New Occurrence of the Early Cambrian Chengjiang Fauna in Haikou, Kunming, Yunnan Province,and Study on Trilobitoidea

This paper gives a brief report on the new occurrence of the Early CambrianQiongzhusi' an Chengjiang fauna in the Haikou area, Kunming and its assemblage feature andthe stratigraphic section of the horizon yielding these fossils. Eight genera and 8 species ofTrilobitoidea are described here, embracing Leanchoilia asiatica Luo et Hu (sp. nov.), Yohoiasinensis Luo et Hu (sp. nov.), Zhongxinia speciosa Luo et Hu (gen. et sp. nov.), Xandarellaspectuculum Hou et al., Dianchia mirabilis Luo et Hu (gen. et sp. nov.), Kuamaia lata Hou,Retifacies longispinus Luo et Hu (sp. nov.) and Sinoburius lunaris Hou et al.     

A New Platy-armored Worm from the Early Cambrian Chengjiang Lagerstatte, South China

The Lower Cambrian Chengjiang Lagerstatte provides a good window to explore the origin and radiation of early bilaterians. Here we describe a netted sclerite-bearing worm Tabelliscolex hexagonus gen. et sp. nov., and tentatively assign it to palaeoscolecidans of priapulids. The cuticle of the animal is covered with two kinds of platy sclerites which are constructed by hexagonally arranged tubercles. Similar structures of the sclerites can be seen on some Cambrian palaeoscolecidans and lobopods, so, this new species is critical for understanding the relationships between lobopods and palaeoscolecidans.     

寒武纪早期澄江生物群小细丝海绵属新材料

文章描述了云南寒武纪早期澄江生物群小细丝海绵属一新种——螺旋小细丝海绵（Leptomitella spiralis sp. nov.）。新种 具有横向骨针束围绕海绵体呈螺旋状排列的典型特征，明显区别于属内其他种；此外，在外部形态和骨针类型上与属内其他种 也存在较大差异。新种的发现增加了Leptomitella种间形态结构的多样性，也为海绵动物的早期演化及寒武纪海绵动物辐射演 化研究提供新的化石资料。     

寒武纪早期澄江生物群小细丝海绵属新材料

文章描述了云南寒武纪早期澄江生物群小细丝海绵属一新种--螺旋小细丝海绵（Leptomitella spiralis sp. nov.）。新种 具有横向骨针束围绕海绵体呈螺旋状排列的典型特征,明显区别于属内其他种;此外,在外部形态和骨针类型上与属内其他种 也存在较大差异。新种的发现增加了Leptomitella种间形态结构的多样性,也为海绵动物的早期演化及寒武纪海绵动物辐射演 化研究提供新的化石资料。     

云南寒武纪早期澄江生物群中原始海绵科似斜纹海绵属新种

本文报道澄江生物群似斜纹海绵属Paradiagoniella的两个新种：橄榄形似斜纹海绵P.oliviformis sp. nov. 和锥形似斜纹海绵P. conicasp. nov. 。两个新种具有似斜纹海绵属的基本特征：体壁薄;由排列不规则、分级的十字骨针为基底构成骨骼;除十字骨针外,还穿插排列有五射针、六射针、弯弓形和针形双尖单轴针等。然而与模式种不同,两个新种十字骨针分级不明显;同时,两个新种在外形、骨针组成、最大十字骨针射长比等方面,与本属的其他种都存在明显差异。新材料的发现增加了寒武纪早期原始海绵物种多样性,为研究海绵动物早期起源演化提供了新信息。     

New Record of Palaeoscolecids from the Early Cambrian of Yunnan, China

A new palaeoscolecid, Guanduscolex minor Hu, Luo et Fu gen. et sp. nov., with preserved soft parts of introvert and intestines comes from the Lower Cambrian Guanshan fauna of Yunnan, South China. Microstructural details of the cuticle revealed by scanning electron microscopy （SEM） show that each annulation bears three rows of plates and each plate bears 9-10 marginal and 4-5 central nodes. This discovery sheds new light on the relationships and evolutionary pathway of the palaeoscolecids and other early priapulids.     

A New Arthropod, Guangweicaris Luo, Fu et Hu gen. nov. from the Early Cambrian Guanshan Fauna, Kunming, China

The Guanshan Fauna is a soft-bodied fauna dominated by arthropods （including trilobites, trilobitoides, Tuzoia, Isoxys, and bradorids） in association with priapulids, brachiopods, anomalocaridids, vetulicoliids, sponges, chancellorids, and echinoderms. This paper reports and describes a new arthropod from the yellowish green mudstone at the lower part of the Wulongqing Formation, Canglangpuan Stage, Lower Cambrian in Kunming, Yunnan, China. The stratigraphic and geographic distribution, classification, fossil preservation, life style of this new arthropod and comparisons with other fossil arthropods are also discussed in details. The discovery and research of the non-mineralized arthropod, Guangweicaris Luo, Fu et Hu gen. nov. from the Guanshan Fauna adds new members to the taxonomic list and provides new information to the evolution of early arthropods. Furthermore, this study would shed new light into the ＂Cambrian Explosion＂ and the evolution of early life.     

A New Platy‐armored Worm from the Early Cambrian Chengjiang Lagerstätte,South China

Abstract The Lower Cambrian Chengjiang Lagerstätte provides a good window to explore the origin and radiation of early bilaterians. Here we describe a netted sclerite‐bearing worm Tabelliscolex hexagonus gen. et sp. nov., and tentatively assign it to palaeoscolecidans of priapulids. The cuticle of the animal is covered with two kinds of platy sclerites which are constructed by hexagonally arranged tubercles. Similar structures of the sclerites can be seen on some Cambrian palaeoscolecidans and lobopods, so, this new species is critical for understanding the relationships between lobopods and palaeoscolecidans.     

Isotopic dating of the Chengjiang Fauna-bearing horizon in Central Yunnan Province, China

Twenty black shale samples, which are free from the influence of weathering, were collected from the Chengjiang Fauna-bearing horizon, central Yunnan Province, yielding a Pb-Pb isochron age of 534±60 Ma. Although this age is younger than both the Rb-Sr isochron age and 40Ar-39Ar age, it should represent the lower isotopic age limit of the Chengjiang Fauna.     

澄江生物群分子在贵州遵义牛蹄塘组发现

澄江生物群是认识寒武纪生命大爆发的重要窗口,它的分布范围一直仅限于滇东地区,员近在贵州遵义松林牛蹄塘组底部页岩中发现有Naronia,Archotuba Conoidalis,Isaxys,Perpicaris,Lingulepis,Tsunyidiscus,Scenella,水母状化石(medusiform fossil),海绵动物化石Leptomitus,类似半索动物秆壁虫Rhabdopleura化石,软舌螺类Hyolithids,宏观藻类和疑源类化石Zunyiphyton perelegans Yang et Zhao,1999,Yuknessia sp.,Longenema Ding,1996,Sphaerocongregus variabilis等。其中有一部分是属于澄江生物群分子,这是云南省外澄江生物群分子的新发现。由于生物群之下40m处还产大量海绵动物、高肌虫和藻类化石等组成的松林生物群,其上又有下寒武统明心寺组古杯动物群,因此,这一生物群的发现对研究早寒武世生物的演化也具有一定的科学意义,同时,对澄江生物群的古生态学研究也具有重要意义。     

Enigmatic ribbon-like fossil from Early Cambrian of Yunnan,China

During the Late Neoproterozoic and Early Cambrian, a series of paleogeographic and paleoecological events occurred associated with deglaciations and the evolution of life. The appearance of fossils representing diverse phyla, novel body plans and complex ecologies in Ediacaran and Cambrian sedimentary successions has sparked diverse hypotheses about potential drivers for the radiation of early animals during this interval. Recently, new macroscopic fossils of carbonaceous compressions with unique features have been found in Anning, Yunnan, China. The fossils’ bodies are ribbon-shaped and bilaterally symmetric, with dense longitudinal features and transversal features. The fossils occur 0.68 m below a bentonite interlayer which has been dated 535.2 ± 1.7 Ma by Ri-xiang Zhu and his team in 2009. The relatively simply morphology of these fossils, coupled with a lack of preserved internal structures challenge efforts taxonomically identify the precursor organism and definitively ally it to a living group. However, the symmetry and unusual features of the body are analogous to members of Platyzoa. The presence of ribbon-shaped fossils in the Zhongyicun Member in Anning indicates that these organisms were at least a locally significant component of Cambrian seafloor ecosystems, and may hold important implications for our understanding of the early evolution of Bilateria.     

澄江动物群一种新的抚仙湖虫类节肢动物

描述抚仙湖虫类一新属双叶尾梁王山虫Liangwangshaniabiloba(gen etsp nov)。Li angwangshania与Fuxianhuia具有相同的体躯分化,但二者在头甲和尾甲的形状、躯体的分节数目等方面有明显差异。Liangwangshania对于诠释节肢动物早期体躯分化具有重要意义。     

New Bradoriid Arthropods from the Early Cambrian Balang Formation of Eastern Guizhou, South China

The Early Cambrian Balang Formation is comprised of mudrock and shale, which was deposited in a shelf environment in the eastern part of Guizhou, south China. The Balang Fauna, which consists of seven phyla, occurs in the middle and upper parts of the Balang Formation. Arthropods are important constituents of the Balang Fauna and include a great number of trilobites, large bivalved arthropods, and newly-discovered well-preserved bradoriid fossils. The bradoriids present include three genera and four species : Comptaluta inflata (Cheng, 1974) emend Hou et al., 2002; Comptaluta kailiensis sp. nov., and Alutella elongeta sp. nov., Aluta sp. This faunal assemblage in the Balang Formation is distinguished from the Tsunyiella Chang, 1964, Songlinella Yin, 1978 and Kunmingella Hou, 1956 assemblage which occurs in the Niutitang and Mingxinsi formations of the Yangtze Platform in middle region of Guizhou and which is earlier than the Balang Formation in age. However, this assemblage resembles the Comptaluta pik, 1968 assemblage from the Early Cambrian Heilinpu Formation in Wuding County, Yuanan Province and from the Ordian Stage of the Cambrian of Australia. The great abundance of Comptaluta pik, 1968 and overall taxonomic diversity of the Comptaluta pik, 1968 assemblage set it distinctly apart from the Alutella Kobayashi et Kato, 1951 and Aluta Hou, 1956 assemblages of the Balang Formation. Alutella Kobayashi et Kato, 1951 and Aluta Hou, 1956 also occur in the Early Cambrian Niutitang Formation of the Yangtze Platform of Guizhou. Individual Bradoriids from the Balang Formation are characterized by large size (3 mm). The discovery of new Bradoriid assemblages not only expands the group’s geographical range and assemblage affinities, but also indicates that Bradoriids migrated eastward from shallow-water to deeper-water environments during the Early Cambrian, indicating that they were capable of life in deeper-water, and adaptation to a new ecological setting.     

New Localities and Palaeoscolecid Worms from the Cambrian (Stage 4, Series 2) Guanshan Biota in Kunming, Yunnan, South China

The Guanshan Biota from the Cambrian Stage 4 Wulongqing Formation is a typical Burgess-shale type Lagerst?tte that had witnessed the Cambrian explosion. Here we report two new localities of the Guanshan Biota, which is located at Baimei village, south of Kunming and Xinglong village, Wuding County, Yunnan Province respectively. Both localities produce soft-body fossils and several new taxa, e.g. a new species of palaeoscolecidans (Palaeoscolex xinglongensis sp. nov.) reported herein. The comparative study of the new species with similar form not only indicates that there is a diversification of palaeoscolecid species in Guanshan Biota, but also strengthens the ties between the older Chengjiang Biota and the younger Kaili Biota (and also the coeval Burgess Shale community). Three paleoecological features, including high diversity, little disparity and richness of palaeoscolecid worms, are summarized as a case study to represent the differences between Guanshan and Chengjiang Biota.     

澄江化石库中的双胚层动物新知

作为观察寒武纪大爆发主幕奥秘的最佳科学窗口,澄江化石库展示了三胚层动物(即两侧对称动物)首次全面大辐射的历史产物,让学术界终于看到了已知最古老的同时包括原口类和后口类两大支系的动物谱系庞大“树冠”的主体轮廓,从而给许多较高级动物类群的起源和早期演化研究提供了十分重要的信息。然而,该化石宝库中却很少见到作为“树干”的双胚层动物(即辐射动物)的化石记录,这给“真动物”的源头探秘蒙上了阴影。尽管广布于前寒武纪末期全球浅海域的“文德生物群”一直被认为是动物界早期一次“失败的演化试验”的牺牲品,但其中一支成功延续到早寒武世的春光虫(Stromatoveris)具有叠覆状梳齿构造,很可能代表着栉水母类的一类原始祖先。文中描述新的羽毛状动物王氏澄江海笔(Chengjiangopennawangii)不同于春光虫,前者具有彼此分离的羽枝,且羽枝上保存了排列规整的珊瑚个体虫室,因而与现代八射珊瑚中海笔类的形态学特征十分一致;新的谱系分析显示,它应该代表刺胞动物门中的一个原始类群。     

New Early Cambrian Chordates from Haikou, Kunming

Two new chordates, Cathaymyrus haikouensis Luo et Hu sp. nov. and Zhongxiniscus intermedius Luo et Hu gen. et sp. nov. are studied in this paper. Both display numerous S-shaped myomeres on their trunk. C. haikouensis shows a long and slim body similar to that of Cathaymyrus diadexus Shu et al., Zhongxiniscus approaches to Myllokunmingia and Haikouichthys in the dorsal fin, but differs in the myomeres. Zhongxiniscus may be the intermediate form between Cathaymyrus and Myllokunmingia and Haikouichthys.     

含澄江动物群地层的Pb-Pb同位素定年

对含澄江动物群的地层样品进行了Pb同位素定年研究.经测定未受风化作用影响的20个黑色页岩样品给出了534±60 Ma左右的Pb-Pb等时线年龄,应代表了澄江动物群的下限年龄.该年龄较以往采用Rb-Sr和40Ar-39Ar等定年方法所获得的年龄更为年轻.     

昆明地区早寒武世关山动物群研究新进展

总结了昆明地区关山动物群自1999年命名以来开展的研究工作和取得的新进展。简要叙述了关山动物群的研究概况、地层位置、生物群组合特征和关山动物群的研究意义。在关山动物群中首次发现了古虫动物门的Vetulicola ganggtoucunenisis Luo,Fu et Hu,吐卓虫Tuzoia sp．nov．,古蠕虫Palaeoscolex sp．nov．,腕足类Heliomedusa sp．nov．及三叶形虫、海绵动物新属种。指出关山动物群是一个由多门类软躯体后生动物化石组成的伯吉斯页岩型的动物群,时代介于澄江动物群与凯里动物群（或伯吉斯页岩动物群）之间,起到了承前启后的作用,在寒武纪生命大爆发和寒武纪早期生物演化的研究中有重要意义。