Sipunculus nudus Linnaeus, 1766 (Sipuncula): cosmopolitan or a group of pseudo‐cryptic species? An integrated molecular and morphological approach

Sipunculan taxonomy relies on a limited set of external morphological and internal anatomical characters. In addition, this marine group is characterized by an unusual large number of putatively cosmopolitan species. However, this ‘cosmopolitan’ status could be an artifact of their conserved morphology and the small number of unambiguous taxonomic characters available for delimiting species. Species delimitation can therefore be aided by molecular techniques. We investigated the case of the widespread and common species Sipunculus nudus Linnaeus, 1766 to determine its systematic validity. We analysed the morphology of multiple specimens of S. nudus collected from 11 localities around the world and undertook phylogenetic analyses using molecular sequence data from four genes (28S rRNA, 16S rRNA, histone H3 and cytochrome c oxidase subunit I). High levels of genetic differentiation are present between distantly related populations of the putative species S. nudus. Five distinct lineages were identified by phylogenetic analyses, three of which – the best‐represented populations – can be distinguished morphologically. Our phylogenetic and morphological analyses thus do not favor the cosmopolitan status of S. nudus, suggesting instead that it constitutes a complex of morphologically similar but distinguishable species.     

Aeolian/fluvial interactions and high-resolution sequence stratigraphy of a non-marine lowstand wedge: the Avilé Member of the Agrio Formation (Lower Cretaceous), central Neuquén Basin, Argentina

Abstract Accumulation within the unconformity‐based Hauterivian Avilé Sandstone of the Neuquén Basin, Argentina, was characterized by a close interaction between fluvial and aeolian processes developed after a major relative sea‐level drop that almost completely desiccated the entire basin and juxtaposed these non‐marine deposits on shallow‐ and deep‐marine facies. Aeolian deposits within the Avilé Member include dune (A1) and sand sheet (A2) units that characterize the lower part of the unit. Fluvial deposits comprise distal flood units (F1) interbedded with aeolian dune deposits in the middle part of the succession, and low‐ (F2) and high‐sinuosity (F3) channels associated with floodplain deposits (F4) towards the top. The internal characteristics of the aeolian system indicate that its accumulation was strongly controlled by water‐table dynamics, with the development of multiple horizontal deflation super surfaces that truncate dune deposits and form the basal boundary of flood deposits and sand sheet units. A long‐term wetting‐upward trend is recorded throughout the entire unit, with an increase in fluvial activity towards the top and the development of a more permanent fluvial system overlying a major erosion surface interpreted as a sequence boundary. The upward increase in water‐table influence might be related to relative sea‐level rise, which controlled the position of the water table and allowed the accumulation of tabular aeolian units bounded by horizontal deflation surfaces. This high‐frequency, eustatically driven process acted together with a long‐term climatic change towards wetter conditions.     

The Upper Jurassic (Kimmeridgian) fluvial–aeolian systems of the southern Neuquén Basin, Argentina

The Kimmeridgian Quebrada del Sapo Formation in the southernmost Neuquén Basin in Argentina represents a succession up to 40 m thick of coarse- to fine-grained fluvial deposits overlain by aeolian deposits. These fluvial–aeolian deposits reflect a significant palaeogeographic change in the basin and are related to a major, tectonically enhanced, relative sea-level fall. The fluvial section is dominated by braided-channel, fine-grained ephemeral, and sheetflood deposits. Aeolian facies are dominated by dune deposits, with minor sandsheet and interdune units. Changes in the nature of both fluvial and aeolian sedimentation within the studied area suggest a regional variability of accommodation/sediment supply conditions. The regional changes of the aeolian succession likely reflect different relative positions within a major erg. In the upwind margin of the erg, a shallow water table promoted water-lain sedimentation in interdune areas, whereas in the central parts of the erg, dry sediment accumulation took place above the regional water-table level. The vertical transition observed in the Quebrada del Sapo Formation, from fluvial to aeolian deposits, may be the result of a local climatic change to drier conditions due to the development of a climatic barrier imposed by growth of a magmatic arc to the west. Alternatively, the vertical transition could be related to a lowering of the water table associated with the compartmentalization of the basin during a period of low sea level.     

Approximate solutions of non-linear circular orbit relative motion in curvilinear coordinates

A compact, time-explicit, approximate solution of the highly non-linear relative motion in curvilinear coordinates is provided under the assumption of circular orbit for the chief spacecraft. The rather compact, three-dimensional solution is obtained by algebraic manipulation of the individual Keplerian motions in curvilinear, rather than Cartesian coordinates, and provides analytical expressions for the secular, constant and periodic terms of each coordinate as a function of the initial relative motion conditions or relative orbital elements. Numerical test cases are conducted to show that the approximate solution can be effectively employed to extend the classical linear Clohessy–Wiltshire solution to include non-linear relative motion without significant loss of accuracy up to a limit of 0.4–0.45 in eccentricity and 40–45\(^\circ \) in relative inclination for the follower. A very simple, quadratic extension of the classical Clohessy–Wiltshire solution in curvilinear coordinates is also presented.     

An interdisciplinary approach to the collapse of the port and degradation of freshwater resources at Panama Viejo (Panama), 1519–1671

The Archaeological Site of Panama Viejo (Panama) comprises a protected area of 28 km² within present-day Panama City, on America's Pacific coast. In 1519, the Spaniards founded the city of Panama Viejo to secure a natural port in an area inhabited by indigenous peoples since at least the eighth century CE. The site, along the coastline and between two rivers, became a principal gateway for goods and people travelling between Europe and Pacific settlements to the east (Realejo) and the west (Trujillo, Lima, Arica). Within one century, however, Panama Viejo's natural port and freshwater infrastructure collapsed, leading to the city's relocation after corsairs attacked it in 1671. This study combines archaeological, geographical and historical analyses to explain how and why human interactions with the local environment made the settlement increasingly untenable.     

Variogram-Based Descriptors for Comparison and Classification of Rock Texture Images

Mathematical Geosciences - Rock characterization is typically performed by geologists in mining companies and involves the analysis of several meters of drill-hole samples to describe distinctive...     

Stock Evaluation of Three Littoral Echinoid Species on the Catalan Coast North-Western Mediterranean

Abstract. We estimated the abundances of the three common littoral echinoid species Paracentrotus lividus, Arbacia lixula and Sphaerechinus granularis in a littoral zone between 0 and 10m depth on the Catalan shores NE Spain. Densities of each species were estimated from a total number of 11 571× 1 m² sampling units deployed at 152 sites along the 13 zones into which the coast was divided. The surface area available in each of these zones was estimated at the scale of the sampling unit 1 m² taking into account the fractal dimension of the shore. The stocks individuals > 2cm test diameter of these species for the whole Catalan shore ≅ 300 km, straight-line distance were estimated to be about 279.1 × 10⁶ individuals for P. lividus , 28.9 × 10⁶ individuals for A. lixula and 10.9 × 10⁶ individuals for S. granularis. These sea-urchins are key species in the regulation of algal communities and some of them P. lividus. S. granularis are of commercial interest. This information will provide a tool for the future monitoring and management of this resource, which now faces diverse pressures of human origin.