The impact of overfishing and El Niño on the condition factor and reproductive success of Peruvian hake, Merluccius gayi peruanus

In this study we examine physiological responses of Peruvian hake (Merluccius gayi peruanus) to changes in their population structure and physical environment during the period 1971–2004. In particular, we assess the relative investment of energy in growth and/or reproduction of small (20–35 cm) and large (35–50 cm) hake. We calculated the (i) condition (Fulton’s K), (ii) gonad and (iii) gut fullness indices for 42,761 female hakes sampled from commercial landings; these indices indicate fish somatic, reproductive and feeding condition, respectively. Using Generalized Additive Models we then examined potential relationships between these indices and sea surface temperature anomalies and date. Drastic energy exhaustion and a decrease in female hake fecundity were observed during El Niño events. The long-term trend showed a general increase in condition factor and a decrease in gonad index for large hake between 1971 and 2004. Small hake exhibited a different trend with an increase in reproductive activity, which was accompanied by an earlier maturation. We hypothesise that the observed low investment of energy in reproduction by large female hake might be related to the lack of large males, due to a sex-selective fishery and the impact of El Niño. We suggest that fishing diminished hake reproductive capacity, modified the sex ratio in favour of females and increased population vulnerability to environmental stress, in particular to the El Niño. The impact of multidecadal variability and predators like the squid, Dosidicus gigas, remain unresolved until longer time series become available.     

Trophic modeling of the Northern Humboldt Current Ecosystem, Part II: Elucidating ecosystem dynamics from 1995 to 2004 with a focus on the impact of ENSO

The Northern Humboldt Current Ecosystem is one of the most productive in the world in terms of fish production. Its location near to the equator permits strong upwelling under relatively low winds, thus creating optimal conditions for the development of plankton communities. These communities ultimately support abundant populations of grazing fish such as the Peruvian anchoveta, Engraulis ringens. The ecosystem is also subject to strong inter-annual environmental variability associated with the El Niño Southern Oscillation (ENSO), which has major effects on nutrient structure, primary production, and higher trophic levels. Here our objective is to model the contributions of several external drivers (i.e. reconstructed phytoplankton changes, fish immigration, and fishing rate) and internal control mechanisms (i.e. predator-prey) to ecosystem dynamics over an ENSO cycle. Steady-state models and time-series data from the Instituto del Mar del Perú (IMARPE) from 1995 to 2004 provide the base data for simulations conducted with the program Ecopath with Ecosim. In simulations all three external drivers contribute to ecosystem dynamics. Changes in phytoplankton quantity and composition (i.e. contribution of diatoms and dino- and silicoflagellates), as affected by upwelling intensity, were important in dynamics of the El Niño of 1997–98 and the subsequent 3 years. The expansion and immigration of mesopelagic fish populations during El Niño was important for dynamics in following years. Fishing rate changes were the most important of the three external drivers tested, helping to explain observed dynamics throughout the modeled period, and particularly during the post-El Niño period. Internal control settings show a mix of predator–prey control settings; however a “wasp-waist” control of the ecosystem by small pelagic fish is not supported.     

The influence of a mature cyclonic eddy on particle export in the lee of Hawaii

Mesoscale eddies may enhance primary production (PP) in the open ocean by bringing nutrient-rich deep waters into the euphotic zone, potentially leading to increased transport of particles to depth. This hypothesis remains controversial, however, due to a paucity of direct particle export measurements. In this study, we investigated particle dynamics using ²³⁴Th–²³⁸U disequilibria within a mesoscale cold-core eddy, Cyclone Opal, which formed in the lee of the Hawaiian Islands. ²³⁴Th samples were collected along two transects across Cyclone Opal as well as during a time-series within the eddy core during a decaying diatom bloom. Particulate carbon (PC), particulate nitrogen (PN) and biogenic silica (bSiO₂) fluxes at 150 m varied spatially and temporally within the eddy and strongly depended on the ²³⁴Th model formulation used (e.g., steady state versus non-steady state, inclusion of upwelling, etc.). Particle fluxes estimated from a steady state model assuming an upwelling rate of 2 m day⁻¹ yielded the best fit to sediment-trap data. These ²³⁴Th-derived particle fluxes ranged from 332±14 to 1719±53 μmol C m⁻² day⁻¹, 27±3 to 114±12 μmol N m⁻² day⁻¹, and 33±20 to 309±73 μmol Si m⁻² day⁻¹. Although PP rates within Cyclone Opal were elevated by a factor of 2–3, PC and PN fluxes were the same, within error, inside and outside of Cyclone Opal. The ratio of PC export to PP remained surprisingly low at <0.03 and similar to those measured in surrounding waters. In contrast, bSiO₂ fluxes within the eddy core were three times higher. Detailed analyses of ²³⁴Th depth profiles consistently showed excess ²³⁴Th at 100–175 m, associated with the remineralization and possible accumulation of suspended and dissolved organic matter from the surface. We suggest that strong microzooplankton grazing facilitated particulate organic matter recycling and resulted in the export of empty diatom frustules. Thus, while eddies may increase PP, they do not necessarily increase PC and PN export to deep waters. This may be a general characteristic of wind-driven cyclonic eddies of the North Pacific Subtropical Gyre and suggests that eddies may preferentially act as a silica pump, thereby playing an important role in promoting silicic-acid limitation in the region.     

Youth Mobility and Territorial Disparities: An Analysis of Urban and Rural Barcelona

Travel behavior of the present generation of youths is being increasingly explored due to their relevance in shaping future accessibility needs and mobility habits. The present study offers an original perspective on this topic by identifying territorial disparities that emerge in youth mobility patterns in rural and urban areas. Unlike most previous research, we propose taking a global view on mobility by analyzing all trip purposes and transportation modes. This is conducted by analyzing a comprehensive mobility survey in the Barcelona Metropolitan Region, which provides data on mobility engagement, trip purposes, modal split, travel times and territorial differences. In general, youngsters account for larger daily travel times than older adults, present a higher attachment to public transportation and walk less on a daily basis. These differences are enhanced in rural territories, where while older adults overcome accessibility issues with higher use of the private vehicle, youngsters are more likely to invest larger travel times on transit.     

Paleogene evolution and demise of the proto‐Paratethys Sea in Central Asia (Tarim and Tajik basins): Role of intensified tectonic activity at ca. 41 Ma

The proto‐Paratethys Sea covered a vast area extending from the Mediterranean Tethys to the Tarim Basin in western China during Cretaceous and early Paleogene. Climate modelling and proxy studies suggest that Asian aridification has been governed by westerly moisture modulated by fluctuations of the proto‐Paratethys Sea. Transgressive and regressive episodes of the proto‐Paratethys Sea have been previously recognized but their timing, extent and depositional environments remain poorly constrained. This hampers understanding of their driving mechanisms (tectonic and/or eustatic) and their contribution to Asian aridification. Here, we present a new chronostratigraphic framework based on biostratigraphy and magnetostratigraphy as well as a detailed palaeoenvironmental analysis for the Paleogene proto‐Paratethys Sea incursions in the Tajik and Tarim basins. This enables us to identify the major drivers of marine fluctuations and their potential consequences on Asian aridification. A major regional restriction event, marked by the exceptionally thick (≤ 400 m) shelf evaporites is assigned a Danian‐Selandian age (ca. 63–59 Ma) in the Aertashi Formation. This is followed by the largest recorded proto‐Paratethys Sea incursion with a transgression estimated as early Thanetian (ca. 59–57 Ma) and a regression within the Ypresian (ca. 53–52 Ma), both within the Qimugen Formation. The transgression of the next incursion in the Kalatar and Wulagen formations is now constrained as early Lutetian (ca. 47–46 Ma), whereas its regression in the Bashibulake Formation is constrained as late Lutetian (ca. 41 Ma) and is associated with a drastic increase in both tectonic subsidence and basin infilling. The age of the final and least pronounced sea incursion restricted to the westernmost margin of the Tarim Basin is assigned as Bartonian–Priabonian (ca. 39.7–36.7 Ma). We interpret the long‐term westward retreat of the proto‐Paratethys Sea starting at ca. 41 Ma to be associated with far‐field tectonic effects of the Indo‐Asia collision and Pamir/Tibetan plateau uplift. Short‐term eustatic sea level transgressions are superimposed on this long‐term regression and seem coeval with the transgression events in the other northern Peri‐Tethyan sedimentary provinces for the 1st and 2nd sea incursions. However, the 3rd sea incursion is interpreted as related to tectonism. The transgressive and regressive intervals of the proto‐Paratethys Sea correlate well with the reported humid and arid phases, respectively in the Qaidam and Xining basins, thus demonstrating the role of the proto‐Paratethys Sea as an important moisture source for the Asian interior and its regression as a contributor to Asian aridification.     

Salt thickness and composition influence rift structural style,northern North Sea,offshore Norway

“Salt” giants are typically halite‐dominated, although they invariably contain other evaporite (e.g. anhydrite, bittern salts) and non‐evaporite (e.g. carbonate, clastic) rocks. Rheological differences between these rocks mean they impact or respond to rift‐related, upper crustal deformation in different ways. Our understanding of basin‐scale lithology variations in ancient salt giants, what controls this and how this impacts later rift‐related deformation, is poor, principally due to a lack of subsurface datasets of sufficiently regional extent. Here we use 2D seismic reflection and borehole data from offshore Norway to map compositional variations within the Zechstein Supergroup (ZSG) (Lopingian), relating this to the structural styles developed during Middle Jurassic‐to‐Early Cretaceous rifting. Based on the proportion of halite, we identify and map four intrasalt depositional zones (sensu Clark et al., Journal of the Geological Society, 1998, 155, 663) offshore Norway. We show that, at the basin margins, the ZSG is carbonate‐dominated, whereas towards the basin centre, it becomes increasingly halite‐dominated, a trend observed in the UK sector of the North Sea Basin and in other ancient salt giants. However, we also document abrupt, large magnitude compositional and thickness variations adjacent to large, intra‐basin normal faults; for example, thin, carbonate‐dominated successions occur on fault‐bounded footwall highs, whereas thick, halite‐dominated successions occur only a few kilometres away in adjacent depocentres. It is presently unclear if this variability reflects variations in syn‐depositional relief related to flooding of an underfilled presalt (Early Permian) rift or syn‐depositional (Lopingian) rift‐related faulting. Irrespective of the underlying controls, variations in salt composition and thickness influenced the Middle Jurassic‐to‐Early Cretaceous rift structural style, with diapirism characterising hangingwall basins where autochthonous salt was thick and halite‐rich and salt‐detached normal faulting occurring on the basin margins and on intra‐basin structural highs where the salt was too thin and/or halite‐poor to undergo diapirism. This variability is currently not captured by existing tectono‐stratigraphic models largely based on observations from salt‐free rifts and, we argue, mapping of suprasalt structural styles may provide insights into salt composition and thickness in areas where boreholes are lacking or seismic imaging is poor.     

Strain analysis of a seismically imaged mass‐transport complex,offshore Uruguay

Strain style, magnitude and distribution within mass‐transport complexes (MTCs) are important for understanding the process evolution of submarine mass flows and for estimating their runout distances. Structural restoration and quantification of strain in gravitationally driven passive margins have been shown to approximately balance between updip extensional and downdip contractional domains; such an exercise has not yet been attempted for MTCs. We here interpret and structurally restore a shallowly buried (c. 1,500 mbsf) and well‐imaged MTC, offshore Uruguay using a high‐resolution (12.5 m vertical and 15 × 12.5 m horizontal resolution) three‐dimensional seismic‐reflection survey. This allows us to characterise and quantify vertical and lateral strain distribution within the deposit. Detailed seismic mapping and attribute analysis shows that the MTC is characterised by a complicated array of kinematic indicators, which vary spatially in style and concentration. Seismic‐attribute extractions reveal several previously undocumented fabrics preserved in the MTC, including internal shearing in the form of sub‐orthogonal shear zones, and fold‐thrust systems within the basal shear zone beneath rafted‐blocks. These features suggest multiple transport directions and phases of flow during emplacement. The MTC is characterised by a broadly tripartite strain distribution, with extensional (e.g. normal faults), translational and contractional (e.g. folds and thrusts) domains, along with a radial frontally emergent zone. We also show how strain is preferentially concentrated around intra‐MTC rafted‐blocks due to their kinematic interactions with the underlying basal shear zone. Overall, and even when volume loss within the frontally emergent zone is included, a strain difference between extension (1.6–1.9 km) and contraction (6.7–7.3 km) is calculated. We attribute this to a combination of distributed, sub‐seismic, ‘cryptic’ strain, likely related to de‐watering, grain‐scale deformation and related changes in bulk sediment volume. This work has implications for assessing MTCs strain distribution and provides a practical approach for evaluating structural interpretations within such deposits.     

The timing of salt structure growth in the Southern Permian Basin (Central Europe) and implications for basin dynamics

In this paper, a literature‐based compilation of the timing and history of salt tectonics in the Southern Permian Basin (Central Europe) is presented. The tectono‐stratigraphic evolution of the Southern Permian Basin is influenced by salt movement and the structural development of various types of salt structures. The compilation presented here was used to characterize the following syndepositional growth stages of the salt structures: (a) “phase of initiation”; (b) phase of fastest growth (“main activity”); and (c) phase of burial’. We have also mapped the spatial pattern of potential mechanisms that triggered the initiation of salt structures over the area studied and summarized them for distinct regions (sub‐basins, platforms, etc.). The data base compiled and the set of maps produced from it provide a detailed overview of the spatial and temporal distribution of salt tectonic activity enabling the correlation of tectonic phases between specific regions of the entire Southern Permian Basin. Accordingly, salt movements were initiated in deeply subsided graben structures and fault zones during the Early and Middle Triassic. In these areas, salt structures reached their phase of main activity already during the Late Triassic or the Jurassic and were mostly buried during the Early Cretaceous. Salt structures in less subsided sub‐basins and platform regions of the Southern Permian Basin mostly started to grow during the Late Triassic. The subsequent phase of main activity of these salt structures took place from the Late Cretaceous to the Cenozoic. The analysis of the trigger mechanisms revealed that most salt structures were initiated by large‐offset normal faults in the sub‐salt basement in the large graben structures and minor normal faulting associated with thin‐skinned extension in the less subsided basin parts.     

Normal fault growth influenced by basement fabrics: The importance of preferential nucleation from pre‐existing structures

Reactivation of pre‐existing intra‐basement structures can influence the evolution of rift basins, yet the detailed kinematic relationship between these structures and overlying rift‐related faults remains poorly understood. Understanding the kinematic as well as geometric relationship between intra‐basement structures and rift‐related fault networks is important, with the extension direction in many rifted provinces typically thought to lie normal to fault strike. We here investigate this problem using a borehole‐constrained, 3D seismic reflection dataset from the Taranaki Basin, offshore New Zealand. Excellent imaging of intra‐basement structures and a relatively weakly deformed, stratigraphically simple sedimentary cover allow us to: (a) identify a range of interaction styles between intra‐basement structures and overlying, Plio‐Pleistocene rift‐related normal faults; and (b) examine the cover fault kinematics associated with each interaction style. Some of the normal faults parallel and are physically connected to intra‐basement reflections, which are interpreted as mylonitic reverse faults formed during Mesozoic subduction and basement terrane accretion. These geometric relationships indicate pre‐existing intra‐basement structures locally controlled the position and attitude of Plio‐Pleistocene rift‐related normal faults. However, through detailed 3D kinematic analysis of selected normal faults, we show that: (a) normal faults only nucleated above intra‐basement structures that experienced late Miocene compressional reactivation, (b) despite playing an important role during subsequent rifting, intra‐basement structures have not been significantly extensionally reactivated, and (c) preferential nucleation and propagation of normal faults within late Miocene reverse faults and folds appears to be the key genetic relationship between contractionally reactivated intra‐basement structures and rift‐related normal faults. Our analysis shows that km‐scale, intra‐basement structures can control the nucleation and development of newly formed, rift‐related normal faults, most likely due to a local perturbation of the regional stress field. Because of this, simply inverting fault strike for causal extension direction may be incorrect, especially in provinces where pre‐existing, intra‐basement structures occur. We also show that a detailed kinematic analysis is key to deciphering the temporal as well as simply the spatial or geometric relationship between structures developed at multiple structural levels.     

Clinoform architecture and along‐strike facies variability through an exhumed erosional to accretionary basin margin transition

Exhumed basin margin‐scale clinothems provide important archives for understanding process interactions and reconstructing the physiography of sedimentary basins. However, studies of coeval shelf through slope to basin‐floor deposits are rarely documented, mainly due to outcrop or subsurface dataset limitations. Unit G from the Laingsburg depocentre (Karoo Basin, South Africa) is a rare example of a complete basin margin scale clinothem (>60 km long, 200 m‐high), with >10 km of depositional strike control, which allows a quasi‐3D study of a preserved shelf‐slope‐basin floor transition over a ca. 1,200 km² area. Sand‐prone, wave‐influenced topset deposits close to the shelf‐edge rollover zone can be physically mapped down dip for ca. 10 km as they thicken and transition into heterolithic foreset/slope deposits. These deposits progressively fine and thin over tens of km farther down dip into sand‐starved bottomset/basin‐floor deposits. Only a few km along strike, the coeval foreset/slope deposits are bypass‐dominated with incisional features interpreted as minor slope conduits/gullies. The margin here is steeper, more channelized and records a stepped profile with evidence of sand‐filled intraslope topography, a preserved base‐of‐slope transition zone and sand‐rich bottomset/basin‐floor deposits. Unit G is interpreted as part of a composite depositional sequence that records a change in basin margin style from an underlying incised slope with large sand‐rich basin‐floor fans to an overlying accretion‐dominated shelf with limited sand supply to the slope and basin floor. The change in margin style is accompanied with decreased clinoform height/slope and increased shelf width. This is interpreted to reflect a transition in subsidence style from regional sag, driven by dynamic topography/inherited basement configuration, to early foreland basin flexural loading. Results of this study caution against reconstructing basin margin successions from partial datasets without accounting for temporal and spatial physiographic changes, with potential implications on predictive basin evolution models.