Paleoecology of two marine oxygen isotope stage 7 sites correlated by the Sheep Creek tephra, northwestern North America

Pollen analysis at two sites, correlated by the presence of the 190,000 yr-old Sheep Creek tephra, documents fluctuations in vegetation and climate consistent with this date and indicates that the records span marine oxygen isotope stage 7 and the stage 6/7 transition. Dawson Cut, near Fairbanks, Alaska, provides a 5.2-m-long pollen record of interglacial boreal forest succeeded by shrub tundra and then forest/tundra. Ash Bend, Stewart River, central Yukon, provides a 9.5-m-long record of interglacial boreal forest succeeded by forest/tundra, shrub tundra, and herbaceous tundra. The replacement of forest at both sites by more open or tundra vegetation indicates warm interglacial conditions giving way to cold and arid climate. It is not clear whether stage 7 was warmer than the present. The warm-cool-warm climate oscillation evident at both sites may correlate to Lake Baikal substages 7a, 7b, and 7c. Sheep Creek tephra fell on forest/tundra vegetation.     

CLIMATIC ZONES OF THE PLEISTOCENE

Although much emphasis has been placed on the effects of Pleistocene ice sheet as the determinant of Pleistocene climate in Europe, only through the combined evaluation of all useful climatic indicators can Pleistocene climatic zones be differentiated. The relationships between a paleoclimatically determined snowline and polar treelines and extent of loess deposition, determined stratigraphically, botanically, and morphologically, also indicate climatic conditions of Pleistocene Europe. Five great climatic-morphologic and plant-geographic zones, namely, forest-rubble tundra, forest tundra, loess tundra, loess steppe, and loess-forest steppe, may be distinguished. Considering these factors and their characteristics and spatial and temporal extents, it is possible to chronologically present the differentiation of the Würm glacial period independent of the retardation effect of ice sheets. --G. E. Denegar.     

The Holocene thermal maximum and late-Holocene cooling in the tundra of NE European Russia

To investigate the Holocene climate and treeline dynamics in the European Russian Arctic, we analysed sediment pollen, conifer stomata, and plant macrofossils from Lake Kharinei, a tundra lake near the treeline in the Pechora area. We present quantitative summer temperature reconstructions from Lake Kharinei and Lake Tumbulovaty, a previously studied lake in the same region, using a pollen–climate transfer function based on a new calibration set from northern European Russia. Our records suggest that the early-Holocene summer temperatures from 11,500 cal yr BP onwards were already slightly higher than at present, followed by a stable Holocene Thermal Maximum (HTM) at 8000–3500 cal yr BP when summer temperatures in the tundra were ca. 3°C above present-day values. A Picea forest surrounded Lake Kharinei during the HTM, reaching 150 km north of the present taiga limit. The HTM ended with a temperature drop at 3500–2500 cal yr BP associated with permafrost initiation in the region. Mixed spruce forest began to disappear around Lake Kharinei at ca. 3500 cal yr BP, with the last tree macrofossils recorded at ca. 2500 cal yr BP, suggesting that the present wide tundra zone in the Pechora region formed during the last ca. 3500 yr.     

Post‐glacial vegetation reconstruction and a possible 8200 cal. yr BP event from the low arctic of continental Nunavut,Canada

Climate models suggest that the global warming during the early to mid‐Holocene may have partly resulted from the northward advance of the northern treeline and subsequent reduction of the planetary albedo. We investigated the Holocene vegetation history of low arctic continental Nunavut, Canada, from a radiocarbon‐dated sediment core from TK‐2 Lake, a small‐lake ca. 200 km north of the limit of the forest‐tundra. The pollen and loss‐on‐ignition data indicate the presence of dwarf shrub tundra in the region since the beginning of organic sedimentation at ca. 9000 cal. yr BP with dominance of Betula, especially since 8700 cal. yr BP. At 8100–7900 cal. yr BP the dominance of the shrub tundra was punctuated by a transient decline of Betula and coincident increases of Ericaceae undiff., Vaccinium‐type, and Gramineae. This suggests an abrupt disturbance of the Betula glandulosa population, approximately simultaneously with the sudden 8200 cal. yr BP event in the North Atlantic. However, in the absence of other sites studied in the area, linkage to the 8200 cal. yr BP event remains tentative. The lack of any evidence of forest‐tundra in the region constrains the northern limit of the mid‐Holocene advance of the forest‐tundra boundary in central northern Canada. Consequently, our results show that the climate models imposing a mid‐Holocene advance of the limit of the forest‐tundra to the arctic coast of Canada may have overestimated the positive climatic feedback effects that can result from the replacement of tundra by the boreal forest. Copyright © 2003 John Wiley & Sons, Ltd.     

The importance of plant macrofossils in the reconstruction of Lateglacial vegetation and climate: examples from Scotland,western Norway,and Minnesota,USA

Lateglacial and early Holocene (ca 14–9000 ¹⁴C yr BP; 15–10,000 cal yr BP) pollen records are used to make vegetation and climate reconstructions that are the basis for inferring mechanisms of past climate change and for validating palaeoclimate model simulations. Therefore, it is important that reconstructions from pollen data are realistic and reliable. Two examples of the need for independent validation of pollen interpretations are considered here. First, Lateglacial-interstadial Betula pollen records in northern Scotland and western Norway have been interpreted frequently as reflecting the presence of tree-birch that has strongly influenced the resulting climate reconstructions. However, no associated tree-birch macrofossils have been found so far, and the local dwarf-shrub or open vegetation reconstructed from macrofossil evidence indicates climates too cold for tree-birch establishment. The low local pollen production resulted in the misleadingly high percentage representation of long-distance tree-birch pollen. Second, in the Minnesotan Lateglacial Picea zone, low pollen percentages from thermophilous deciduous trees could derive either from local occurrences of the tree taxa in the Picea/Larix forest or from long-distance dispersal from areas further south. The regionally consistent occurrence of low pollen percentages, even in sites with local tundra vegetation, and the lack of any corresponding macrofossil records support the hypothesis that the trees were not locally present. Macrofossils in the Picea zone represent tundra vegetation or Picea/Larix forest associated with typically boreal taxa, suggesting it was too cold for most thermophilous deciduous trees to grow. Any long-distance tree pollen is not masked by the low pollen production of tundra and Picea and Larix and therefore it is registered relatively strongly in the percentage pollen spectra.Many Lateglacial pollen assemblages have no recognisable modern analogues and contain high representations of well-dispersed ‘indicator’ taxa such as Betula or Artemisia. The spectra could have been derived from vegetation types that do not occur today, perhaps responding to the different climate that resulted from the different balance of climate forcing functions then. However, the available contemporaneous plant-macrofossil assemblages can be readily interpreted in terms of modern vegetation communities, suggesting that the pollen assemblages could have been influenced by mixing of locally produced pollen with long-distance pollen from remote vegetation types that are then over-represented in situations with low local pollen production. In such situations, it is important to validate the climate reconstructions made from the pollen data with a macrofossil record.     

论青藏高原苔原——成因、分布与分类的研究

通过分析青藏高原形成苔原的综合因素,论证了高原边缘高山和高原本体都发育有苔原,是全球中纬度独特的大陆苔原带.在划分高山苔原与山原苔原的基础上,将山原苔原进一步划分为湿地苔原、草甸苔原、草原苔原和荒漠苔原等4类,丰富了全球苔原结构.阐明青藏高原既有区域环境的独特性,也有适应全球地带性规律的统一性,使高原自然地带的定位更为准确,也有益于高原环境垂直带理论的完善.     

长白山自然保护区历史森林景观的初步重建

本文在对环境因子(包括年均温、年降水量、海拔高度、坡度和坡向)进行空间表达的基础上,建立了长白山自然保护区当前森林景观垂直带(包括阔叶红松林、云冷杉林、岳桦林和苔原)与这些环境因子间的逻辑斯蒂回归模型,然后利用1997年各环境因子的空间数字面来反推1975年森林景观带的空间分布,并根据1975年MSS遥感影像计算机监督分类的结果,用Kappa指数对模型预测结果进行验证。结果表明,苔原、岳桦林、云冷杉林和阔叶红松林1975年的预测分布面积分别为7243.83hm2、6517.08hm2、125570.16hm2和34264.80hm2,与遥感分类结果相比,苔原和云冷杉林的面积分别减少了29.04%和3.05%;岳桦林和阔叶红松林的面积分别增加了246.45%和8.6%。用逻辑斯蒂回归模型可以较好地预测苔原在保护区内的分布范围,同时在一定程度上也能预测阔叶红松林的分布范围,但是对岳桦林和云冷杉林分布范围的预测结果较差。尽管如此,作为一种方法性的探讨,本文可以为森林景观空间分布和环境因子变化的关系以及森林景观对气候变化的反应研究提供一种新思路。     

The pollen complex from postglacial sediments of the Laptev Sea as a bioindicator

The first results of comparison of palynological analysis (pollen of terrestrial plants), SEM analysis of pollen morphology, and radiocarbon age dating (AMS14C) of sediments of the eastern shelf of the Laptev Sea show that the diverse taxonomic composition of pollen spectra provides an integrated idea of the vegetation and climate of the region over 11.2 calendar kiloyears. It is found that phases of the tree and shrub vegetation development (maxima of pollen of Betula sect. Nanae and Pinus s/g Haploxylon) correspond to the warm epochs in the Holocene. It is obvious that birch phytocoenoses first settled in the southern tundra subzone with increasing temperature, and then coniferous communities of forest tundra. An occurrence of pollen of shrubby birches (Nanae) suggests compliance of permafrost landscapes with cold climate conditions, i.e., with an annual average temperature of ?2°C and amount of precipitation of less than 500 mm. Owing to a progressive increase in summer temperatures, dwarf cedar and pine communities advanced toward the seashore. The SEM analysis results show that a significant proportion of regional coniferous pollen belongs to representatives of Pinus pumila (Pall.) and P. sylvestris L. In addition, the SEM study of the exine of Pinus sylvestris L. and P. pumila (Pall.) Regel pollen grains confirmed polymorphism in coniferous pollen. According to the inverse relationship between climate and vegetation, frequent climate fluctuations that are typical of progressive and differential postglacial transgression were revealed. An increase in arboreal pollen transfer onto the shelf later than 9.1 cal. ka coincides with the time of forest boundary migration to the north due to the warming of the Earth’s climate. At that time, the tundra vegetation was replaced by forest-tundra vegetation, the maximum stage of sea transgression began, and there appeared a trend of increasing temperature and moisture.     

Lithological and biochemical properties in sediments of Lama Lake as indicators for the late Pleistocene and Holocene ecosystem development of the southern Taymyr Peninsula, Central Siberia

In order to evaluate the sensitivity of aquatic and terrestrial ecosystems to climatic changes, lithological (sediment structure and color, grain size, physical properties) and biochemical (TOC, TOC/TN, δ¹³C of TOC and carbonates) investigations were carried out on an 11.12 m-long sediment core from Lama Lake (Central Siberia, Putorana Plateau). According to the pollen data, the sequence represents the termination of the Pleistocene, and the entire Holocene. It is composed of highly terrigenous and stratified clays and silts. Sediment structure, grain-size distribution, carbonate contents and physical properties of the sediment indicate that glaciers were present in the catchment area of Lama Lake during the period Oldest Dryas to AllerØd. For the same time period, δ¹³C values of TOC give indications of a perennial ice cover. Since the AllerØd, organic matter accumulation has increased, caused by an increasing input of land vegetation and aquatic primary production as revealed by relations TOC contents, TOC/TN ratios and δ¹³C values of TOC. During the Holocene climatic optimum, in late Preboreal and Boreal times, biogenic primary production in Lama Lake reached its maximum and the vegetation in the catchment area changed from grassy tundra to a dense forest. From the Atlantic period to the present, small variations in δ¹³C values of TOC and TOC contents are probably related to the location of Lama Lake on the border between grass and wooded steppe, leading to responses of environmental conditions to even small changes in climate.     

The Eemian interglacial and its termination

The development and termination of the Eemian interglacial is important because it may serve as a model showing how the present warm period might end in the event of no anthropogenic impact. The most important methods for studying the Eemian are outlined and critically evaluated. In spite of interpretation and dating problems, the various proxy data seem consistent enough to allow the conclusion that some 120,000 years ago the warm Eemian climate deteriorated rapidly and drastically. The forest vegetation in West Europe was replaced by a tundra type vegetation, and within 5000 or 10,OOO years the volume of continental ice grew to at least double the present volume, corresponding to a sea level 65 m, perhaps 90 m, below that of today. There is considerable disagreement between sea level estimates deduced from geological evidence and from benthic foraminifera oxygen isotope records.     

北方针叶林对气候变化响应的研究进展

北方针叶林地处北半球中高纬度地区,增温剧烈,是受气候变化影响最显著的地区之一.据预测,在未来气候变化情景下,北方针叶林将向北迁移,且可能完全移出我国,从而将对气候系统及全球碳收支产生重要的反馈作用.因此,北方针叶林在气候变化与全球碳收支中具有十分重要的作用.从气候变化的影响及碳循环研究方面综述了北方针叶林近年来的最新研究进展,探讨了未来北方针叶林拟重点加强的研究领域,为深入理解北方针叶林对气候变化的响应、制定适宜的气候变化对策提供参考依据.     

Soil carbon stock of the Central-East Asia in the climate warming

Water samples from the Wujiang River, a typical karst river system, were analyzed for major ion concentrations and δ^34S values of dissolved sulfate in order to identify the sources of sulfate, quantify the sulfate export flux and understand the role of sulfur cycling in chemical weathering rate of carbonate. Spatial variations in sulfate concentration and sulfur isotopic composition of tributaries over the catchment area are obvious, allowing to decipher S sources between rocks and atmosphere. According to the variations in sulfate concentration and isotopic composition, it is inferred that sulfate ions in the upper-reach river waters may have three sources, rain water, sulfate resultant from oxidation of pyrite in coal, and sulfate from sulfide deposits. In the lower reaches, the S isotopic composition of the samples lies mainly on a mixing trend between evaporite sulfate and rainwater sulfate, the contribution of sulfate from oxidation of pyrite being lesser. A pronounced seasonal variation in both content and isotopic composition of sulfate characterizes the Wujiang River. The average sulfate concentration of the waters is 0.65 mmol/L in winter, 0.17 mmol/L higher than that in summer. River water δ^34S values range from -15.7‰ to 18.9‰ in winter, while the δ^34S values of river waters in summer vary to a lesser extent than in winter, from -11.5‰ to 8.3‰. The δ^34S values of the main stream range from -6.7‰ to -3.9‰ in summer, averaging 3‰ lower than in winter. This indicates that in summer, when the discharge increases, the contribution of a source enriched in light isotopes to the atmosphere or the oxidation of pyrite in coal is more important.     

Chemical and isotopic characteristics of weathering and nitrogen release in non-glacial drainage waters on Arctic tundra

Soil-water interactions in coastal tundra soils are a potential source of nutrients for surrounding fjordal and coastal ecosystems. Changes in water chemistry and stable isotope composition from three streams in west Spitsbergen were examined to assess the sources and losses of nitrogen, sulfur and carbon in thin organic tundra soils overlying sediments. Studies were undertaken from snowmelt (mid June) through to the end of the summer (September) in both 2001 and 2002. Drainage water chemistry was dominated by the solution of Ca-Mg carbonates with δ¹³C values in the waters being uncharacteristically high (approx. −2‰ at the end of the season), reflecting a largely open system in which the CO₂ is derived equally from the atmosphere and plant/soil sources. Early melt waters had δ³⁴S values dominated by sea salt reflecting the close proximity to the ocean. However, as the season progressed the marine influence lessened. Extrapolation of the data suggests that the origin of non-sea salt δ³⁴S was the oxidation of reduced sulfur from coal particles in the subsoil. Concentrations of inorganic N in stream waters were generally very low. However, values were found to increase as the season progressed, possibly through increased microbial activity in the soil and the early senescence of tundra plants reducing demand. Dual isotope analysis of δ¹⁵N and δ¹⁸O suggested that the vast majority of snow-pack was assimilated by the soil microbial biomass before being released, recycled and lost to drainage waters. Organic N concentrations in drainage waters were generally equal to or greater than losses of inorganic N from tundra soils. The study demonstrated the effectiveness of stable isotope data for understanding biogeochemical cycling and soil-water interactions in tundra ecosystems. The implications of the results are discussed in relation to climate warming.     

Plant and insect fossils at Norwood in south-central Minnesota: A record of late-glacial succession

The Norwood site in Sibley Co., Minnesota, contains 1.6 m of silt resting on till and overlain by peat. The base of the peat has been radiocarbon dated at 12,400 ± 60 and the top at 11,200 ± 250 yr B.P. The pollen, plant macrofossils, and insect remains in the basal silt consist of boreal species inhabiting open environments, but not tundra. No modern analogue exists for the insect assemblage, which includes elements of boreal forest, tundra-forest, and western affinities. The transition from an unstable open environment to a stable coniferous forest is reflected by both plant and insect fossils and is interpreted as a successional rather than a climatic event. During this time of significant biologic change, the climate is inferred to have been relatively uniform, with temperatures similar to those presently existing in the boreal forest south of the tundra-forest transition zone. The geologic and ecologic succession at Norwood is generally similar to that presently associated with ice stagnation of the Klutlan Glacier in the Yukon Territory. Localized successional sequences similar to those at Norwood are conceived to have occurred repeatedly during the melting of the Laurentide ice, and thus the proposed model has potentially broad application to the interpretation of late-glacial sequences.     

Vegetation and climate during the Weichselian Early Glacial and Pleniglacial in the Niederlausitz,eastern Germany — macrofossil and pollen evidence

Cryoturbated organic beds and channel fills, intercalated with sandy and gravelly fluvial units, have been studied in an opencast brown‐coal mine near Nochten (Niederlausitz), eastern Germany. The fluvial–aeolian sequence covers parts of the Early, Pleni‐ and Late‐glacial. The detailed chronology is based on 11 radiocarbon and 12 OSL dates, covering the period between ca. 100 kyr and 11 kyr BP. Basal peat deposits are correlated with an Early Weichselian interstadial. During this period boreal forests were present and minimum mean summer temperatures were > 13°C. Early Pleniglacial deposits are absent. The Middle and Late Pleniglacial environments were treeless and different types of tundra vegetation can be recognised. Minimum mean summer temperatures varied between 10 and 15°C. Vegetation and climate is reconstructed in detail for the periods around 34–38 kyr BP and 24–25 kyr BP. Around 34–38 ka, a mixture between a low shrub tundra and a cottongrass tussock–subshrub tundra was present. The botanical and sedimentological data suggest that from the Middle to the Late Pleniglacial, the climate became more continental, aridity and wind strength increased, and the role of a protecting winter snow cover decreased. A sedge–grass–moss tundra dominated around 24 and 25 kyr BP. Copyright © 2001 John Wiley & Sons, Ltd.     

Weichselian and Holocene palaeoenvironmental history of the Bol'shoy Lyakhovsky Island, New Siberian Archipelago, Arctic Siberia

Cryolithological, ground ice and fossil bioindicator (pollen, diatoms, plant macrofossils, rhizopods, insects, mammal bones) records from Bol'shoy Lyakhovsky Island permafrost sequences (73°20′N, 141°30′E) document the environmental history in the region for the past c. 115 kyr. Vegetation similar to modern subarctic tundra communities prevailed during the Eemian/Early Weichselian transition with a climate warmer than the present. Sparse tundra‐like vegetation and harsher climate conditions were predominant during the Early Weichselian. The Middle Weichselian deposits contain peat and peaty soil horizons with bioindicators documenting climate amelioration. Although dwarf willows grew in more protected places, tundra and steppe vegetation prevailed. Climate conditions became colder and drier c. 30 kyr BP. No sediments dated between c. 28.5 and 12.05 ¹⁴C kyr BP were found, which may reflect active erosion during that time. Herb and shrubby vegetation were predominant 11.6–11.3 ¹⁴C kyr BP. Summer temperatures were c. 4 °C higher than today. Typical arctic environments prevailed around 10.5 ¹⁴C kyr BP. Shrub alder and dwarf birch tundra were predominant between c. 9 and 7.6 kyr BP. Reconstructed summer temperatures were at least 4 °C higher than present. However, insect remains reflect that steppe‐like habitats existed until c. 8 kyr BP. After 7.6 kyr BP, shrubs gradually disappeared and the vegetation cover became similar to that of modern tundra. Pollen and beetles indicate a severe arctic environment c. 3.7 kyr BP. However, Betula nana, absent on the island today, was still present. Together with our previous study on Bol'shoy Lyakhovsky Island covering the period between about 200 and 115 kyr, a comprehensive terrestrial palaeoenvironmental data set from this area in western Beringia is now available for the past two glacial–interglacial cycles.     

Late Saalian and Eemian palaeoenvironmental history of the Bol''shoy Lyakhovsky Island (Laptev Sea region, Arctic Siberia)

Palaeoenvironmental records from permafrost sequences complemented by infrared stimulated luminescence (IRSL) and [Formula: See Text]Th/U dates from Bol'shoy Lyakhovsky Island (73°20'N, 141°30'E) document the environmental history in the region for at least the past 200 ka. Pollen spectra and insect fauna indicate that relatively wet grass-sedge tundra habitats dominated during an interstadial c. 200-170 ka BP. Summers were rather warm and wet, while stable isotopes reflect severe winter conditions. The pollen spectra reflect sparser grass-sedge vegetation during a Taz (Late Saalian) stage, c. 170-130 ka BP, with environmental conditions much more severe compared with the previous interstadial. Open Poaceae and Artemisia plant associations dominated vegetation at the beginning of the Kazantsevo (Eemian) c. 130 ka BP. Some shrubs (Alnus fruticosa, Salix, Betula nana) grew in more protected and wetter places as well. The climate was relatively warm during this time, resulting in the melting of Saalian ice wedges. Later, during the interglacial optimum, shrub tundra with Alnus fruticosa and Betula nana s.l. dominated vegetation. Climate was relatively wet and warm. Quantitative pollen-based climate reconstruction suggests that mean July temperatures were 4-5°C higher than the present during the optimum of the Eemian, while late Eemian records indicate significant climate deterioration.     

Geology,palynology, and climatic significance of two pre-Pinedale Lake sediment sequences in and near Yellowstone National Park

Pollen analysis of a section of lake sediments at Grassy Lake Reservoir indicates a vegetational sequence changing from tundra, to spruce-fir-pine forest, to pine forest, to tundra at the top. Pollen analysis of a section of lake sediments on Beaverdam Creek indicates a tundra vegetation at the base, followed by a brief episode of spruce-fir forest and a return to a tundra vegetation at the top. The analyses of both sections suggest a cold to cool to cold climatic sequence, interpreted as interstadial in character. However, differences suggest that they represent separate interstadials. Pinedale Till disconformably overlies the lake deposits at Grassy Lake Reservoir. The upper sediments contain wood ¹⁴C dated at >42,000 yr; the lowermost interfinger with till shown to be more than about 70,000 yr old. The deposits at Beaverdam Creek grade upward into proglacial Pinedale deposits, contain an ash that is probably about 70,000 yr old near their base, and rest comformably on gravel that grades down into lake sediments containing wood debris suggestive of an older climatic amelioration. We conclude that the warmest part of the interstadial at Grassy Lake Reservoir is probably more than 70,000 yr old, and that the warmest part of the interstadial analyzed at Beaverdam Creek is slightly younger than 70,000 yr old.     

Late Quaternary vegetation in the Southwestern Columbia Basin,Washington

A 33,000-yr pollen record from Carp Lake provides information on the vegetation history of the forest/steppe border in the southwestern Columbia Basin. The site is located in the Pinus ponderosa Zone but through much of late Quaternary time the area was probably treeless. Pollen assemblages in sediments dating from 33,000 to 23,500 yr B.P. suggest a period of temperate climate and steppe coinciding with the end of the Olympia Interglaciation. The Fraser Glaciation (ca. 25,000–10,000 yr B.P.) was a period of periglacial steppe or tundra vegetation and conditions too dry and cold to support forests at low altitudes. Aridity is also inferred from the low level of the lake between 21,000 and 8500 yr B.P., and especially after about 13,500 yr B.P. About 10,000 yr B.P. Chenopodiineae and other temperate taxa spread locally, providing palynological evidence for a shift from cold, dry to warm, dry conditions. Pine woodland developed at the site with the onset of humid conditions at 8500 yr B.P.; further cooling is suggested at 4000 yr B.P., when Pseudotsuga and Abies were established locally.     

吉林辉南二龙湾玛珥湖12.0 ka B.P.以来孢粉记录与气候变化

东北二龙湾玛珥湖孢粉记录显示,12.0 ka B.P.以来,该区植被和气候经历了以下变化：11.76～9.81 ka B.P. 发育针阔叶混交林,气候冷干。9.81～4.11 ka B.P.为落叶阔叶林植被景观,前期（9.81～8.86 ka B.P.）温干,为增温期;后期（8.86～4.11 ka B.P.）暖湿,为气候适宜期。4.11 ka B.P.至今,再次发育针阔叶混交林, 其中1.62 ka B.P.以前,气候温凉;之后湿度下降,气候凉干。研究表明:YD（11.56～10.21 ka B.P.）事件在二龙湾玛珥湖孢粉资料中有突出记录,4.0 ka B.P.左右开始的降温现象在东北东部地区广泛出现。