Qinling Faunal Region-The Third Ordovician Faunal Region: International Correlation

The Ordovician conodont faunal provinces were previously divided into the Midcontinent and Atlantic Faunal Regions situated respectively in low and high latitudes, where warm- and cold-water type conodont faunas flourished respectively. According to the international correlation this paper proposes the third Ordovician conodont faunal region-Qinling Faunal Region, in which cold-water conodont faunas were well developed in the Early to middle Middle Ordovician and warm-water conodont faunas were well developed in the late Middle and Late Ordovician, indicating that the Qinling Region was situated in high latitudes earlier and in low latitudes later. The origin was only due to plate movement In the Qinling Region the time interval of the change of the conodont fauna from the cold- to warm-water type was 4 Ma (from 474 to 470 Ma), during which the fauna geographically spanned 40° of latitudes, with a movement velocity of nearly 1.12 m/a, indicating that the high-latitude plates were divorced and reduced i     

中国奥陶纪三叶虫生物地理

三叶虫证据表明,除北疆区的阿尔泰地体和兴安区的额尔古纳-兴安地体与西伯利亚和劳伦陆块在动物群方面紧密关联外,中国奥陶纪的所有板块和大部分地体均系当时东泛冈瓦纳的重要组成部分,在生物地理上彼此密切相关。对这些属于东泛冈瓦纳的中国区块的台地或内陆棚三叶虫动物群的综合分析研究证明,它们在特马豆克期（Tremadocian）以及凯迪晚期-赫南特期（late Katian-Hirnantian）或阿石极期（Ashgill）应归属同一生物地理区,而在弗洛期-凯迪早期（Floian-early Katian）或阿仑尼克期-卡拉多克期（Arenig-Caradoc）则可划分为两个生物地理亚区：一个由华南、塔里木板块和安南或印支地体组成,另一个包括华北板块以及滇缅马、藏南、中天山-北山地体（可能还有海南地体）。饶有趣味的是在凯迪晚期或阿石极期开始发生的生物地理亚区解体之前,有关陆块的深水相三叶虫动物群已经历了从达瑞威尔中期（mid Darriwilian）至凯迪早期的长期频繁交流以及组成逐渐趋于齐一的过程。因此深水动物群之间的属种交流和一体化进程的发生、发展和完成显然均先于浅水一侧的三叶虫。     

The Upper Ordovician of northeastern Gorny Altai: Stratigraphy and deposition environments

Comprehensive lithofacies and biofacies analysis provided constraints on the origin of Upper Ordovician clastic and carbonate deposits in northeastern Gorny Altai, which form large low-elevated flat carbonate banks located relatively close to the shore. The sediments were deposited during the Sandbian and early-middle Katian stages, according to new conodont data. Upper Ordovician sections in northeastern Gorny Altai store record of two global regressions: the early Sandbian (Vollen Lowstand) and early Katian (Frognerkilen Lowstand) events.     

Darriwilian (Middle Ordovician) graptolite faunas of the Sandia Region,southern Peru

Ordovician graptolite faunas of Peru are restricted to a short interval in the Middle to basal Upper Ordovician, found in three regions of the country. All Peruvian graptolite faunas are strongly dominated by shallow water elements of the Atlantic Faunal Realm, represented largely by Didymograptus s. str. and Aulograptus, but a number of faunal elements of the pandemic isograptid biofacies have recently been discovered in the Sandia Region of SE Peru. Peruvian graptolite faunas are reviewed and the new records from the Sandia Region are discussed in detail. The faunas from the Purumpata and Iparo members of the San José Formation range in age approximately from the Undulograptus austrodentatus Biozone to the Holmograptus lentus Biozone (early to middle Darriwilian). The faunas provide a better understanding of faunal composition and diversity in this region and help to correlate shallow water and deeper water graptolite faunas from this time interval. Biserial graptolites are rare in most samples and usually indeterminable, but a single identifiable specimen of Undulograptus austrodentatus was found, indicating a level close to the base of the Darriwilian. A number of specimens of the genera Isograptus and Arienigraptus from the Sandia Region represent pandemic graptolite faunas of the isograptid biofacies, described for the first time from this region. Copyright © 2009 John Wiley & Sons, Ltd.     

中国主要块体奥陶纪达瑞威尔期（Darriwilian）晚期—凯迪期（Katian）早期海相红层及其构造意义

根据中国主要块体奥陶纪达瑞威尔期地层学、古生物学和岩石学的特点,本文主要论述了中国9个块体,其中包括5个主要块体（扬子克拉通、中朝克拉通、塔里木克拉通、拉萨地块和喜马拉雅北坡）中达瑞威尔晚期—凯迪期早期海相红层的分布、岩性特征和古动物群。它们的岩性特征：如红色灰岩;古生物群特征：如牙形石Pygodus serra动物群,P.anserinus动物群和Hamarodus europaeus动物群,珊瑚Yohophyllum动物群,以及头足类Sinoceras chinense动物群均十分相似或相同。这就充分阐明,中华古陆块群（包括扬子、中朝和塔里木克拉通等）与冈瓦纳古陆北缘古陆块群（包括拉萨地块和喜马拉雅北坡等）之间的关系十分密切,可能在此之前它们同属于冈瓦纳古陆。这对于阐明中华古陆块群与冈瓦纳古陆之间的关系具有重要意义。此外,还将海相红层分为两类：陆棚红层和大洋红层。陆棚红层主要见于寒武纪至三叠纪,大洋红层主要见于白垩纪至现代。     

The Hirnantian δ<Superscript>13</Superscript>C Positive Excursion in the Nabiullino Section (South Urals)

The upper Sandbian, Katian, and Hirnantian complexes of conodonts in the upper Ordovician section of the western slope of the Southern Urals near the village of Nabiullino were studied. The δ¹³C positive excursion with a maximum of 3.3‰ associated with the global Hirnantian isotopic event, HICE, was fixed for the first time. This excursion shows the beginning of the Hirnantian stage in the terrigenous–carbonate section of the upper Ordovician in the Southern Urals. It coincides with the first occurrence of the Hirnantian conodont species of Gamachignathus ensifer and the conodonts of shallow-water biophacies, Aphelognathus-Ozarkodina, reflecting the global glacio-eustatic event.     

华北板块奥陶纪牙形石生物地层研究回顾及在西北缘区新进展

牙形石在以碳酸盐岩为主的华北奥陶系划分对比中占有举足轻重的地位。针对华北奥陶纪牙形石的研究已持续近半个世纪,总体上可以分为两个阶段:第一阶段自20世纪70年代到21世纪初,第二阶段为最近10年(2010—2020)。第一阶段以建立牙形石生物地层序列为目标,主要为解决石油勘探过程中地层时代的确定和地层对比的需求;第二阶段的研究以修订化石带为主,目的是与国际地层研究接轨。近年来在华北板块西北缘的工作显示,该区奥陶纪牙形石在纵向上具有显著的生态变化,可分为达瑞威尔期中期、达瑞威尔期晚期—桑比期中期、桑比期晚期—凯迪期中期3个时段。第一时段以介于北美中大陆区和北大西洋区之间的热带台地边缘型牙形石为特征;第二时段以北大西洋型为主混有少量亚澳型牙形石为特征;第三时段以亚澳型和北美中大陆型牙形石混生为特征。在华北西北缘尽可能使用广布性标准牙形石属种进行化石带厘定,共识别牙形石带12个,自下而上分别是达瑞威尔期Histiodella cf. holodentata间隔带、Histiodella kristinae谱系带、Histiodella bellburnensis延限带、Dzikodus tablepointensis间隔带、Eoplacognathus suecicus间隔带、Pygodus serra间隔带和Pygodus anserinus(早期型)间隔亚带,桑比期Pygodus anserinus(晚期型)间隔亚带和Belodina compressa间隔带,凯迪期Belodina confluens间隔带、Yaoxianognathus neimengguensis间隔带和Yaoxianognathus yaoxianensis间隔带。由于部分化石带与国际同名带的对比还存在一些矛盾,尚需今后进一步解决。     

新疆巴楚中—晚奥陶世牙形刺生物地层和沉积环境研究

塔里木盆地中央隆起区的中—上奥陶统灰岩相地层露头分布在巴楚良里塔格地区的一间房—唐王城。以牙形刺动物群为依据厘定3个组的时代,从下至上为一间房组(Periodus flabellum层、Pygodus serra层,属达瑞威尔阶)、吐木休克组(Pygodus anserinus层、Baltoniodus alobatus带,属桑比阶)和良里塔格组(含B.confluens动物群层,属凯迪阶下部)。一间房组的开始标志了新一期的海进,此组下段发育1期藻丘,中段发育1期藻丘、1期瓶筐石礁丘和3—4期瓶筐石—棘屑滩,上段代表了海水加深至浪基面之下的过程;吐木休克组沉积过程中达到了海进最大值,为凝缩沉积;良里塔格组代表了逐渐海退的过程,由浅滩和3期藻丘建造组成。     

Reefs in the Early Paleozoic Taebaek Group, Korea: A Review

Various early Paleozoic (Cambrian Series 3–Middle Ordovician) reefs are found in the Taebaek Group, eastern Korea, located in the eastern margin of the Sino-Korean Block. They occur in every carbonate-dominant lithostratigraphic unit of the group, but their morphology and composition differ markedly. The Daegi Formation (middle Cambrian: Cambrian Series 3) contains siliceous sponge-Epiphyton reefs formed in a shallow subtidal environment, which is one of the earliest metazoan-bearing microbial reefs after the archaeocyath extinction. The Hwajeol Formation (upper Cambrian: Furongian) encloses sporadic dendrolites consisting of Angulocellularia, which developed in a relatively deep subtidal environment, representing a rare deeper water example. The onset of the Ordovician radiation resulted in the formation of microbialite–Archaeoscyphia–calathiid patch reefs in shallow subtidal deposits of the Lower Ordovician Dumugol Formation. Subsequent late Early Ordovician relative sea-level fall established extensive peritidal environments, forming microbial mats and stromatolites of the Lower–Middle Ordovician Makgol Formation. Ensuing Ordovician radiation resulted in one of the earliest metazoan skeletal reefs of the Middle Ordovician Duwibong Formation, constructed by stromatoporoid Cystostroma and bryozoan Nicholsonella, and developed around shallow shoals. These reefs reflect ongoing evolution and sea-level change during the early Paleozoic, and exemplify a rare glimpse of peri-Gondwanan records of reef evolution, which warrant detailed investigations and comparison with their counterparts in other regions.     

Structure,Age, and Settings of Formation of Ordovician Complexes of the Northwestern Frame of the Kokchetav Massif,Northern Kazakhstan

This work presents the data on the structure, geochronology, and formation settings of the Ordovician sedimentary and volcanogenic-sedimentary complexes of the Sterlitamak, Mariev, and Imanburluk structural and formational zones located in the western and northwestern frames of the Kokchetav massif (Northern Kazakhstan). In addition, the results of detailed stratigraphic, geochemical, and geochronological studies of the reference section of the Ordovician deposits of the Mariev Zone are given. The studied section is composed of carbonate, terrigenous, and less commonly volcanogenic-sedimentary deposits, confined to a wide stratigraphic interval from Tremadocian Stage of the Lower Ordovician to the lower Sandbian Stage of the Upper Ordovician. For the first time, the study of conodont assemblages made it possible to establish the Early to Middle Ordovician age of the most ancient limestone–dolomite sequence, which was previously conventionally attributed to the Cambrian. The above-lying tuffaceous–terrigenous Kupriyanovka Formation is now attributed to the Middle Ordovician. On the basis of compositional features of the lithoclastic tuffs composing the middle part of the formation, we assume that it was formed within the island arc zone. Limestones from the base of the youngest terrigenous–carbonate Kreshchenovka Formation are attributed to the lower part of the Sandbian Stage of the Upper Ordovician. The study of the geochronology of detrital zircons from terrigenous rocks of the limestone–dolomite sequence has shown that the Early Neoproterozoic quartzite–schist sequences of the Kokchetav massif were the most probable provenance area during its deposition. It was established that there was the change of sedimentation environments from closed lagoons to a relatively deep sea basin with normal salinity and intense circulation of water masses in the northwestern frame of the Kokchetav massif during the Ordovician. During this period of time, there was a sufficiently high level of erosion of provenance areas that resulted in the deposition of thick strata of terrigenous material. A general tendency of the deepening of sedimentation environments from the Early to Late Ordovician was interrupted by sea level rises in the Dapingian and early Darriwilian ages.     

Aragonite depositional facies in a Late Ordovician calcite sea,Eastern Laurentia

The Black River (Upper Ordovician – Sandbian) and Trenton (Upper Ordovician – Katian) groups are traditionally interpreted as a deepening-upward succession deposited in a progressively subsiding Appalachian Basin margin that contained warm-water, marine, photozoan deposits that pass upward into cool-water, marine, heterozoan carbonates. This succession is customarily interpreted to reflect an incursion of cold, high-latitude ocean waters into the area. This view is herein confirmed for coeval carbonates in the northern part of the basin, particularly the St. Lawrence Platform. They are now well explained in this study on the basis of recent studies of cool-water carbonates and calcite–aragonite seas. Overall the succession is one of Sandbian photozoan ramp deposits succeeded by Katian heterozoan ramp carbonates that changed back to photozoan ramp deposits prior to the Hirnantian glaciation. The current interpretation, that deposition took place throughout a calcite sea time, seems at odds with this series of strata. Instead it is herein proposed that deposition took place during an aragonite sea time wherein calcite sea-like sediments accumulated under cold ocean-water temperatures. Such an interpretation is supported by recent experimental data that supports the importance of seawater temperature on CaCO₃ polymorph precipitation. If correct, this means that some of the evidence for calcite sea deposition through time brought about by global tectonics, should be re-evaluated to make sure it was not simply cool-water carbonate production.     

A new species of the Gondwanan genus Cardiolaria Munier-Chalmas in the Sandbian of northwestern Argentina: Paleobiogeographic considerations

A new species of afghanodesmatid, Cardiolaria benicioi, is recorded from Sandbian strata of northwestern Argentina. This species confirms the strong paleobiogeographic relationships between the western Argentina basin and other peri-Gondwanan areas. The Mid-Late Ordovician distribution of bivalves fit well into the Mediterranean Province defined upon brachiopod and trilobite faunas. Similitudes between Tremadocian and Floian bivalves from the western Gondwana and the peri-Gondwanan areas indicate that such ‘Mediterranean’ paleobiogeographic patterns can be traced back well into the Early Ordovician.     

Lower and Middle Ordovician conodonts of Laurentian affinity from blocks of limestone in the Rosroe Formation,South Mayo Trough,western Ireland and their palaeogeographic implication

The Middle Ordovician Rosroe Formation consists of some 1350 m of coarse, mainly siliciclastic to volcaniclastic sedimentary rocks, deposited in a submarine fan environment, and is restricted to the southern limb of the South Mayo Trough, western Ireland. Discrete allochthonous blocks, reaching 5 m in size, are present in the formation at several localities. Conodonts recovered from these blocks, collected from two separate locations, are of late Early and mid Mid Ordovician age. The conodonts have high conodont‐alteration indices (CAI 5) indicative of temperatures as high as 300^o to max. 480 °C; some found in the Lough Nafooey area have abnormally high indices (CAI 6), which correspond to temperatures of about 360^o to max. 550 °C. The oldest fauna is dominated by Periodon aff. aculeatus and characterized by Oepikodus evae typical of the Oepikodus evae Zone (Floian Stage; Stage Slices Fl2–3, Lower Ordovician). The younger conodont assemblage, characterized by Periodon macrodentatus associated with Oistodella pulchra, is referred to the P. macrodentatus conodont Biozone (lower Darriwilian; Stage Slices Dw1–2). The Rosroe conodont assemblages are of Laurentian affinity; comparable faunas are well known from several locations along the east to south‐eastern platform margin of Laurentia and the Notre Dame subzone of central Newfoundland, Canada. The faunal composition from the limestone blocks suggests a shelf edge to slope (or fringing carbonate) setting. The faunal assemblages are coeval with, respectively, the Tourmakeady Formation (Floian–Dapingian) and Srah Formation (Darriwilian) in the Tourmakeady Volcanic Group in the eastern part of the South Mayo Trough and probably are derived from the same or similar laterally equivalent short‐lived carbonate successions that accumulated at offshore ‘peri‐Laurentian’ islands, close to and along the Laurentian margin. During collapse of the carbonate system in the late Mid Ordovician, the blocks were transported down a steep slope and into deep‐water by debris flows, mixing with other rock types now found in the coarse polymict clastics of the Rosroe Formation. The faunas fill the stratigraphical ‘gap’ between the Lower Ordovician Lough Nafooey Volcanic Group and the upper Middle Ordovician Rosroe Formation in the South Mayo Trough and represent a brief interval conducive to carbonate accumulation in an otherwise siliciclastic‐ and volcaniclastic‐dominated sedimentary environment. Copyright © 2015 John Wiley & Sons, Ltd.     

二叠-三叠纪之交牙形石生态新模式

通过对浙江长兴煤山剖面二叠－三叠系界线层沉积相分析及牙形石演化规律研究,提出了该时期Ｃｌａｒｋｉｎａ（Ｎｅｏｇｏｎｄｏｌｅｌａ）,Ｈｉｎｄｅｏｄｕｓ两类重要牙形石属的生态新模式．（１）Ｃｌａｒｋｉｎａ是一种底栖自由游泳生物,可出现于远岸浅海至盆地相较深水环境;（２）Ｈｉｎｄｅｏｄｕｓ是一种洋面浮游型生物,死亡之后可沉于海底的滨、浅海至深水盆地各种环境之中,并可渡过二叠－三叠纪之交海底缺氧事件这一难关,在早三叠世得到发展;因此,Ｈｉｎｄｅｏｄｕｓ可作为全球二叠－三叠系界线划分对比的可靠证据．     

Rugose corals from the Upper Ordovician Sholeshook Limestone of southwest Wales with an assessment of the coral affinities and biofacies

Limestone horizons of Upper Ordovician (Katian) age in southwest Wales contain diverse fossil faunas including rugose corals. The existence of Ordovician Rugosa in Wales was first reported by Murchison in the 1830s, but since then hardly any specimens have been documented systematically until this present study. Newly collected material from the area around Llanddowror (Carmarthenshire) has now confirmed the diversity of rugose corals in the Sholeshook Limestone (Katian age), an arenaceous limestone originating from the shelf edge of the palaeocontinent Avalonia. The majority of the specimens are preserved as moulds. This means that in many instances preservation of the fossils was insufficient for specific identification; nevertheless, it was possible to document a diverse rugose coral fauna, including Helicelasma, probable Grewingkia and Kenophyllum, and a potential early mucophyllid. While associated with considerable difficulties, as some diagnostic features of Rugosa are not visible in moulds, it is demonstrated here that the work with such specimens can result in faunal information which would otherwise be unobtainable. An assemblage of rugose and tabulate corals in the Sholeshook Limestone can be differentiated from a similar assemblage in the neighbouring Robeston Wathen Limestone which has a slightly different lithology. The fauna has strong similarities with other Avalonian (Irish, English, Belgian) as well as Baltic (Estonian and Norwegian) rugose coral faunas. Copyright © 2012 John Wiley & Sons, Ltd.     

滇西保山地区晚泥盆世至早石炭世早期牙形石生物相

一、绪言保山地区是滇西泥盆系和下石炭统出露的主要地区之一,以前已有不少报道。近年来,由于该地区特殊的大地构造位置,引起了国内外广大地质工作者的密切注意,因而对泥盆石炭纪地层及其古生物的研究也更加深入。保山地区的上泥盆统目前暂分为何元寨组和整合其上的大寨门组,前者底部尚属中泥盆统(龚大明等,待刊)。下石炭统下部称鱼洞组。何元寨组在其命名剖面施甸何元寨主要是一套中层灰岩和泥灰岩,厚约220m,腕足、珊瑚、层孔及苔藓动物丰富,向东北至大寨门、由     

Similar morphological and chemical variations of Gloeocapsomorpha prisca in Ordovician sediments and cultured Botryococcus braunii as a response to changes in salinity

Most Ordovician source rocks consist of accumulation of a colonial marine microorganism, Gloeocapsomorpha prisca (G. prisca) whose nature, ecology and affinity with extant organisms have been in dispute for years. Furthermore, recent studies have shown major differences in phenol moieties between two G. prisca-rich samples. Examination of five G. prisca-rich kerogens by electron microscopy and pyrolysis studies revealed (i) the occurrence of two markedly distinct “morpho/chemical” types: a “closed/phenol-rich” type (Baltic samples) and an “open/phenol-poor” one (North American samples) and (ii) the selective preservation of the resistant micromolecular material building up the thick cell walls in the original organism. Comparison with extant Botryococcus braunii (a widespread green microalga) grown on media of increasing salinity suggests that G. prisca is likely to be a planktonic green microalga related to B. braunii, which can adapt to large salinity variations which, in turn, control its polymorphism. The large differences in colony morphology and in the content of phenol moieties observed in fossil G. prisca and the resulting occurrence of two “morpho/chemical” types, should therefore reflect depositional environments with different salinities. The presence of thick, highly aliphatic, resistant walls in G. prisca selectively preserved during fossilization, accounts for the major contribution of this organism to Ordovician organic-rich sediments and for the resulting typical signature of Ordovician oils.     

Carbon isotope composition of Upper Cambrian to Lower Ordovician carbonate in Korea,and its bearing on the Cambrian–Ordovician boundary and Lower Ordovician paleoceanography

This study presents an example of locating Cambrian–Ordovician boundary in the lower Paleozoic carbonate succession in Korea using carbon isotope stratigraphy. The Yeongweol Unit of the lower Paleozoic Joseon Supergroup comprises the Upper Cambrian Wagok Formation and the Lower Ordovician Mungok Formation in the Cambrian–Ordovician transition interval. Conventionally, the boundary was placed at the lithostratigraphic boundary between the two formations. This study reveals that the boundary is positioned in the basal part of the Mungok Formation based on the carbon isotope stratigraphy coupled with biostratigraphic information of conodont and trilobite faunas. The δ¹³C curve of the Lower Ordovician Mungok Formation shows a similar trend to that of the coeval stratigraphic interval of Argentine Precordillera (Buggisch et al., 2003), suggesting that the δ¹³C curve of the Mungok Formation reflects the Early Ordovician global carbon cycle.     

Early Ordovician CA-IDTIMS U–Pb zircon dating and conodont biostratigraphy,Canning Basin,Western Australia

A continuously cored section of more than 300 m through the Nambeet Formation and the basal part of the conformably overlying Willara Formation in the Olympic 1 petroleum well, drilled in the Canning Basin of northern Western Australia, yields valuable information that increases by more than 40% the number of precise isotopic ages available to constrain the Ordovician Period. New CA-IDTIMS U–Pb zircon ages for seven bentonite layers in the Olympic 1 core are integrated into a new conodont biostratigraphic framework for the Early Ordovician comprising four biozones recognised in this well. The weighted mean U–Pb dates range from 479.37 ± 0.16 Ma within the late Tremadocian Paroistodus proteus conodont Biozone, to 470.18 ± 0.13 Ma near the boundary between the Floian and Dapingian stages within the Jumudontus gananda conodont Biozone. The intervening Prioniodus oepiki–Serratognathus bilobatus conodont Biozone (early Floian) and succeeding Oepikodus communis conodont Biozone (middle Floian) are similarly well constrained by isotopic dates centred on ca 477 Ma for the early Floian and by three ages of 473–471 Ma for the middle Floian. The seven new isotopic dates significantly increase the precision of dating for the Early Ordovician, where previously only two ages with limited or imprecise biostratigraphic control were known globally.