Spatial patterns and storage composition of woody debris in a natural secondary forest dominated by <Emphasis Type="Italic">Pinus tabulaeformis</Emphasis> on Loess Plateau,China

Woody debris (WD) is an important part of natural Pinus tabulaeformis mixed stands, and it affects the forest ecosystem stability and development. The WD spatial patterns are especially important structural characteristics that can provide insights into forest dynamics. In this paper, the WD storage, WD spatial patterns and WD associations among the main species were examined in the natural secondary forest on Loess Plateau in northwest China. Data were collected in a 1 ha (100 m × 100 m) permanent plot, and all the trees with a diameter at breast height of more than 3 cm were measured and stem-mapped. Ripley’s K functions from the spatial-point-pattern-analysis method were used to analyze the spatial distribution and associations. The results showed that: (1) The total storage of WD was 10.73 t/ha, fallen wood was the main source of WD, and the majority diameters were greater than 20 cm, and in intermediate levels of decay; (2) The overall spatial pattern was closely related to the spatial scale, which exhibited an aggregated pattern on a small scale, and a random pattern on a large scale. The spatial patterns of coarse woody debris also gradually transitioned from an aggregated pattern in fine scales to a random pattern in broader spatial scales, which matched the overall spatial pattern. The spatial intensity was gradually decreased with the increasing diameters, and increased with the decomposition classes; (3) The WD of Pinus tabulaeformis species was negatively associated with Betula platyphylla and Populus davidiana on a small scale but positively associated with these species on a large scale. The spatial pattern and interspecies relations were the results of long-term interactions between the natural secondary forest community and the surrounding natural environment. These findings would provide a scientific basis for the sustainable management and protection of natural secondary forest ecosystems on Loess Plateau.     

Effects of climate change on potential habitats of the cold temperate coniferous forest in Yunnan province,southwestern China

We built a classification tree(CT) model to estimate climatic factors controlling the cold temperate coniferous forest(CTCF) distributions in Yunnan province and to predict its potential habitats under the current and future climates, using seven climate change scenarios, projected over the years of 2070-2099. The accurate CT model on CTCFs showed that minimum temperature of coldest month(TMW) was the overwhelmingly potent factor among the six climate variables. The areas of TMW-4.05 were suitable habitats of CTCF, and the areas of -1.35 TMW were non-habitats, where temperate conifer and broad-leaved mixed forests(TCBLFs) were distribute in lower elevation, bordering on the CTCF. Dominant species of Abies, Picea, and Larix in the CTCFs, are more tolerant to winter coldness than Tsuga and broad-leaved trees including deciduous broad-leaved Acer and Betula, evergreen broadleaved Cyclobalanopsis and Lithocarpus in TCBLFs. Winter coldness may actually limit the cool-side distributions of TCBLFs in the areas between -1.35°C and -4.05°C, and the warm-side distributions of CTCFs may be controlled by competition to the species of TCBLFs. Under future climate scenarios, the vulnerable area, where current potential(suitable + marginal) habitats(80,749 km~2) shift to nonhabitats, was predicted to decrease to 55.91%(45,053 km~2) of the current area. Inferring from the current vegetation distribution pattern, TCBLFs will replace declining CTCFs. Vulnerable areas predicted by models are important in determining priority of ecosystem conservation.     

Structure and regeneration dynamics of three forest types at different succession stages of spruce – fir mixed forest in Changbai Mountain,northeastern China

Analyzing and understanding the structure and growth dynamics of forests at different stages is helpful to promote forest succession, restoration and management. Three spots representing three succession stages of spruce-fir mixed forest(SF: polar-birch secondary forest, MF: spruce-fir mixed forest and PF: spruce-fir near primary forest) were established. Structure, growth dynamics during two growth seasons for dominant tree species, regeneration were examined, and a univariate O-ring function statistic was used to analyze the spatial patterns of main regeneration tree species. Results showed that,(1) composition of tree species, periodic annual increment(PAI) of the diameter at breast height(DBH), basal area for overstory trees and of ground diameter(DGH) for saplings, were significantly different with the succession;(2) the current species composition and regeneration dynamics of SF suggested a development towards spruce-fir mixed forests. Pioneer species like Betula platyphyllaa will gradually disappear while climax species, such as Abies nephrolepis, Pinus koraiensis, Picea koraiensis and Tilia amurensis will dominate forest stands;(3) Despite the highest volume occurring in PF, and saplings in it grew better than in the others, this forest type is unstable because of its unsustainable structure of DBH class and insufficient regeneration; and(4) MF had the most reasonable distribution of DBH class for adult trees(DBH 5.0 cm) and DGH class for saplings(H ≥30 cm and DBH ≤5 cm), as well as an optimal volume increment. Limiting canopy opening size can lessen the physiological stress and promote the growth and competitive status of regeneration. Management implications for increasing the gaps and thus creating better growth conditions for understory saplings and facilitating forest succession were discussed.     

Study on vegetation and climate changes in Beijing region since late pleistocene

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神农架林区植被分布与地形的关系研究

神农架林区是我国物种多样性最为丰富的地区之一,地形地貌复杂,对植被分布影响巨大。本文利用该地区2007年数字高程数据、2007年植被分布图以及2017年野外实地调查数据,基于最大熵模型和空间分析理论,从植被类型和种群两个角度研究该地区不同尺度植被空间分布的地形特征,分别量化植被类型和种群空间分布的地形范围,得到植被类型与地形因子关系模型、植被种群与地形因子关系模型。结果表明:①神农架林区影响植被空间分布的地形因子不同,其中影响针叶林分布的最重要的地形因子是高程和高程变异系数,影响阔叶林分布的是高程和坡向,影响灌丛分布的是坡向变率和坡向,影响草丛分布的较为分散;②典型植被种群分布的地形范围和植被类型的基本一致,其中90%针叶林分布在高程1600~2600 m间,典型种群巴山冷杉和华山松主要分布在高程1700~3200 m和1700~2200 m;85%的阔叶林分布在高程1000~2000 m间,典型种群青冈类和鹅耳枥主要分布在高程1200~2200 m间;95%的灌丛分布在坡向变率0~40°间,典型种群杜鹃和蔷薇主要分布在坡向变率小于40°的范围,但相应的关系模型存在差异,植被类型与地形因子为高斯模型,典型种群与地形因子关系模型相对复杂,不同种群的分布模式不同;③虽然坡度常作为数字地形的重要因子,但本文研究发现该地区坡度对植被类型和种群分布的影响不明显。研究结果可为神农架林区植被保护和恢复,以及植被规划和管理提供基础参考。     

Impact factors identification of spatial heterogeneity of herbaceous plant diversity on five southern islands of Miaodao Archipelago in North China

Herbaceous plants are widely distributed on islands and where they exhibit spatial heterogeneity. Accurately identifying the impact factors that drive spatial heterogeneity can reveal typical island biodiversity patterns. Five southern islands in the Miaodao Archipelago, North China were studied herein. The spatial distribution of herbaceous plant diversity on these islands was analyzed, and the impact factors and their degree of impact on spatial heterogeneity were identified using CCA ordination and ANOVA. The results reveal 114 herbaceous plant species, belonging to 94 genera from 34 families in the 50 plots sampled. The total species numbers on different islands were significantly positively correlated with island area, and the average α diversity was correlated with human activities, while the β diversity among islands was more affected by island area than mutual distances. Spatial heterogeneity within islands indicated that the diversities were generally high in areas with higher altitude, slope, total nitrogen, total carbon, and canopy density, and lower moisture content, pH, total phosphorus, total potassium, and aspect. Among the environmental factors, pH, canopy density, total K, total P, moisture content, altitude, and slope had significant gross effects, but only canopy density exhibited a significant net effect. Terrain affected diversity by restricting plantation, plantation in turn influenced soil properties and the two together affected diversity. Therefore, plantation was ultimately the fundamental driving factor for spatial heterogeneity in herbaceous plant diversity on the five islands.     

Dynamics and Relationships of Ca,Mg,Fe in Litter,Soil Fauna and Soil in Pinus koraiensis-Broadleaf Mixed Forest

The Liangshui Natural Reserve in Heilongjiang Province of China was selected as the study area.The authors collected the samples of forest litter (Tilia amurensis,Fraxinus mandshurica,Pinus koraiensis,Acer mono,Betula costata,and mixed litter),soil in humus horizon (0-5cm) and soil horizon (5-20cm),and soil macrofauna (Oligochaeta,Geophiloporpha and Juliformia) from 2001 to 2002.The role of soil macrofauna in the material cycle was analyzed through comparing the macro-element contents among various parts of the subsystems and using enrichment index (EI).The results indicate that dynamic changes of various litters are very complicated.The contents of Fe in each kind of litter increase firstly,and then decrease in the study period.The changes of macro-element contents are greater in the broad-leaf litter than in the coniferous litter,and the mixed litter is in the middle level,but the differences among them are not significant.The contents of Mg and Fe in humus are higher than those in soil,but the contents of Ca in soil are higher than that in humus.The dynamic changes of macro-element contents in soil and soil fauna are not consistent with those in litter.The diplopod presented obvious enrichment of Ca and Mg (EI>1),but it does not significantly enrich Fe.Earthworm has a stronger enrichment ability of Fe than diplopod and scolopendra,but EI<1.Soil fauna can make great influences on the material cycle of the subsystems.     

Omni-directional distribution patterns of montane coniferous forest in the Helan Mountains of China

Slope aspect has significant effect on the development and distribution of montane forest,especially in arid and semiarid regions.This paper,using SPOT5 images and 1:50,000 DEM,digitally extracts and analyzes the spatial information of montane coniferous forest(mainly Qinghai spruce),and thereby explores how the upper and lower limits and the altitudinal range of coniferous forest vary and how the area of coniferous forest is related with annual insolation with all aspects in the Helan Mountains.The results show that: 1) In the eastern flank,the lower limit of coniferous forest is between 1,600 m and 2,000 m a.s.l.,and the upper limit between 2,800 m and 3,000 m a.s.l.;in the western flank,the lower limit of coniferous forest is between 2,000 m and 2,300 m a.s.l.,and the upper limit between 2,800 m and 3,100 m a.s.l.2) The altitudinal ranges of coniferous forest are 806~1,435 m,438~1,140 m for eastern flank and western flank,respectively.3) The area of coniferous forest takes on a normal distribution with aspect,and it has a close relationship with annual insolation.This distribution model developed in this paper quantitatively reveals the significant effect of slope aspect on the distribution of coniferous forest in arid and semi-arid land.     

Spatial pattern analysis and associations of different growth stages of populations of Abies georgei var.smithii in Southeast Tibet,China

Abies georgei var.smithii is an important plant species in Southeast Tibet,China.It has high ecological value in terms of biodiversity protection,as well as soil and water conservation.We analyzed the spatial pattern and associations of A.georgei var.smithii populations at different growth stages by using Ripley's L function for point pattern analysis.The diameter structure was a nearly reverse 'J' shape.The amount of saplings and medium-sized trees accounts for a large part of the entire population,suggesting a high regeneration rate and an expanding population.In the transition from saplings to medium trees or to large trees,saplings show a significant aggregation distribution at small scales,while medium trees and large trees show a random distribution.There are significant inverse associations between saplings and medium trees and large trees at small scales,while there are no obvious associations between medium trees and large trees.The natural regeneration was affected by interspecific competition,and it was also affected by intraspecific competition.The joint effects of biological characteristics and environmental factors contribute to the spatial distribution pattern and associations of this A.georgei var.sm ithii population.     

Dynamics and relationships of Ca, Mg, Fe in litter, soil fauna and soil in Pinus koraiensis-broadleaf mixed forest

The Liangshui Natural Reserve in Heilongjiang Province of China was selected as the study area. The authors collected the samples of forest litter （Tilia amurensis, Fraxinus mandshurica, Pinus koraiensis, Acer mono, Betula costata, and mixed litter）, soil in humus horizon （0--5cm） and soil horizon （5-20cm）, and soil macrofauna （Oligochaeta, Geophiloporpha and Juliformia） from 2001 to 2002. The role of soil macrofauna in the material cycle was analyzed through comparing the macro-element contents among various parts of the subsystems and using enrichment index （El）. The results indicate that dynamic changes of various litters are very complicated. The contents of Fe in each kind of litter increase firstly, and then decrease in the study period. The changes of macro-element contents are greater in the broad-leaf litter than in the coniferous litter, and the mixed litter is in the middle level, but the differences among them are not significant. The contents of Mg and Fe in humus are higher than those in soil, but the contents of Ca in soil are higher than that in humus. The dynamic changes of macro-element contents in soil and soil fauna are not consistent with those in litter. The diplopod presented obvious enrichment of Ca and Mg （E1〉1）, but it does not significantly enrich Fe. Earthworm has a stronger enrichment ability of Fe than diplopod and scolopendra, but E1〈1. Soil fauna can make great influences on the material cycle of the subsystems.     

Diversity,composition and host-species relationships of epiphytic orchids and ferns in two forests in Nepal

Epiphytic plant species are an important part of biological diversity. It is therefore essential to understand the distribution pattern and the factors influencing such patterns. The present study is aimed at observing the patterns of species richness, abundances and species composition of epiphytic orchids and ferns in two subtropical forests in Nepal. We also studied the relationship of host plants (Schima wallichii and Quercus lanata) and epiphyte species. Data were collected in Naudhara community forest (CF) and the national forest (NF) in Shivapuri Nagarjun National Park. The data were analyzed using univariate and multivariate tests. In total, we recorded 41 species of epiphytes (33 orchid and 8 fern species). Orchid species abundance is significantly higher in CF compared to NF. Orchid species richness and abundance increased with increasing southern aspect whereas it decreased with increasing canopy cover, and fern species richness increased with host bark roughness. Orchid abundance was positively correlated with increasing bark pH, stem size, tree age and tree height and negatively correlated with increasing steepness of the area. Likewise, fern abundances were high in places with high canopy cover, trees that were tall and big, but decreased with increasing altitude and southern aspect. The composition of the orchid and fern species was affected by altitude, aspect, canopy cover, DBH, number of forks and forest management types. We showed that the diversity of orchid and fern epiphytes is influenced by host characteristics as well as host types. The most important pre-requisite for a high epiphyte biodiversity is the presence of old respectively tall trees, independent of the recent protection status. This means: (i) for protection, e.g. in the frame of the national park declaration, such areas should be used which host such old tall trees; and (ii) also in managed forests and even in intensively used landscapes epiphytes can be protected by letting a certain number of trees be and by giving them space to grow old and tall.     

PALEOSOLS OF SANDY LANDS AND ENVIRONMENTAL CHANGES IN THE WESTERN PART OF NORTHEAST PLAIN OF CHINA WESTERN DURING HOLOCENE

ＰＡＬＥＯＳＯＬＳＯＦＳＡＮＤＹＬＡＮＤＳＡＮＤＥＮＶＩＲＯＮＭＥＮＴＡＬＣＨＡＮＧＥＳＩＮＴＨＥＷＥＳＴＥＲＮＰＡＲＴＯＦＮＯＲＴＨＥＡＳＴＰＬＡＩＮＯＦＣＨＩＮＡＷＥＳＴＥＲＮＤＵＲＩＮＧＨＯＬＯＣＥＮＥ￥ＱｉｕＳｈａｎｗｅｎ（裘善文）ＬｉＱｕｓ...     

Spatial pattern analysis and associations of different growth stages of populations of <Emphasis Type="Italic">Abies georgei</Emphasis> var. <Emphasis Type="Italic">smithii</Emphasis> in Southeast Tibet,China

Abies georgei var. smithii is an important plant species in Southeast Tibet, China. It has high ecological value in terms of biodiversity protection, as well as soil and water conservation. We analyzed the spatial pattern and associations of A. georgei var. smithii populations at different growth stages by using Ripley’s L function for point pattern analysis. The diameter structure was a nearly reverse ‘J’ shape. The amount of saplings and medium-sized trees accounts for a large part of the entire population, suggesting a high regeneration rate and an expanding population. In the transition from saplings to medium trees or to large trees, saplings show a significant aggregation distribution at small scales, while medium trees and large trees show a random distribution. There are significant inverse associations between saplings and medium trees and large trees at small scales, while there are no obvious associations between medium trees and large trees. The natural regeneration was affected by interspecific competition, and it was also affected by intraspecific competition. The joint effects of biological characteristics and environmental factors contribute to the spatial distribution pattern and associations of this A. georgei var. smithii population.     

典型深街谷内树木空间配置对行人呼吸高度处气流的影响

为揭示树木的不同空间配植方案对行人呼吸高度气流的影响,本文将树木视为均匀多孔介质,通过附加源项法从空气动力学角度用CFD模拟了H/W=2的典型深街谷几何内4种树木配植情景,实验表明,不同空间配置下树木对街谷内行人呼吸高度处局地气流的影响强弱在空间分布模式上差异悬殊：① 均匀种植的树木对街谷内行人呼吸高度的气流起到阻碍作用,不均匀种植则有效提升街谷的整体流速。4种空间配植方案下树木对气流的影响程度不同,阻碍作用从大到小的顺序为均匀间距8 m（Spa8m）>均匀间距6 m（Spa6m）>均匀间距20 m（Spa20m）>不均匀配植（Non-uniform）;对应的平均气流增强指标顺序为$\bar{D}_{spa8}$（-19.31%）<$\bar{D}_{spa6}$（-16.14%）<$\bar{D}_{spa20}$（-10.73%）<$\bar{D}_{non-uniform}$（1.25%）。② 对比不均匀和均匀的种植方案,不均匀植树的街谷内部行人呼吸高度的气流流速比其对照案例（均匀植树Spa8m方案）整体增强了106.49%。街谷中部不种树,在街谷两端配置树木并预留足够的自由空间的不均匀植树方案,能够让角涡渗入街谷中部,促使街谷内部的垂直漩涡和两端的水平角涡运动,增强湍流和垂直交换,有效减少了街道两端“风口效应”和街道中部“风影效应”的区域,改善了整个街谷行人呼吸平面的风环境。④ 合理空间配置的树木能够改善街谷内部的行人风环境。街谷内行人呼吸高度处的气流对局地条件很敏感,树木的局部配置（空间簇集、密度）将引起强烈的空间变化。在既有城市建筑布局条件下,如何通过谨慎的景观设计,利用树木等城市绿化措施有效地改善城市的行人风环境,缓解污染扩散、疾病传播等问题,本文的方法可提供一定的参考。     

基于DEM和地统计的森林资源空间格局分析--以武夷山山区为例

鉴于森林资源空间分布与地形的紧密相关,本文采用DEN与地统计学方法探讨武夷山森林主要树种蓄积量的空间格局。研究结果表明,地带性植被阔叶树由于受人为干扰趋向分布于高海拔、坡度陡的山地:杉木林在长年人工经营的过程中,逐渐占据林分的首要地位,因此主要分布在500m以下的低山丘陵:马尾松适生性较广,且明显倾向分布于(半)阳坡。地统计的分析发现各向同性的块金方差与基台值之比为0.5,即随机因素和空间自相关引起的空间异质性均各占一半,变程约24kin。在各向异性的变异中,0°和45°方向的曲线变化规律一致,随着空间距离的增加,半变异函数上升,达到基台值附近趋平,但90°和135°方向上,曲线的斜率则急剧上升。结果表明,虽然武夷山林区本身优越的地带性自然条件带来丰富多样的森林资源,地形也深刻影响着树木的天然分布,但是源于森林经营和管理的人为选择,较大程度地改变了武夷山林区森林天然的空间分布格局,森林蓄积量的空间异质性并不十分明显。由此可推断,随着城镇和经济的不断发展,该区域植被的水平地带性特征将愈加模糊,森林资源的异质性也将日渐减少。     

Forest structure of a maple old-growth stand: a case study on the Apennines mountains (Southern Italy)

In Europe, very small forest areas can be considered to be old-growth, and they are mainly located in Eastern Europe. The typical structures of old growth forests infrequently occur in Mediterranean mountainous environments, since they have been affected by human activities for centuries. This study focused on a remote and almost pure Italian maple stand located in southern Italy, which has not been managed for long time due to its inaccessibility. The effects of natural evolution on the forest stand were evaluated through the analysis of the spatial and chronological structure and the regeneration patterns, then estimating the amounts and quality of deadwood occurrence. Across the whole stand, all the trees with DBH (diameter at breast height) larger than 50 cm (LLT, large living trees) were measured (DBH and height) and age was also determined through a dendrochronological approach. The diameters observed ranged between 50 and 145 cm with ages of 120 to 250 years. The Latham index calculated for trees within the sample plot highlighted a multilayered canopy with a dominant layer of large living trees (age > 120 years). The size-class distribution of stems had a reverse-J shape, and basal area was 52 m² ha^-1. Deadwood was exclusively constituted by standing dead trees and CWD and its volume was on average 31 m³ ha^-1.Pure Italian maple forests are generally rare in Europe, and it was unexpected to find a forest stand characterized by a so complex structure with old growth attributes. The study of complex forest stand, even if small, could give precious information on the forest evolution, clarifying also diverse auto-ecological traits of tree species that usually are not common in our forests.     

Responses of soil fauna structure and leaf litter decomposition to effective microorganism treatments in Da Hinggan Mountains, China

Microorganisms are nutritious resources for various soil fauna.Although soil fauna grazing affects microorganism composition and decomposition rate,the responses of soil fauna and leaf litter decomposition to added microorganism is little understood.In this study,in the coniferous and broad-leaved mixed forest of Tahe County in the northern Da Hinggan Mountains,China,three sampling sites(each has an area of 10 m2) were selected.The first two sites were sprinkled with 250 times(EM1) and 1000 times(EM2) diluted effective microorganism(EM) preparations evenly,and the third site was sprinkled with the same volume of water as a control site.The responses of soil fauna structure and leaf litter decomposition to EM treatment were conducted during three years.The results revealed that EM treatment resulted in significant increase of soil organic matter.The number of soil fauna in the EM1 and EM2 sites increased by 12.88% and 2.23% compared to the control site,and among them springtails and mites showed the highest increase.However,the groups of soil fauna in the EM1 and EM2 sites decreased by 6 and 9,respectively.And the changes in the diversity and evenness index were relatively complicated.EM treatment slowed the decomposition of broad-leaved litter,but accelerated the decomposition of coniferous litter.However,the decomposition rate of broad-leaved litter was still higher than that of coniferous litter.The results of this study suggested that the added microorganisms could help individual growth of soil fauna,and this method led to a change in the process of leaf litter decomposition.This paper did not analyze the activity of soil microorganisms,thus it is difficult to clearly explain the complex relationships among litter type,soil fauna and soil microorganisms.Further research on this subject is needed.     

牟平、乳山地区早元古代地层

牟平、乳山地区的荆山群出露较全,依其原岩性质、变质岩石组合及接触关系,自下而上划分为禄格庄组、野头组和陡崖组,各组根据岩石组合特征可进一步划分为上、下两段。荆山群是继晚太古代胶东群沉积之后,在以升降运动为主且较稳定的浅海陆棚环境中形成的,经历过中压相系高角闪岩相区域变质。其成岩年龄为１８４８～２４８４Ｍａ。年代地层划归下元古界。     

ANALYSIS OF BIOLOGICAL GEOCHEMISTRY OFCHEMICAL ELEMENTS IN Betula ermanii FORESTIN CHANGBAI MOUNTAINS,CHINA

匠he 乙hangbal Mountains Nature Reserve Is a al，1982ZHUANGetal，1989；HUANGetal，1982；state－grade nature rese。e，also Is a component part of LIU，1993；LIU，1988； ZHU et al，1991）．Most ofreserve net of International blosphere．its area Is large，their research works focus on the chemical structure andthe natural environment and ecosystem lsprotected material cycling In alpine tundra zone andcompletely．the natural elements and ecological envl－broadleatKorean pine fores…     

Influence of human pressure on forest resources and productivity at stand and tree scales: The case study of Yunnan pine in SW China

This paper examines human impact on stands and individual trees of Pinus yunnanensis growing near the small mountain villages of Pianshui and Yangjuan in southwestern Sichuan Province,China.In an effort to assess whether use of these forests was sustainable,we examined the effects of human use in two ways.First,we directly measured the effect of cutting branches,for fuel and fodder,on tree growth.We hypothesized that branch cutting would negatively impact tree growth.We established 12 plots on four hills and compared 14 pairs of trees,one tree in each pair with an apparently full crown and the other with a considerable portion of the crown removed.Second,we assessed stand and tree properties over a 500 m elevation gradient above the villages where we hypothesized that as elevation increases,stand and tree properties should show fewer human impacts.Although extensive branch cutting reduced the live crown,tree height and diameter,compensatory processes likely enabled trees to recover and to add basal area increments(BAIs) similar to those added by trees with full crowns.Trees and stands close to villages showed less growth and lower basal areas,respectively,than stands and trees at intermediate or distant elevations from villages.Areas relatively close to the villages showed considerable effects of human-related disturbances such as branch cutting,grazing,tree and shrub removal,losses of litter,and human and animal trails.Such areas had increased soil erosion and often loss of the ‘A’ horizon.Stands close to villages had younger trees,lower stand basal areas,smaller basal area increments,and more stumps.Our results suggest an increasingly vulnerable interface between occupants of these two villages and their surrounding forests.