An integrative study of larval organogenesis of American shad <Emphasis Type="Italic">Alosa sapidissima</Emphasis> in histological aspects

We describe organogenesis at a histological level in American shad (Alosa sapidissima) larvae from 0 until 45 days after hatching (DAH). Larval development was divided into four stages based on the feeding mode, external morphological features, and structural changes in the organs: stage 1 (0–2 DAH), stage 2 (3–5 DAH), stage 3 (6–26 DAH) and stage 4 (27–45 DAH). At early stage 2 (3 DAH), American shad larvae developed the initial digestive and absorptive tissues, including the mouth and anal opening, buccopharyngeal cavity, oesophagus, incipient stomach, anterior and posterior intestine, differentiated hepatocytes, and exocrine pancreas. The digestive and absorptive capacity developed further in stages 2 to 3, at which time the pharyngeal teeth, taste buds, gut mucosa folds, differentiated stomach, and gastric glands could be observed. Four defined compartments were discernible in the heart at 4 DAH. From 3 to 13 DAH, the excretory systems started to develop, accompanied by urinary bladder opening, the appearance and development of primordial pronephros, and the proliferation and convolution of renal tubules. Primordial gills were detected at 2 DAH, the pseudobranch was visible at 6 DAH, and the filaments and lamellae proliferated rapidly during stage 3. The primordial swim bladder was first observed at 2 DAH and started to inflate at 9 DAH; from then on, it expanded constantly. The spleen was first observed at 8 DAH and the thymus was evident at 12 DAH. From stage 4 onwards, most organs essentially manifested an increase in size, number, and complexity of tissue structure.     

俯冲带的后撤与弧后扩张

西太平洋地壳年龄较老,因而岩石层较冷和比重较大,俯冲带的角度也较大,活动和成熟的弧后盆地则较多;条件与之相反的东太平洋弧后盆地则较少.本文探讨这种相关关系的力学成因,计算了俯冲板块诱生的弧后上涌地幔流动.计算表明,俯冲角度大及存在后撤俯冲时,有利于在弧后地区产生明显的上涌地幔流,这种深部热物质的上涌会导致弧后扩张.反之,年龄较轻的海洋地块较热和较轻,俯冲角度一般也较小,不易诱生上涌地幔物质流动和弧后扩张.大陆地壳密度小于地幔物质,大陆碰撞区就更不具备弧后扩张的条件.