The number of immature stages and the seasonal patterns of development are basic life history features of a stream dwelling species and knowledge about these important components are essential for understanding its adaptations to its dynamic environment. The life cycle of Agapetus fuscipes (Trichoptera, Glossosomatidae), one of the dominant scrapers in the upper and middle reaches of the Breitenbach, a first-order upland stream in central Germany, was analysed. The pronotum length and the relationship between pronotum length, larval biomass and case length showed seven distinct larval instars, contrary to earlier findings from the Breitenbach. In addition to a few trichopteran species from other functional feeding groups, A. fuscipes is the only scraping caddis fly reported to have more than five larval instars. The moult increments of pronotum length and larval biomass were distinctly lower than in glossosomatid species with five larval instars. A. fuscipes is clearly univoltine in the Breitenbach. First-instar larvae were found from July to the beginning of December, and second-instar larvae from July to January. At the beginning of December the population consisted of the instars I to V, and development did not cease during winter. The sixth-instar larvae occurred mostly in January, and the seventh-instar larvae were never present before January. The prepupae and pupae occurred in April. The last pupae were found at the beginning of September, although most of the emergence took place in June and July. At least five different immature stages with different ecological demands were present at any time throughout the year. The ecological advantage having two additional larval instars compared to other glossosomatid species may be to compensate for the high rate of mouthpart wear that occurs while the larvae feed on the rough Bunter Sandstone substratum. A further advantage may be to spread the risk of high mortality under unfavourable environmental conditions. 相似文献
In the study of soil erosion, specifically on detachment of soil particles by raindrop impact, kinetic energy is a commonly suggested indicator of the raindrop's ability to detach soil particles from the soil mass. Since direct measurement of kinetic energy requires sophisticated and costly instruments, the alternative approach is to estimate it from rainfall intensity. The present study aims at establishing a relationship between rainfall intensity and kinetic energy for rainfalls in Central Cebu, Philippines as a preface of a wider regional investigation.
Drop size distributions of rainfalls were measured using the disdrometer RD-80. There are two forms of kinetic energy considered here. One is kinetic energy per unit area per unit time (KER, J m−2 h−1) and the other is kinetic energy per unit area per unit depth (KE, J m−2 mm−1). Relationships between kinetic energy per unit area per unit time (KER) and rainfall intensity (I) were obtained using linear and power relations. The exponential model and the logarithmic model were fitted to the KE–I data to obtain corresponding relationships between kinetic energy per unit area per unit depth of rainfall (KE) and rainfall intensity (I). The equation obtained from the exponential model produced smaller standard error of estimates than the logarithmic model. 相似文献
Denitrification rates along a salinity gradient in the eutrophic Neuse River Estuary, North Carolina, were quantified using
membrane inlet mass spectrometry (MIMS) within short-term batch incubations. Denitrification rates within the system were
highly variable, ranging from 0 to 275 μmol N m−2 h−1. Intrasite variability increased with salinity, but no significant differences were observed across the salinity gradient.
Denitrification rates were positively correlated with sediment oxygen demand at the upstream sampling site where sediment
organic carbon levels were lowest. This relationship was not observed in the more saline sampling sites. Denitrification rates
were highest during winter. On an annual basis, denitrification accounted for 26% of the dissolved inorganic nitrogen and
12% of the total nitrogen supplied to the system. 相似文献
We present a general stratigraphic synthesis for the Upper Rhine Graben (URG) and the Swiss Molasse Basin (SMB) from Eocene to Pliocene times. The stratigraphic data were compiled both from literature and from research carried out by the authors during the past 6 years ; an index of the stratigraphically most important localitites is provided. We distinguish 14 geographical areas from the Helvetic domain in the South to the Hanau Basin in the North. For each geographical area, we give a synthesis of the biostratigraphy, lithofacies, and chronostratigraphic ranges. The relationships between this stratigraphic record and the global sea-level changes are generally disturbed by the geodynamic (e.g., subsidence) evolution of the basins. However, global sea-level changes probably affected the dynamic of transgression–regression in the URG (e.g., Middle Pechelbronn Beds and Serie Grise corresponding with sea-level rise between Ru1/Ru2 and Ru2/Ru3 sequences, respectively) as well as in the Molasse basin (regression of the UMM corresponding with the sea-level drop at the Ch1 sequence). The URGENT-project (Upper Rhine Graben evolution and neotectonics) provided an unique opportunity to carry out and present this synthesis. Discussions with scientists addressing sedimentology, tectonics, geophysics and geochemistry permitted the comparison of the sedimentary history and stratigraphy of the basin with processes controlling its geodynamic evolution. Data presented here back up the palaeogeographic reconstructions presented in a companion paper by the same authors (see Berger et al. in Int J Earth Sci 2005). 相似文献
The relative importance of climate, forest fires and human population size on long‐term boreal forest composition were statistically investigated at regional and local scales in Fennoscandia. We employ pollen data from lakes, reflecting regional vegetation, and small forest hollows, reflecting local vegetation, from Russia, Finland and Sweden to reconstruct the long‐term forest composition. As potential drivers of the Holocene forest dynamics we consider climate, generated from a climate model and oxygen isotope data, past forest fires generated from sedimentary charcoal data and human population size derived from radiocarbon dated archaeological findings. We apply the statistical method of variation partitioning to assess the relative importance of these environmental variables on long‐term boreal forest composition. The results show that climate is the main driver of the changes in Holocene boreal forest composition at the regional scale. However, at the local scale the role of climate is relatively small. In general, the importance of forest fires is low both at regional and local scales. The fact that both climate and forest fires explain relatively small proportions of variation in long‐term boreal vegetation in small forest hollow records demonstrates the complexity of factors affecting stand‐scale forest dynamics. The relative importance of human population size was low in both the prehistorical and the historical time periods. However, this is the first time that this type of data has been used to statistically assess the importance of human population size on boreal vegetation and the spatial representativeness of the data may cause bias to the analysis. 相似文献
Following Appalachian orogenesis, metamorphic rocks in central Newfoundland were exhumed and reburied under Tournaisian strata. New zircon fission‐track (ZFT) ages of metamorphic rocks below the Tournaisian unconformity yield post‐depositionally reset ages of 212–235 Ma indicating regional fluid‐absent reheating to at least ≥220°C. Post‐Tournaisian sedimentary thicknesses in surrounding basins show that burial alone cannot explain such temperatures, thus requiring that palaeo‐geothermal gradients increased to ≥30–40°C/km before final late Triassic accelerated cooling. We attribute these elevated palaeo‐geothermal gradients to localized thermal blanketing by insulating sediments overlying radiogenic high‐heat‐producing granitoids. Late Triassic rifting and magmatism before break up of Pangaea likely also contributed to elevated heat flow, as well as uplift, triggering late Triassic accelerated cooling and exhumation. Thermochronological ages of 240–200 Ma are seen throughout Atlantic Canada, and record rifting and basaltic magmatism on the conjugate margins of the Central Atlantic Ocean preceding the onset of oceanic spreading at ~190 Ma. 相似文献