In two waters: contemporary evolution of lagoonal and marine white seabream (Diplodus sargus) populations

Brackish water ecosystems are often exposed to wide variations in environmental variables, including temperature and salinity, which may cause strong selective pressures on organisms modifying the genetic patterns of species. The aim of this work was to test whether there is a ‘divergence-with-gene flow’ in coastal lagoon populations of white seabream (Diplodus sargus) (Ria Formosa, S Portugal and Mar Menor, SE Spain) respect to four marine populations, by using partial sequences of cyt b mitochondrial gene and information from nine microsatellite loci. Genetic diversity was highest in both coastal lagoons (Mar Menor and Ria Formosa) considering mitochondrial and nuclear markers. Although some of F_ST population pairwise comparisons were not significant, analyses of molecular variance (AMOVAs) detected differences between groups (coastal lagoon and marine) close to significance. Also, only two haplotypes (Cytb-17 and Cytb-18) were detected in both coastal lagoon sampling sites and these localities (Mar Menor and Ria Formosa) showed the highest number of singletons, some of them with a high number of mutations, as has been already described for other Mar Menor populations (Pomatochistus marmoratus and Holothuria polii). Also, several tests detected significant positive and balancing selection considering mtDNA and microsatellite data. These data support the hypothesis of selection as one of the drivers of the genetic differences found between coastal lagoon and marine populations. The life strategy adopted by Diplodus sargus in coastal lagoons allows it to decrease its mortality rate and improve the heritability of its genes. Also, the increase time spent in coastal lagoons with different temperatures and salinities favours the fitness selection and the maintenance of exclusive haplotypes and genotypes in coastal lagoon inhabitants favouring the ‘divergence-with-gene-flow’.     

West versus East Mediterranean Sea: origin and genetic differentiation of the sea cucumber Holothuria polii

We studied the genetic structure of the sea cucumber Holothuria (Roweothuria) polii (Delle Chiaje 1823) by analysing the mitochondrial DNA variation in two fragments of cytochrome oxidase I (COI) and 16S genes. Individuals were collected in seven locations along the Mediterranean Sea, which cover a wide range of the species distribution. We found high haplotype diversity for COI and moderate diversity for 16S, and low nucleotide diversity for both genes. Our results for the COI gene showed many recent and exclusive haplotypes with few mutational changes, suggesting recent or ongoing population expansion. The Western and Eastern Mediterranean populations exhibited slight but significant genetic differentiation (COI gene) with higher genetic diversity in the East. The most ancient haplotype was not present in the westernmost sampling location (SE Spain). The oldest expansion time was observed in Turkey, corresponding to mid‐Pleistocene. Turkey had also the highest genetic diversity (number of total and exclusive haplotypes, polymorphisms, haplotype and nucleotide diversity). This suggests that this region could be the origin of the subsequent colonizations through the Mediterranean Sea, a hypothesis that should be assessed with nuclear markers in future research.     

Artificial Marine Micro‐Reserves Networks (AMMRNs): an innovative approach to conserve marine littoral biodiversity and protect endangered species

Coastal engineering works, especially the outer breakwaters of ports where environmental conditions are usually good, often constitute new habitats for marine littoral species. Their potential indirect contribution to the regeneration and conservation of protected species opens up a new perspective in protection of marine biodiversity. This is particularly true when these works harbour, as we show here, important reproductive populations of species that are threatened or even in danger of extinction. We provide integrated data on protected marine species in Southern Iberia established on different substrata (quarried dolomitic rocks, concrete cubic blocks, concrete tetrapods and vertical concrete walls) that are used in the construction of artificial levees that could potentially be Artificial Marine Micro‐Reserves (AMMRs). We also present the first evidence of the ‘reserve effect’ in potential AMMRs and we discuss the need for the creation of AMMRs including their effective networking (AMMRNs), supporting data for which has been observed by studying coastal facilities subjected to strict access restrictions (for security reasons, not conservation). These facilities have acted as valuable proxies and, in reality, potentially act as AMMRs, as well as having a role in helping to detect and control invasive species. We discuss the ecological engineering aspects related to the design and type of materials used in coastal constructions with a view to aiding the integration of these into existing coastal ecosystems as well as promoting greater settlement of species and therefore increases in biodiversity. Finally, the environmental implications of AMMRNs within the future implementation strategy of the Water Framework Directive 2000/60 EC are also discussed.     

Beach morphodynamics forcements in oiled shorelines: Coupled physical and chemical processes during and after fuel burial

In November 2002, the sinking of the Prestige oil tanker off the Galician coast (N.W. Spain) caused the largest ecological catastrophe in the history of Spain, affecting the coast called the 'Costa da Morte' (Galicia, N.W. Spain). This work is focused on the study of the oil contamination of the intertidal area of two beaches located on this stretch of coast. The study of twenty cores extracted from both beaches has identified fuel embedded in the sedimentary column up to a depth of 2.38 m (this being the maximum depth of extraction). This, along with the presence of oil below the groundwater indicates the existence of a new factor which determines the burial of oil: the morphodynamic behaviour of the beach. Furthermore, this morphodynamic variation conditions the physical appearance of the buried oil. Four different types have been identified: tar-balls (cm), particles (mm), oil coatings on sediment grains and on emulsion, with distribution patterns conditioned by the degree of wave exposure. The analysis of the information obtained have permitted the development of a conceptual model of the burial and oil evolution in the sedimentary column in relation to wave exposure, and thus to the morphodynamic variability of the beach.     

CME Classification Based on Wavelet Spectra

We study the internal structure of coronal mass ejections (CMEs) using wavelet analysis. We derive wavelet spectra, spatially integrated over regions of interest within LASCO C2 white-light coronographic images. These spectra show an inflection point, which we use to characterize each spectrum. In a diagram of flux vs. spatial scale of the inflection point, we find that the analyzed structures fall into two, distinct groups: a low-flux, small-spatial-scale group (which we call the “homogeneous” type), and a high-flux, larger-spatial-scale group (the “collimated” type). Interestingly, if we study different regions of a given image, all of the structures fall into one of the two groups described above. From a qualitative comparison with the images, it is clear that the two groups identified by the wavelet analysis correspond to two types of morphologies, which are seen as either more-homogeneous or more-collimated structures.     

European temperature records of the past five centuries based on documentary/instrumental information compared to climate simulations

Two European temperature reconstructions for the past half-millennium, January-to-April air temperature for Stockholm (Sweden) and seasonal temperature for a Central European region, both derived from the analysis of documentary sources and long instrumental records, are compared with the output of climate simulations with the model ECHO-G. The analysis is complemented by comparisons with the long (early)-instrumental record of Central England Temperature (CET). Both approaches to study past climates (simulations and reconstructions) are burdened with uncertainties. The main objective of this comparative analysis is to identify robust features and weaknesses in each method which may help to improve models and reconstruction methods. The results indicate a general agreement between simulations obtained with temporally changing external forcings and the reconstructed Stockholm and CET records for the multi-centennial temperature trend over the recent centuries, which is not reproduced in a control simulation. This trend is likely due to the long-term change in external forcing. Additionally, the Stockholm reconstruction and the CET record also show a clear multi-decadal warm episode peaking around AD 1730, which is absent in the simulations. Neither the reconstruction uncertainties nor the model internal climate variability can easily explain this difference. Regarding the interannual variability, the Stockholm series displays, in some periods, higher amplitudes than the simulations but these differences are within the statistical uncertainty and further decrease if output from a regional model driven by the global model is used. The long-term trend of the CET series agrees less well with the simulations. The reconstructed temperature displays, for all seasons, a smaller difference between the present climate and past centuries than is seen in the simulations. Possible reasons for these differences may be related to a limitation of the traditional ‘indexing’ technique for converting documentary evidence to temperature values to capture long-term climate changes, because the documents often reflect temperatures relative to the contemporary authors’ own perception of what constituted ‘normal’ conditions. By contrast, the amplitude of the simulated and reconstructed inter-annual variability agrees rather well.     

Tidal and wind-induced circulation within the Southeastern limit of the Bay of Biscay: Pasaia Bay,Basque Coast

The Basque coastal area, in the southeastern Bay of Biscay, can be characterised as being more influenced by land climate and inputs, than other typically ‘open sea’ areas. The influence of coastal processes, together with the presence of irregular and steep topography, complicate greatly the water circulation patterns. Water movement along the Basque coastal area is not well understood; observations are scarce and long-term current records are lacking. The knowledge available is confined to the surface currents: the surface water circulation is controlled mainly by wind forcing, with tidal and density currents being weak. However, there is a lack of knowledge available on currents within the lower levels of the water column; likewise, on the main time-scales involved in the water circulation. This study quantifies the contribution of the tidal and wind-induced currents, to the overall water circulation; it identifies the main time-scales involved within the tidal and wind-induced flows, investigating difference in such currents, throughout the water column, within Pasaia Bay (Basque coast). Within this context, extensive oceanographic and meteorological data have been obtained, in order to describe the circulation. The present investigation reveals that the circulation, within the surface and the sub-surface waters, is controlled mainly by wind forcing fluctuations, over a wide range of meteorological frequencies: third-diurnal, semidiurnal and diurnal land–sea breezes; synoptic variability; frequencies, near fortnightly periods; and seasonal. At the lower levels of the water column, the main contribution to the water circulation arises from residual currents, followed by wind-induced currents on synoptic time-scales. In contrast, tidal currents contribute minimally to the overall circulation throughout the water column.     

Modelling of Seismic Ground Motion in Santiago de Cuba City from Earthquakes in Oriente Fault Seismic Zone

— We present the results of complete P-SV and SH waves modelling, up to a maximum frequency of 1?Hz, along two profiles in Santiago de Cuba city. The seismic sources are located in the depth range from 10 to 40?km on the Oriente fault zone at distances of several tens of kilometres from the city. The calculation has been made by a hybrid method: Modal summation in the regional anelastic model (one-dimensional) where the source is buried, and finite differences in the local sedimentary anelastic models (two-dimensional). The analysis of the influence of the depth and of the distance of the source on the site effects shows that standard traditional methods, based on the deconvolution analysis of the rock outcrop motion, can lead to erroneous results.     

Edge wave and non-trapped modes of the 25 april 1992 Cape Mendocino tsunami

The 25 April 1992 Cape Mendocino earthquake generated a tsunami characterized by both coastal trapped edge wave and non-trapped tsunami modes that propagated north and south along the U.S. West Coast. Both observed and synthetic time series at Crescent City and North Spit are consistent with the zero-order edge wave mode solution for a semi-infinite sloping beach depth profile. Wave amplitudes at Crescent City were about twice that observed at North Spit, in spite of the fact that the source region was three times farther from Crescent City than North Spit. The largest observed amplitude was due to an edge wave which arrived almost three hours after the initial onset of the tsunami; since such waves are highly localized nearshore, this suggests that the enhanced responsiveness at Crescent City is at least partly due to local dynamic processes. Furthermore, the substantially delayed arrival of this wave, which was generated at the southern end of the Cascadia Subduction Zone, has significant implications for hazard mitigation efforts along the entire U.S. West Coast. Specifically, this study demonstrates that slow-moving but very energetic edge wave modes could be generated by future large tsunamigenic earthquakes in the CSZ, and that these might arrive unexpectedly at coastal communities several hours after the initial tsunami waves have subsided.     

Regional significance of kilometric-scale north-east vergent recumbent folds associated with east to south-east directed shear on the southern border of the Central Iberian Zone (Hornachos-Oliva region,Variscan belt,Iberian Peninsula)

On the southern border of the Central Iberian Zone there are two sectors with different styles of deformation. To the south-west, in the Hornachos sector, large-scale recumbent folds associated with ductile shearing can be seen. This shearing is characterized by a direction of movement parallel to the fold axes and can be correlated for 150 km along strike. The K-values of the strain ellipsoid range from 0.8 to 2.0. Stretching in the X direction, parallel to the recumbent fold axes, is more than 100%. To the north-east, in the Oliva sector, first-phase folds are upright and the strain intensity is lower than in the Hornachos sector. Metamorphic, geometric and kinematic considerations lead us to conclude that the shearing in the Hornachos sector is better explained as conjugate to a main shear zone along which the southern border of the Central Iberian Zone is moved onto the Ossa-Morena Zone. This main thrust is at present obliterated by a left-lateral extensional shear zone that affects a high pressure exotic unit located between the Central Iberian and the Ossa-Morena Zones. This high pressure unit constitutes a suture of the Variscan belt in the Iberian Peninsula.