Tsunamis Observed on and Near the Turkish Coast

For centuries, inhabitants of coastal areas have suffered from the effects of tsunamis. Turkey, with a coastline of 8333 km, has experienced many tsunamis.Historical records reveal that, during the observation period over 3000 years, the coastal and surrounding areas of Turkey have been affected by more than ninety tsunamis. These tended to cluster around the Marmara Sea, the city of Istanbul and the gulfs of Izmit, Izmir, Fethiye and Iskenderun. Each of the tsunami occurrences surveyed in this paper deserves further individual study. The most extensive available information concerns the tsunamis associated with the Istanbul Earthquakes of 1509 and 1894, the Eastern Marmara Earthquake in 1963 and that of Izmit in 1999,which disturbed the Marmara Sea; the Earthquake of 1939 in Erzincan ineastern Anatolia; and the 1968 Bartn Earthquake, which affected Fatsa and Amasra on the Black Sea. In addition to these, it is known that a tsunami occurred in 1598 on the shores of the Black Sea in connection with an earthquake at Amasya in northern Anatolia.     

Carbonate characteristics of waters of the Arctic Ocean continental slope

Carbonate characteristics of the water mass of the deepwater part of the Arctic Ocean (AO) in the continental slope area were determined, and the range and reasons of their variability during summer-fall season were revealed. The AO water area is a meaningful sink for atmospheric carbon dioxide. The warm intermediate Atlantic waters (AW) are also undersaturated with carbon dioxide relative to its content in the atmosphere. While these waters move along AO continental slope, the value pCO₂ in the AW core decreases to 8–10 μatm (mainly, due to drop in the water temperature). The potential absorption capacity of the AO deepwater basin is estimated at approximately 48 Tg of carbon (without sea ice taken into account). Joint analysis of carbonate and hydrological parameters showed that near-bottom waters formed on the shallow shelf of the Laptev Sea, which is rich in inorganic and organic carbon of terrestrial and marine genesis, take part in formation of halocline waters of the AO. They are modified due to interaction with AW penetrating to the shelf and are transferred to the deepwater AO segment, where they occur in the halocline according to their density. Transformed near-bottom waters of the Laptev Sea shelf, similar to waters of the halocline of Pacific origin in the eastern sector of the AO, are traced above the continental slope in Amundsen Basin on the basis of higher CO₂ concentrations.     

Land-based sources of pollution along the Izmit Bay and their effect on the coastal waters

The degree and the contribution of each point source to the pollution were determined in the Izmit Bay during the period 1999–2000. During 8 campaigns, samples from 11 points in the channels and water samples from 5 points in the coastal sea were collected for chemical analysis. The important pollutant parameters taken into account are inflow of total organic carbon (TOC), total suspended solids (TSS), total phosphorus (TP), total nitrogen (TN), nitrate, ortho-phosphate, ammonia and total polycyclic aromatic hydrocarbons (t-PAHs) in the discharge channels, and TOC, TSS, nitrate, ortho-phosphate, chlorophyll-a, temperature, dissolved oxygen (DO), and salinity in the coastal stations of the Bay. It should be pointed out that the industrial wastewaters entering the bay are partially treated but domestic wastes are discharged directly into the surface waters without any treatment. Of the pollution parameters measured in the channels, the highest concentrations, except TP, were observed in the Dil River and in the Eastern Channel. Concentrations of TOC, TSS, TN, TP, ammonia, nitrate and o-phosphate were found at concentrations of 231, 290, 152, 3.8, 16, 79, and 3.07 mg/L, respectively. Annual inflows of TOC were 21,301, 580, and 775 t/year and for TSS were 26,742, 585, and 1505 t/year in the western, central and eastern parts, respectively. The results show that the water quality of the bay has been deteriorated and 80% of the pollution was caused by Dil River for all parameters measured.     

Characteristics of the 1912 co-seismic rupture along the North Anatolian Fault Zone (Turkey): implications for the expected Marmara earthquake

Evidence of right‐lateral offsets associated with the 1912 earthquake (M_w 7.4) along the North Anatolian Fault (Gaziköy–Saros segment) allow us to survey (using DGPS) the co‐seismic and cumulative slip distribution. The damage distribution and surface breaks related with the earthquake show an elongated zone of maximum intensity (X MSK) parallel to the fault rupture on land but this may extend offshore to the north‐east and south‐west. Detailed mapping of the fault using topographic maps and aerial photographs indicates the existence of pull‐apart basins and pressure ridges. At several localities, the average 1912 offset along strike is 3.5–4 m and cumulative slip is 2–6 times that of individual movement. The fault rupture geometry and slip distribution suggest the existence of three subsegments with a combined total length of 110–120 km, a fault length and maximum slip similar to those of the 1999 Izmit earthquake. The amount of slip at the north‐easternmost section and in the coastal region of the Sea of Marmara reaches an average 4 m, thereby implying the offshore extension of the 1912 rupture. The results suggest that the 1912 event generated up to 150 km of surface faulting, which would imply a M_w 7.2–7.4 earthquake and which, added with rupture lengths of the 1999 earthquakes, help to constrain the remaining seismic gap in the Sea of Marmara.     

Evaluation of earthquake potential along the Northern Anatolian Fault Zone in the Marmara Sea using comparisons of GPS strain and seismotectonic parameters

Seismotectonic parameters including the Gutenberg-Richter b-value and multifractal dimensions D₂ and D₁₅ of seismicity patterns (both spatial and temporal) were compared to GPS-derived maximum shear and dilatation strains measured in the Marmara Sea region of western Turkey along the Northern Anatolian Fault Zone (NAFZ). Comparisons of seismotectonic parameters and GPS-derived maximum shear and dilatation strain along the NAFZ in the vicinity of the 1999 M7.4 Izmit earthquake reveal a positive correlation (r = 0.5, p = 0.05) between average dilatation and the Gutenberg-Richter b-value. Significant negative correlation (r = − 0.56, p = 0.03 and r = − 0.56, p = 0.02) was also observed between the spatial fractal dimension D₂ and GPS-derived maximum geodetic and shear strain. This relationship suggests that, as maximum geodetic and shear strains increase, seismicity becomes increasingly clustered.Anomalous interrelationships are observed in the Marmara Sea region prior to the Izmit event along a bend in the NAFZ near the eastern end of the Marmara Sea known as the Northern Boundary Fault (NBF). An asperity is located near the northwest end of the NBF. Along the 50-km length of the NBF, GPS strains become slightly compressive. The correlation between b-value and GPS-derived dilatation suggests that regions in compression have increased probability of larger magnitude rupture. The NBF appears to serve as an impediment to the transfer of strain from east to west along the NAFZ. Recurrence times for large earthquakes along the NBF are larger than in surrounding areas. Temporal clustering of seismicity in the vicinity of the NBF may represent foreshocks of an impending rupture.     

Earthquake risk to industry in Istanbul and its management

Istanbul is home to 40% of the industrial facilities in Turkey. Thirty percent of the population working in industry lives in the city. Past earthquakes have evidenced that the structural reliability of residential and industrial buildings in the country is questionable. In the Marmara region the earthquake hazard is very high with a 2% annual probability of occurrence of a magnitude 7+ earthquake on the main Marmara fault. These facts make the management of industrial risks imperative for the reduction of socioeconomic losses. In this paper we present a first-order assessment of earthquake damage to the industry in Istanbul and raise issues for better characterization and quantification of industrial losses and management of urban industrial risks. This paper borrows from the project report entitled ‘Earthquake Risk Assessment for Industrial Facilities in Istanbul’. The full report can be found at http://www.koeri.boun.edu.tr/depremmuh.html under the link ‘Research and Applied Projects’.     

Current accelerating seismic excitation along the northern boundary of the Aegean microplate

According to previous observations [Geophys. Res. Lett. 27 (2000) 3957], the generation of large (M≥7.0) earthquakes in the western part of the north Anatolian fault system (Marmara Sea) is followed by strong earthquakes along the Northern Boundary of the Aegean microplate (NAB: northwestermost Anatolia–northern Aegean–central Greece–Ionian islands). Therefore, it can be hypothesized that a seismic excitation along this boundary should be expected after the occurrence of the Izmit 1999 earthquake (M=7.6). We have applied the method of accelerating seismic crustal deformation, which is based on concepts of critical point dynamics in an attempt to locate more precisely those regions along the NAB where seismic excitation is more likely to occur. For this reason, a detailed parametric grid search of the broader NAB area was performed for the identification of accelerating energy release behavior.Three such elliptical critical regions have been identified with centers along this boundary. The first region, (A), is centered in the eastern part of this boundary (40.2°N, 27.2°E: southwest of Marmara), the second region, (B), has a center in the middle part of the boundary (38.8°N, 23.4°E: East Central Greece) and the third region, (C), in the westernmost part of the boundary (38.2°N, 20.9°E: Ionian Islands). The study of the time variation of the cumulative Benioff strain in two of the three identified regions (A and B) revealed that intense accelerating seismicity is observed especially after the occurrence of the 1999 Izmit mainshock. Therefore, it can be suggested that the seismic excitation, at least in these two regions, has been triggered by the Izmit mainshock.Estimations of the magnitudes and origin times of the expected mainshocks in these three critical regions have also been performed, assuming that the accelerating seismicity in these regions will lead to a critical point, that is, to the generation of mainshocks.     

A study of microearthquake seismicity and focal mechanisms within the Sea of Marmara (NW Turkey) using ocean bottom seismometers (OBSs)

We have carried out seismological observations within the Sea of Marmara (NW Turkey) in order to investigate the seismicity induced after Gölcük–İzmit (Kocaeli) earthquake (M_w 7.4) of August 17, 1999, using ocean bottom seismometers (OBSs). High-resolution hypocenters and focal mechanisms of microearthquakes have been investigated during this Marmara Sea OBS project involving deployment of 10 OBSs within the Çınarcık (eastern Marmara Sea) and Central-Tekirdağ (western Marmara Sea) basins during April–July 2000. Little was known about microearthquake activity and their source mechanisms in the Marmara Sea. We have detected numerous microearthquakes within the main basins of the Sea of Marmara along the imaged strands of the North Anatolian Fault (NAF). We obtained more than 350 well-constrained hypocenters and nine composite focal mechanisms during 70 days of observation. Microseismicity mainly occurred along the Main Marmara Fault (MMF) in the Marmara Sea. There are a few events along the Southern Shelf. Seismic activity along the Main Marmara Fault is quite high, and focal depth distribution was shallower than 20 km along the western part of this fault, and shallower than 15 km along its eastern part. From high-resolution relative relocation studies of some of the microearthquake clusters, we suggest that the western Main Marmara Fault is subvertical and the eastern Main Marmara Fault dips to south at 45°. Composite focal mechanisms show a strike-slip regime on the western Main Marmara Fault and complex faulting (strike-slip and normal faulting) on the eastern Main Marmara Fault.     

Structural relationship between the Sea of Marmara Basin and the North Anatolian Fault Zone

The North Anatolian Fault (NAF) zone is 1500 km long, extending almost up to the Greek mainland in the west. It is a seismically active right-lateral strike-slip fault that accommodates the relative motion between the Turkish block and Black Sea plate. The Sea of Marmara lies along the western part of the NAF and shows evidence of subsidence. In this area pure strike-slip motion of the fault zone changes into extensional strike-slip movement that is responsible for the creation of the Sea of Marmara and the North Aegean basins. The northern half of the Sea of Marmara is interpreted as a large pull-apart basin. This basin is subdivided into three smaller basins separated by strike-slip fault segments of uplifted blocks NE-SW. Basinal areas are covered by horizontally layered sedimentary sequences. Uplifted blocks have undergone compressional stress. All the blocks are subsiding and are undergoing vertical motions and rotations relative to one another. The uplifted blocks exhibit positive Bouguer gravity anomalies. According to gravity interpretation, there is relative crustal thinning under the Sea of Marmara. The northern side of the Sea of Marmara is marked by a distinctive deep-rooted magnetic anomaly, which is dissected and shifted southward by strike-slip faulting. The southern shelf areas of the Sea of Marmara are dominated by short-wavelength magnetic anomalies of shallow origin.     

Stress modeling to determine the through-going active fault geometry of the Western North Anatolian Fault,Turkey

The North Anatolian Fault (NAF) is a 1200 km long dextral strike-slip fault which is part of an east-west trending dextral shear zone (NAF system) between the Anatolian and Eurasian plates. The North Anatolian shear zone widens to the west, complicating potential earthquake rupture paths and highlighting the importance of understanding the geometry of active fault systems. In the central portion of the NAF system, just west of the town of Bolu, the NAF bifurcates into the northern and southern strands, which converge, then diverge to border the Marmara Sea. At their convergence east of the Marmara Sea, these two faults are linked through the Mudurnu Valley. The westward continuation of these two fault traces is marked by further complexities in potential active fault geometry, particularly in the Marmara Sea for the northern strand, and towards the Biga Peninsula for the southern strand. Potential active fault geometries for both strands of the NAF are evaluated by comparing stress models of various fault geometries in these regions to a record of focal mechanisms and inferred paleostress from a lineament analysis. For the Marmara region, the best-fit active fault geometry consists of the northern and southern bounding faults of the Marmara basin, as the model representing this geometry better replicated primary stress orientations seen in focal mechanism data and stress field interpretations. In the Biga Peninsula region, the active geometry of the southern strand has the southern fault merging with the northern fault through a linking fault in a narrow topographic valley. This geometry was selected over the other two as it best replicated the maximum horizontal stresses determined from focal mechanism data and a lineament analysis.     

Seismic behaviour of the North Anatolian Fault beneath the Sea of Marmara (NW Turkey): implications for earthquake recurrence times and future seismic hazard

Possible long-term seismic behaviour of the Northern strand of the North Anatolian Fault Zone, between western extreme of the 1999 İzmit rupture and the Aegean Sea, after 400 AD is studied by examining the historical seismicity, the submarine fault mapping and the paleoseismological studies of the recent scientific efforts. The long-term seismic behaviour is discussed through two possible seismicity models devised from M _S ≥ 7.0 historical earthquakes. The estimated return period of years of the fault segments for M1 and M2 seismic models along with their standard deviations are as follows: F4 segment 255 ± 60 and 258 ± 12; F5 segment 258 ± 60 and 258 ± 53; F6 segment 258 ± 60 and 258 ± 53; F7 segment 286 ± 103 and 286 ± 90; F8 segment 286 ± 90 and 286 ± 36. As the latest ruptures on the submarine segments have been reported to be during the 1754–1766 earthquake sequence, and the 1912 mainshock rupture has been evidenced to extend almost all over the western part of the Sea of Marmara, our results imply imminent seismic hazard and, considering the mean recurrence time, a large earthquake to strike the eastern part of the Sea of Marmara in the next two decades.     

Asymmetric slip partitioning in the Sea of Marmara pull‐apart: a clue to propagation processes of the North Anatolian Fault?

Between 1939 and 1999 the North Anatolian fault (NAF) experienced a westward progression of eight large earthquakes over 800 km of its morphological trace. The 2000-km-long North Anatolian transform fault has also grown by westward propagation through continental lithosphere over a much longer timescale (∼10 Myr). The Sea of Marmara is a large pull-apart that appears to have been a geometrical/mechanical obstacle encountered by the NAF during its propagation. The present paper focuses on new high-resolution data on the submarine fault system that forms a smaller pull-apart beneath the Northern Sea of Marmara, between two well-known strike-slip faults on land (Izmit and Ganos faults). The outstandingly clear submarine morphology reveals a segmented fault system including pull-apart features at a range of scales, which indicate a dominant transtensional tectonic regime. There is no evidence for a single, continuous, purely strike-slip fault. This result is critical to understanding of the seismic behaviour of this region of the NAF, close to Istanbul. Additionally, morphological and geological evidence is found for a stable kinematics consistent both with the long-term displacement field determined for the past 5 Myr and with present-day Anatolia/Eurasia motion determined with GPS. However, within the Sea of Marmara region the fault kinematics involves asymmetric slip partitioning that appears to have extended throughout the evolution of the pull-apart. The loading associated with the westward propagation process of the NAF may have provided a favourable initial geometry for such a slip separation.     

The sources of metal contents in the shelf sediments from the Marmara Sea, Turkey

The shelf area is the largest morphological unit of the Marmara Sea and is subjected to increasing population, urbanization, and industrial activities. Metal contents (Al, Fe, Mn, Cu, Pb, Zn, Ni, Cr, Co and Hg) of the surface sediments from the shelf areas of the Marmara Sea generally do not indicate shelf-wide pollution. The variability of the metal contents of the shelf sediments is mainly governed by the geochemical differences in the northern and southern hinterlands. Northern shelf sediments contain lower values compared to those of the southern shelf, where higher Ni, Cr, Pb, Cu and Zn are derived from the rock formations and mineralized zones. However, besides from the natural high background in the southern shelf, some anthropogenic influences are evident from EF values of Pb, Zn and Cu, and also from their high mobility in the semi-isolated bay sediments. Anthropogenic influences are found to be limited at the confluence of Istanbul Strait in the northern shelf. However, suspended sediments along the shallow parts of the northern shelf were found to be enriched in Pb and Hg and to a lesser degree in Zn, reflecting anthropogenic inputs from Istanbul Metropolitan and possibly from the Black Sea via the Istanbul Strait.     

Rupture progression along discontinuous oblique fault sets: implications for the Karadere rupture segment of the 1999 Izmit earthquake, and future rupture in the Sea of Marmara

Large earthquakes in strike-slip regimes commonly rupture fault segments that are oblique to each other in both strike and dip. This was the case during the 1999 Izmit earthquake, which mainly ruptured E–W-striking right-lateral faults but also ruptured the N60°E-striking Karadere fault at the eastern end of the main rupture. It will also likely be so for any future large fault rupture in the adjacent Sea of Marmara. Our aim here is to characterize the effects of regional stress direction, stress triggering due to rupture, and mechanical slip interaction on the composite rupture process. We examine the failure tendency and slip mechanism on secondary faults that are oblique in strike and dip to a vertical strike-slip fault or “master” fault. For a regional stress field well-oriented for slip on a vertical right-lateral strike-slip fault, we determine that oblique normal faulting is most favored on dipping faults with two different strikes, both of which are oriented clockwise from the strike-slip fault. The orientation closer in strike to the master fault is predicted to slip with right-lateral oblique normal slip, the other one with left-lateral oblique normal slip. The most favored secondary fault orientations depend on the effective coefficient of friction on the faults and the ratio of the vertical stress to the maximum horizontal stress. If the regional stress instead causes left-lateral slip on the vertical master fault, the most favored secondary faults would be oriented counterclockwise from the master fault. For secondary faults striking ±30° oblique to the master fault, right-lateral slip on the master fault brings both these secondary fault orientations closer to the Coulomb condition for shear failure with oblique right-lateral slip. For a secondary fault striking 30° counterclockwise, the predicted stress change and the component of reverse slip both increase for shallower-angle dips of the secondary fault. For a secondary fault striking 30° clockwise, the predicted stress change decreases but the predicted component of normal slip increases for shallower-angle dips of the secondary fault. When both the vertical master fault and the dipping secondary fault are allowed to slip, mechanical interaction produces sharp gradients or discontinuities in slip across their intersection lines. This can effectively constrain rupture to limited portions of larger faults, depending on the locations of fault intersections. Across the fault intersection line, predicted rakes can vary by >40° and the sense of lateral slip can reverse. Application of these results provides a potential explanation for why only a limited portion of the Karadere fault ruptured during the Izmit earthquake. Our results also suggest that the geometries of fault intersection within the Sea of Marmara favor composite rupture of multiple oblique fault segments.     

Heterogeneous response of hydrogeological systems to the Izmit and Düzce (Turkey) earthquakes of 1999

At four sites in Turkey and Armenia the physico-chemical properties of thermal and mineral waters were monitored continuously during the Izmit and Düzce earthquakes that occurred along the North Anatolian fault in August and November 1999. The epicentral distances between the moment magnitude (M_w) 7.6 Izmit earthquake and the monitoring locations were 313, 488, 1,161, and 1,395 km. At the most distant site, the specific electrical conductivity of mineral water from a flowing artesian well dropped co-seismically and postseismically by 7%. No changes were observed at the other sites, although the estimated earthquake strains and peak ground accelerations are much higher. A similar pattern was observed after the Düzce earthquake, which happened three months after the Izmit event. The response of a hydrogeological system seems to depend on the site characteristics rather than on the nature of the earthquake. A hydrogeological model for the sensitive observation site farthest from the Izmit earthquake explains the observations in terms of a changed mixing ratio between two fluid components. Passing seismic waves may trigger a local pore-pressure increase according to the mechanism of advective overpressure. The preconditions for this mechanism, free gas bubbles in the aquifer in combination with a trap for rising bubbles, is probably not fulfilled by the other groundwater systems. Electronic Publication     

基于离散元方法对走滑拉分盆地演化及次级断裂扩展过程研究

拉分盆地是一种与走滑断裂带密切相关的特殊拉张构造,因其重要的构造意义,及其与火山活动、中小地震群集、特殊的成矿作用间的伴生关系而受到研究者的高度重视。关于拉分盆地的形成演化过程,已有较多的研究成果,但是由于研究手段的限制,缺少对盆地演化中次级断裂扩展过程的研究。基于离散元的数值计算方法是研究断裂扩展方式的理想方法。本文采用基于离散元的颗粒流方法,揭示纯走滑拉分盆地发育过程中的断裂扩展和连接过程,为拉分盆地演化机理和断裂扩展提供新的研究方法。同时,根据主走滑断层与块体运动方向的夹角不同,建立不同的张扭性拉分盆地模型,系统研究张扭性盆地的断裂扩展和演化机理。将上述理论研究结果与死海盆地等经典拉分盆地实例相结合,探讨了死海盆地、土耳其Cinarcik盆地、哥伦比亚El Paraiso盆地等的形成演化机理和断裂扩展方式。      

Circulation of water masses in the Baltic Proper revealed through iodine isotopes

Tracer technology has been used to understand water circulation in marine systems where the tracer dose is commonly injected into the marine waters through controlled experiments, accidental releases or waste discharges. Anthropogenic discharges of ¹²⁹I have been used to trace water circulation in the Arctic and North Atlantic Ocean. Here, ¹²⁹I, together with ¹²⁷I, is utilized as a tracer of water pathways and circulation in the Baltic Sea through collection of seawater depth profiles. The results indicate the presence of ¹²⁹I signatures which are distinct for each water mass and provide evidence for: (1) inflow water masses through the Drogden Sill that may reach as far as the SW of the Arkona Sea, (2) a portion of North Atlantic water in the bottom of Arkona basin, (3) cyclonic upwelling which breaks through the halocline in a pattern similar to the Baltic haline conveyor belt and (4) more influx of fresher water from the Gulf of Finland and Bothnian Sea in August relative to April. These findings provide advances in labeling and understanding water pathways in the Baltic Sea.     

Geochemical evidence for a large methane release during the last deglaciation from Marmara Sea sediments

Recent methane inventories have revealed the potential impact of gas hydrates on the global carbon cycle, and hence in climate change (Milkov, 2004). However, only a few studies have traced methane release in the geologic record. Here, we show geochemical evidence for a large scale methane release at mid-latitudes during the last deglaciation. The Sea of Marmara, an enclosed sea between the Mediterranean and Black Seas, is located in a tectonically active basin with gas hydrate expulsion and the formation of shallow gas hydrates. Since depths in the basin are shallower than 1100 m, future global temperatures are expected to have a great influence in destabilizing methane clathrates. Among the suite of biomarkers, we have focused on diplopterol and diploptene profiles in core MD012430, retrieved from the central basin in the Marmara Sea. Our results indicate that during the last 15,000 years, hopanoids showed important concentration variations with a pronounced peak during the deglaciation.The lack of a relationship between diplopterol/diploptene and phytoplanktonic biomarker concentrations, as well as a depleted isotopic composition, have linked the hopanoid maxima to methanotrophic activity, suggesting that an intense methane release occurred at the onset of deglaciation in the Marmara Sea. The vulnerability of the hydrate stability zone to changes in temperature and pressure under this range of shallow water depths, as well as the relative timing of the hopanoid maxima and sea surface temperature rise, points to thermal destabilization of hydrates as a trigger for methane release in the water column.     

Fish assemblage structure in relation to environmental variation in a Texas Gulf coastal wetland

We described seasonal fish-assemblages in an estuarine marsh fringing Matagorda Bay, Gulf of Mexico. Habitat zones were identified by patterns of fish species abundance and indicator species optima along gradients in salinity, dissolved oxygen (DO), and depth in our samples. Indicators of the lower brackish zone (lower lake and tidal bayou closest to the bay) were gulf menhaden (Brevoortia patronus), bay anchovy (Anchoa mitchilli), silver perch (Bairdiella chrysoura), and spotted seatrout (Cynoscion nebulosus) at salinity >15‰, DO 7–10 mg l⁻¹, and depth <0.5 m. Indicators of the upper brackish zone (lake and fringing salt marsh) were pinfish (Lagodon rhomboides) and spot (Leiostomus xanthurus) at salinity 10–20‰, DO >10 mg l⁻¹, and depth <0.5 m. In the freshwater wetland zone (diked wetland, ephemeral pool, and perennial scour pool), indicators were sheepshed minnow (Cyprinod on variegatus), rainwater killifish (Lucania parva), mosquitofish (Gambusia affinis), and sailfin molly (Poecilia latipinna) at salinity <5‰, DO <5 mg l⁻¹, and depth ≥1 m. In the freshwater channelized zone (slough and irrigation canal), indicators were three sunfish species (Lepomis), white crappie (Pomoxis annularis), and gizzard shad (Dorosoma cepedianum) at salinity <5‰, DO <5 mg l⁻¹, and depth >1.5 m. In brackish zones, seasonal variation in species diversity among sites was positively correlated with temperature, but assemblage structure also was influenced by depth and DO. In the freshwater zones, seasonal variation in species diversity among sites was positively correlated with depth, DO, and salinity, but assemblage structure was weakly associated with temperature. Species diversity and assemblage structure were strongly affected by the connectivity between freshwater wetland and brackish zones. Uncommon species in diked wetlands, such as tarpon (Megalops atlanticus) and fat sleeper (Dormitator maculatus), indicated movement of fishes from the brackish zone as the water level rose during natural flooding and scheduled (July) releases from the diked wetland. From September to July, diversity in the freshwater wetland zone decreased as receding waters left small isolated pools, and fish movement became blocked by a water-control structure. Subsequently, diversity was reduced to a few species with opportunistic life histories and tolerance to anoxic conditions that developed as flooded vegetation decayed.     

Middle Pleistocene bivalves of the İznik lake basin (Eastern Marmara, NW Turkey) and a new paleobiogeographical approach

?znik Lake is a tectonically originated basin mainly controlled by the E–W trending middle strand of the North Anatolian Fault (NAF) system. Pleistocene sediments occurring in front of the faults are well exposed both in the northern and in the southern shorelines of the basin. In this study, two endemic brackish water bivalve species, Didacna subpyramidata Pravoslavkev 1939 and Didacna nov. sp. were found in the oldest terrace of the northern Pleistocene sequence. Having characterized morphology, these species serve as stratigraphic indicators in the regional Pleistocene stratigraphy of the Ponto-Caspian region, and thus are well correlated to the assemblages of the early Khazarian subhorizon (Middle Pleistocene). Hence, these data demonstrate that the early Khazarian brackish water sea covered the study area. Additionally, a model for the formation of the basin is proposed: the ?znik lake basin was a gulf of the former Marmara Sea in the early Khazarian, connecting the Marmara to the Black Sea and the Caspian Sea. The subsequent regional prograding uplifts, main dextral strike-slip fault and many normal faults of the NAF Zone cut off the marine connections to the basin, leading to its present location and topographic level.