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141.
142.
Properties of the light saturation curve of photosynthesis and ribulose-1,5-bisphosphate carboxylase (RuBPC) activity are shown to change qualitatively in a natural population of marine phytoplankton during a spring bloom. Evidence is presented to show that these changes constitute photoadapative responses to increasing irradiance. As irradiance increased during the bloom, both the level of light-saturated photosynthesis (Pm) and the initial slope of the light saturation curve (α = photosynthetic efficiency) increased whether those parameters were normalized to chlorophyll a concentration (Pmb, αb) or to cell numbers (Pmc, αc). The magnitudes of these changes were such that Ik (= Pm/α, the photoadaptation parameter) did not change, but Im, the light intensity at which photosynthesis becomes saturated, increased. RuBPC activity, both chlorophyll a (RuBPCb) and cell number normalized (RuBPCc), also increased during the bloom. We suggest that these adaptations were achieved by simultaneously increasing the number of photosynthetic units, proportionately decreasing the photosynthetic unit size, and increasing both the concentrations of the enzymes of the dark reactions and possibly also of photosynthetic electron transport components.We also observed diminished levels of photoinhibition in the high light adapted cells late in the bloom and have suggested that this was a consequence of the same suite of physiological changes.In situ carbon fixation per cell increased during the bloom whereas no change occurred in this parameter when normalized to chlorophyll a concentration. Although these photoadaptive responses thus permitted carbon to be fixed in situ more rapidly per cell, at a constant efficiency with respect to investment of energy in the photosynthetic apparatus, they did not result in a change in growth rate. Based on consideratios of the role of time scale in physiological adaptation, however, it is suggested that the observed alterations in photosynthesis with increasing irradiance might permit a cell to more rapidly fill an energy quota for division, possibly an advantage in a mixing environment in which energy is patchily distributed, both spatially and temporalyy.Phosphoenolpyruvate carboxylase activity when normalized to chlorophyll a (PEPCb) did not change during the bloom while chlorophyll a normalized dark carbon fixation decreased sharply and was quantitatively small compared to PEPCb. On this basis and considering that RuBPCb increased during the bloom, it is suggested that, although PEPC may be involved in dark carbon fixation, its most important quantitative role is probably an indirect one in light dependent photosynthesis.We have also considered the relevance of laboratory results on photoadaptation to interpretations of field studies and have suggested that batch culture studies must be treated with caution but that turbidistat and semi-continuous methods provide reasonable simulations of natural conditions. 相似文献
143.
大沽河是胶州湾最大的入海河流,每年都会给胶州湾带来大量泥沙。本文利用三维的ECOMSED水动力-泥沙数值模型对洪水期的大沽河进行了数值模拟,得到了洪水存在期间以及洪水结束后胶州湾的余流分布、大沽河河道上中下游各层盐度、泥沙浓度随时间变化的趋势以及入海冲淡水和泥沙对胶州湾的影响。 相似文献
144.
The mixing characteristics of particles such as dredged sediment of variable size discharged into cross flow are studied by a 3D numerical model, which is developed to model the particle-fluid two-phase flow. The Eulerian method with the modified k-ε parameterization of turbulence for the fluid phase is used to solve fluid phase, while a Lagrangian method for the solid phase (particles), both the processes are coupled through the momentum sources. In the model the wake turbulence induced by particles has been included as additional source term in the k-ε model; and the variable drift velocities of the particles are treated efficiently by the Lagrangian method in which the particles are tracked explicitly and the diffusion process is approximated by a random walk model. The hydrodynamic behavior of dumping a cloud of particles is governed by the total buoyancy of the cloud, the drag force on each particle and the velocity of cross-flow. The computed results show a roughly linear relationship between the displacement of the frontal position and the longitudinal width of the particle cloud. The particle size in the cloud and the velocity of cross flow dominate the flow behavior. The computed results are compared with the results of laboratory experiments and satisfactory agreement is obtained. 相似文献
145.
声波在海底界面的反射和透射是海底散射、海底混响、海底目标探测的重要问题。利用Biot多孔介质声传播理论对声波在水-多孔海底界面上的反射和透射进行了分析,具体给出反射声波的反射系数,3种透射声波的透射系数以及声能透射系数随入射波入射角和频率(10~40 kHz)的变化关系,分析了各种透射波对透射声能的贡献。多孔海底介质参数分别采用Stoll和Chotiros给出的2组参数进行理论计算。 相似文献
146.
用电生理学方法观察了4种不同浓度的高纯河豚毒素(tetrodotoxin,TTX)对蛙离体坐骨神经动作电位的影响.实验结果表明:1μmol/dm3TTX在给药后7 min能显著抑制动作电位的上相幅度(P<0.05),但在给药后10 min才显著抑制动作电位的下相幅度(P<0.01);20μmol/dm3TTX在给药后3 min内能完全抑制动作电位的形成.1μmol/dm3TTX在给药后1~5min对动作电位传导速度的影响没有统计学意义,5μmol/dm3TTX在5min时可显著抑制传导速度(P<0.05),而10μmol/dm3TTX在2min时便显著抑制传导速度(P<0.05).实验结果提示高纯河豚毒素对蛙离体坐骨神经动作电位的影响存在剂量效应关系. 相似文献
147.
Two distinct series of slumps deform the upper part of the sedimentary sequence along the continental margin of the Levant.
One series is found along the base of the continental slope, where it overlies the disrupted eastern edge of the Messinian
evaporites. The second series of slumps transects the continental margin from the shelf break to the Levant Basin. It seemed
that the two series were triggered by two unrelated, though contemporaneous, processes. The shore-parallel slumps were initiated
by basinwards flow of the Messinian salt, that carried along the overlying Plio-Quaternary sediments. Seawater that percolated
along the detachment faults dissolved the underlying salt to form distinctly disrupted structures. The slope-normal slumps
are located on top of large canyons that cut into the pre-Messinian sedimentary rocks. A layer of salt is found in the canyons,
and the Plio-Quaternary sediments were deposited on that layer. The slumps are bounded by large, NW-trending faults where
post-Messinian faulted offset was measured. We presume that the flow of the salt in the canyons also drives the slope-normal
slumps. Thus thin-skinned halokynetic processes generated the composite post-Tortonian structural patterns of the Levant margin.
The Phoenician Structures are a prime example of the collapse of a distal continental margin due to the dissolution of a massive
salt layer. 相似文献
148.
149.
150.
利用漳州港海域无围堰填海造地前后的水深测量数据,结合海域地质钻探、表层和柱状沉积物资料,用数字地形模型方法计算了该区海底的冲淤量,分析了冲淤变化及其影响因素,结果显示:(1)淤积主要发生在近岸海域,其主要原因是近岸3-10m厚的海底淤泥层在回填土的挤压和推移下发生形变,产生淤浅假象;另一个原因是回填时部分泥沙流失入海,淤积在近岸海域,致使淤积强度增大。(2)由近岸往外,淤积强度逐渐减弱,并过渡为产中刷状态。 相似文献