strongly reflect temporal patterns of soil moisture; because Australian deserts receive more variable rainfall than most others, there is a distinctive spectrum of life-histories. Low levels of phosphorus (together with abundant soil moisture on irregular occasions) favour plants producing a relative excess of carbohydrate (C). In turn, C-rich plant products sometimes lead to fire-prone ecosystems, assemblages dominated by consumers of sap and other C-based products, and abundant detritivores (particularly termites). Fluctuations in production due to variable rainfall provide openings for consumers with opportunistic life-histories, including inhabitants of extensive but ephemeral rivers and lakes. Most consumer species exhibit some dietary flexibility or utilise more dependable resources; these strategies give rise to greater stability in species dynamics and composition of assemblages than might first be imagined under the variable rainfall regime. Aboriginal people have had long-standing ecological influence as they accessed resources. For each proposition we suggest the extent to which it is ‘different’, ‘accentuated’ or ‘universal’ in comparison with other deserts of the world, recognising that this categorisation is in need of critical testing. Further tests of each proposition are also suggested to fill the many gaps that still exist in our knowledge of the structure and functioning of Australia’s deserts.     

Vector fragility surfaces for reinforced concrete frames in Europe

Traditional approaches to damage estimation in earthquake loss modelling make use of relationships between scalar intensity measures and scalar engineering demand parameters. In this study we present a series of vector-valued fragility surfaces computed for low- and mid-rise reinforced concrete frames typical of those found in Europe. The use of vectors of intensity measures can result in conditional standard deviations of logarithmic engineering demand parameters that are up to 50 % smaller than those from traditional scalar methods. These reductions have significant implications for the shapes of loss curves, particularly for long return periods. The most efficient vector corresponds to a combination of spectral acceleration and a spectral shape parameter, \(\langle \ln S_a, \ln N_p\rangle \) , when used to predict maximum interstorey drifts. The study also demonstrates that engineering demand parameters have significant heteroskedasticity with respect to various intensity measures and that this feature must be modelled correctly when constructing fragility curves. This feature of the models presented herein has not previously been accounted for during the development of fragility curves or surfaces.     

A predictive model for Arias intensity at multiple sites and consideration of spatial correlations

Arias intensity, I_a, has been identified as an efficient intensity measure for the estimation of earthquake‐induced losses. In this paper, a new model for the prediction of Arias intensity, which incorporates nonlinear site response through the use of the average shear‐wave velocity and a heteroskedastic variance structure, is proposed. In order to estimate the effects of ground motions on spatially‐distributed systems, it is important to take into account the spatial correlation of the intensity measure. However, existing loss‐estimation models, which use I_a as an input, do not take this aspect of the ground motion into account. Therefore, the potential to model the spatial correlation of Arias intensity is also investigated. The empirical predictive model is developed using recordings from the Pacific Earthquake Engineering Research Center Next Generation of Attenuation database whereas the model for spatial correlation makes use of the well‐recorded events from this database, that is the Northridge and Chi‐Chi earthquakes. Copyright © 2011 John Wiley & Sons, Ltd.     

An evaluation of the applicability of the NGA models to ground-motion prediction in the Euro-Mediterranean region

The first phase of the Next Generation Attenuation (NGA) project has now finished, resulting in the publication of five new sets of empirical ground-motion models for PGA, PGV and response spectral ordinates. These models mark a significant advancement in the state-of-the-art in empirical ground-motion modelling and include many effects that are not accounted for in existing European equations. Under the assumption that the Euro-Mediterranean database from which the European relationships are derived is unlikely to drastically change in the near future, a prudent question to ask is: can the NGA models be applied in Europe? In order to answer this question, the NGA model of Boore and Atkinson (PEER Report 2007/01, Pacific Earthquake Engineering Research Center, Berkeley, CA, 234 pp., 2007), which is shown to be representative of the NGA models as a suite, is compared with the dataset used for the development of the most recent European empirical ground-motion models for response spectral ordinates and peak ground velocity. The comparisons are made using analyses of model residuals and the likelihood approach of Scherbaum et al. (Bull Seism Soc Am 94(6):2164–2185, 2004). The analyses indicate that for most engineering applications, and particularly for displacement-based approaches to seismic design, the NGA models may confidently be applied within Europe. Furthermore, it is recommended that they be used in conjunction with existing European models to provide constraint on finite-fault effects and non-linear site response within logic-tree frameworks. The findings also point to the potential benefits of merging the NGA and European datasets.     

Numbers of scaled and matched accelerograms required for inelastic dynamic analyses

Selecting, scaling and matching accelerograms are critically important to engineering design and assessment, enabling structural response to be determined with greater confidence and through fewer analyses than if unscaled accelerograms are employed. This paper considers the response of an 8‐storey multiple‐degree‐of‐freedom reinforced concrete structure to accelerograms selected, linearly scaled or spectrally matched using five different techniques. The first method consists of selecting real records on the basis of seismological characteristics, while the remaining methods make an initial selection on the basis of magnitude and spectral shape before (1) scaling to the target spectral acceleration at the initial period; (2) scaling to the target spectrum over a range of periods; (3) using wavelet adjustments to match the target spectrum and (4) using wavelet adjustments to match multiple target spectra for multiple damping ratios. The analyses indicate that the number of records required to obtain a stable estimate of the response decreases drastically as one moves through these methods. The exact number varies among damage measures and is related to the predictability of the damage measure. For measures such as peak roof and inter‐storey drift, member end rotation and the Park and Ang damage index, as few as one or two records are required to estimate the response to within ±5% (for a 64% confidence level) if matching to multiple damping ratios is conducted. Bias checks are made using predictive equations of the expected response derived from the results of 1656 nonlinear time‐domain analyses of the structure under the action of unscaled accelerograms. Copyright © 2008 John Wiley & Sons, Ltd.