Aminostratigraphy of European marine interglacial deposits

Molluscan fossils collected from shallow water marine sediment across NW Europe and nearby Arctic regions have been analysed for the extent of isoleucine epimerization ( ratio) in indigenous protein residues. The ratios confirm that essentially all ‘classical’ Eemian sites from NW Europe are of the same age, and are correlative with the type locality near Amersfoort in the Netherlands; shells from interglacial marine sediment beneath the type Weichselian till in Poland also correlate with the type Eemian site. ratios in Holsteinian marine shells (0.29) are substantially higher than in their Eemian counterparts (0.17); ‘Late Cromerian’ shells yield even higher ratios (0.46). ratios in late glacial shells (0.06) and Middle Weichselian shells (0.09) permit differentiation from modern (0.01) and last interglacial material. Based on the position of the Matuyama-Brunhes boundary and the differences in ratios, the Eemian must correlate with isotope substage 5e, whereas the Holsteinian is most likely substage 7c, possibly stage 9 but certainly younger than stage 11. Intra-Saalian warm periods may be terrestrial equivalents of the younger substages of stage 7. Extensive pre-Eemian marine sediments along the SW coast of Denmark previously correlated with the Holsteinian are shown to be of ‘Late Cromerian’ age. The underlying till there is the first widespread evidence of a pre-Elsterian till in NW Europe. ratios in molluscs from last interglacial sites along the Arctic coast of the USSR, the Arctic Islands and eastern Greenland are substantially lower than in their European counterparts due to their low thermal histories. The combined mid- and high-latitude data are used to develop a predictive model for the expected ratio in any of several moderate epimerization-rate taxa for last interglacial sites with mean temperatures between −20 and +15°C.Not all sites could be unambiguously assigned to an established interglacial. The Fjøsanger (Norway) and Margareteberg (Sweden) sites previously thought to be Eemian, yield ratios higher than in secure nearby Eemian material. It is yet unresolved whether these are aberrant sites or if they predate the last interglacial. In situ shoreline deposits encountered in borings in SW Belgium and in exposures on the Belgium coastal plain contain molluscs that yield ratios intermediate between secure Eemian and Late Weichselian ratios, raising the possibility that a late stage 5 high-sea-level event attained near-modern levels in the southern North Sea basin. Resolution of these uncertainties is the focus of future work.     

Land biotas of the last interglacial/glacial cycle on Jameson Land, East Greenland

Macrofossil plant and insect remains from nearshore marine sediments in Jameson Land, central East Greenland show that the land biotas of the last interglacial stage, the Langelandselv stage, were more diverse than those of the Holocene. Rich dwarf shrub heaths with a diverse assemblage of ericaceous plants occupied low land areas with copses of Betula pubescens on sheltered sites. Many southern extra-limital species were present, and the mean summer temperature was c . 5°C higher than today. The subarctic bioclimatic zone was displaced from southernmost Greenland/Iceland to central East Greenland. The diverse beetle fauna was of palaearctic affinity and strikingly different from the Plio-Pleistocene and the Holocene Greenlandic beetle faunas. A few fossil assemblages from the Hugin Sø Interstade, which is correlated with oxygen isotope stage 5c (early last glacial stage), point to poor, perhaps entirely herbaceous vegetation with a mean summer temperature that was perhaps 3 4°C lower than today.     

Elsterian‐Holsteinian deposits at Kås Hoved,northern Denmark: sediments,foraminifera, ostracods,and stable isotopes

The coastal cliff section at Kås Hoved in northern Denmark represents one of the largest exposures of marine interglacial deposits in Europe. High‐resolution analyses of sediments, foraminifera, ostracods, and stable isotopes (oxygen and carbon) in glacial‐interglacial marine sediments from this section, as well as from two adjacent boreholes, are the basis for an interpretation of marine environmental and climatic change through the Late Elsterian‐Holsteinian glacial‐interglacial cycle. The overlying glacial deposits show two ice advances during the Saalian and Weichselian glaciations. The assemblages in the initial glacier‐proximal part of the marine Late Elsterian succession reveal fluctuations in the inflow of sediment‐loaded meltwater to the area. This is followed by faunal indication of glacier‐distal, open marine conditions, coinciding with a gradual climatic change from arctic to subarctic environments. Continuous marine sedimentation during the glacial‐interglacial transition is presumably a result of a large‐scale isostatic subsidence caused by the preceding extended Elsterian glaciation. The similarity of the climatic signature of the interglacial Holsteinian and Holocene assemblages in this region indicates that the Atlantic Ocean circulation was similar during these two interglacials, whereas Eemian interglacial assemblages indicate a comparatively high water temperature associated with an enhanced North Atlantic Current. The foraminiferal zones are correlated with other Elsterian‐Holsteinian sites in Denmark, as well as those in the type area for the Holsteinian interglacial in northern Germany and the southern North Sea. Correlation of the NW European Holsteinian succession with the marine isotope stages MIS 7, 9 or 11 is still unresolved.     

A multiproxy macrofossil record of Eemian palaeoenvironments from Klaksvík,the Faroe Islands

Studies of interglacial successions are critical to our understanding of the environmental history of an area. Analyses of macrofossil remains of plants and invertebrates from Eemian sediments exposed in a coastal cliff section at Borðoyarvík near Klaksvík, Bordoy, northeastern Faroe Islands, indicate that the sediments accumulated in a coastal lagoon. The fossil flora comprises tree birch Betula sect. Albae and we suggest that birch forests were found locally at sheltered sites in the area. Tree birch also occurred on the islands during the mid‐Holocene. The only other woody plant recovered from the Eemian deposit is the dwarf‐shrub Empetrum nigrum, which is common on the islands today. Remains of herbaceous plants are rare but include Viola, Ajuga, Myosotis, Urtica dioica and Ranunculus. The bryophyte flora is species‐rich and most of the fragments belong either to stream species or to species of humid or wet habitats. The fossil flora and fauna also comprise a number of freshwater species that probably lived in an oligotrophic lake and in streams in the catchment of the lagoon. The climate during deposition of the lagoonal sediments was similar to the Holocene oceanic climate of the Faroe Islands. The study adds to our understanding of Eemian environments in the North Atlantic region and helps to fill a knowledge gap about the history of the flora and fauna of the Faroe Islands, which is of biogeographical importance.     

Middle to Late Pleistocene lake‐level fluctuations of Lake El'gygytgyn,far‐east Russian Arctic

Lake El'gygytgyn, located in central Chukotka, Russian Arctic, was the subject of an international drilling project that resulted in the recovery of the longest continuous palaeoclimatic and palaeoenvironmental record for the terrestrial Arctic covering the last 3.6 million years. Here, we present the reconstruction of the lake‐level fluctuations of Lake El'gygytgyn since Marine Isotope Stage (MIS) 7 based on lithological and palynological as well as chronological studies of shallow‐water sediment cores and subaerial lake terraces. Reconstructed lake levels show an abrupt rise during glacial–interglacial terminations (MIS 6/5 and MIS 2/1) and during the MIS 4/3 stadial–interstadial transition. The most prominent lowstands occurred during glacial periods associated with a permanent lake‐ice cover (namely MIS 6, MIS 4 and MIS 2). Major triggering mechanisms of the lake‐level fluctuations at Lake El'gygytgyn are predominantly changes in air temperature and precipitation. Regional summer temperatures control the volume of meltwater supply as well as the duration of the lake‐ice cover (permanent or seasonal). The duration of the lake‐ice cover, in turn, enables or hampers near‐shore sediment transport, thus leading to long‐term lake‐level oscillations on glacial–interglacial time scales by blocking or opening the lake outflow, respectively. During periods of seasonal ice cover the lake level was additionally influenced by changes in precipitation. The discovered mechanism of climatologically driven level fluctuations of Lake El'gygytgyn are probably valid for large hydrologically open lakes in the Arctic in general, thus helping to understand arctic palaeohydrology and providing missing information for climate modelling.     

The last glacial cycles in East Greenland, an overview

Marine, fluvial and glacigene sediments exposed in coastal cliffs and stream-cut sections in East Greenland between latitudes 69° and 78° N display a record of Quaternary climatic and environmental change going back to pre-Saalian times (> 240 ka), but with main emphasis on the last interglacial/glacial cycle. The stratigraphical scheme is based on studies on the Jameson Land peninsula, and contains five glacial stages and stades with the Greenland ice sheet or its outlets reaching the outer coasts. Individual sites are correlated and dated by a combination of biostratigraphy, luminescence dating, amino acid analyses, as well as ¹⁴C- and uranium series dating. The pre-Weichselian Lollandselv and Scoresby Sund glaciations were the most extensive. During the Weichselian the Inland Ice margin in this part of East Greenland was apparently very stable. The Aucellaelv, Jyllandselv and Flakkerhuk stades mark the advance and subsequent retreat of outlet glaciers from the Inland Ice which advanced through the wide Scoresby Sund basin and reached the inner shelf. In-between the glacier advances, three interglacial or interstadial periods have been recognized. During the Langelandselv interglacia-tion (≅ Eemian) the advection of warm Atlantic water was higher than during the Holocene, and the terrestrial flora and insect faunas show that summer temperatures were 3–4°C higher than during the Holocene optimum. There is no unambiguous evidence for cooling in the sediments from this interval. Later, in isotope stage 5, there were apparently two ice-free periods. During the Hugin Sø interstade, stable Polar water dominated Scoresby Sund, and the terrestrial flora suggests summer temperatures 2° -3° lower than the present. The marine and fluvial sediments from the second ice-free period, the Mønselv interstade, are devoid of organic remains.     

Early Wisconsinan (MIS 4) Arctic ground squirrel middens and a squirrel-eye-view of the mammoth-steppe

Fossil arctic ground squirrel (Spermophilus parryii) middens were recovered from ice-rich loess sediments in association with Sheep Creek-Klondike and Dominion Creek tephras (ca 80 ka) exposed in west-central Yukon. These middens provide plant and insect macrofossil evidence for a steppe-tundra ecosystem during the Early Wisconsinan (MIS 4) glacial interval. Midden plant and insect macrofossil data are compared with those previously published for Late Wisconsinan middens dating to ～25–29¹⁴C ka BP (MIS 3/2) from the region. Although multivariate statistical comparisons suggest differences between the relative abundances of plant macrofossils, the co-occurrence of steppe-tundra plants and insects (e.g., Elymus trachycaulus, Kobresia myosuroides, Artemisia frigida, Phlox hoodii, Connatichela artemisiae) provides evidence for successive reestablishment of the zonal steppe-tundra habitats during cold stages of the Late Pleistocene. Arctic ground squirrels were well adapted to the cold, arid climates, steppe-tundra vegetation and well-drained loessal soils that characterize cold stages of Late Pleistocene Beringia. These glacial conditions enabled arctic ground squirrel populations to expand their range to the interior regions of Alaska and Yukon, including the Klondike, where they are absent today. Arctic ground squirrels have endured numerous Quaternary climate oscillations by retracting populations to disjunct “interglacial refugia” during warm interglacial periods (e.g., south-facing steppe slopes, well-drained arctic and alpine tundra areas) and expanding their distribution across the mammoth-steppe biome during cold, arid glacial intervals.     

Tepsankumpu revisited — pollen evidence of stable Eemian climates in Finnish Lapland

Several till-covered organic deposits, principally lake gyttja, in Finnish Lapland have been correlated with the last (i.e. Eemian) interglacial on the basis of their lithostratigraphic position and pollen stratigraphy. Most of the sequences are short, but together with three longer sequences from Finnish Lapland and one from Swedish Lapland (Leveäniemi) they provide a complete picture of Eemian vegetational and climatic development. The Tepsankumpu site was revisited, and the till-covered thick freshwater gyttja deposit was studied in detail for pollen in order to search for signals of rapid climatic fluctuations postulated for the earlier part of the Eemian on the basis of Greenland ice core studies. The Eemian pollen stratigraphy in Finnish Lapland closely resembles the Holocene pollen stratigraphy of the area. The abundance of spruce and alder pollen suggests, however, more northerly limits for forest vegetation zones during the Eemian than during the Holocene. Oak also grew closer to Lapland, indicating a wanner climate than during the Holocene climatic optimum. The Tepsankumpu pollen stratigraphy indicates climatic stability over the entire time-span it covers, i.e. the major part of the interglacial. This finding is in conflict with results from Greenland GRIP ice core studies and interpretations of some Continental European Eemian pollen diagrams.     

A multi‐proxy palaeoenvironmental and geochronological reconstruction of the Saalian‐Eemian‐Weichselian succession at Klein Klütz Höved,NE Germany

Here we present a multi‐proxy investigation of the Klein Klütz Höved (KKH) coastal cliff section in northeastern Germany, involving lithofacies analysis, micromorphology, micropalaeontology, palynology and luminescence dating of quartz and feldspar. We subdivide the local stratigraphy into three depositional phases. (i) Following a Saalian advance (MIS 6) of the Scandinavian Ice Sheet, the penultimate deglaciation (Termination II) at the site occurred between c. 139 and 134 ka, leading to the establishment of a braided river system and lacustrine basins under arctic‐subarctic climate conditions. (ii) In the initial phase of the Eemian interglacial lacustrine deposits were formed, containing warm‐water ostracods and a pollen spectrum indicating gradual expansion of woodlands eventually containing thermophile deciduous forest elements. A correlation of the local pollen assemblages with Eemian reference records from central Europe suggests that fewer than 750 years of the last interglacial period are preserved at KKH. The occurrence of brackish ostracods dates the onset of the Eemian marine transgression at the section at c. 300–750 years after the beginning of the last interglacial period. (iii) Directly above the Eemian record a ~10‐m‐thick sedimentary succession of MIS 2 age was deposited, implying a significant hiatus of c. 90 ka encompassing the time from middle and upper MIS 5e to late MIS 3. During the Late Weichselian, KKH featured a depositional shift from (glacio‐)lacustrine to subglacial to recessional terminoglacial facies, with the first documented Weichselian ice advance post‐dating 20±2 ka. Overall, the KKH section represents an exceptional sedimentary archive for palaeoenvironmental reconstructions, covering the period from the Saalian glaciation and subsequent Termination II to the early Eemian and Late Weichselian. The results refine the existing palaeogeographical and geochronological models of the late Quaternary history in the southwestern Baltic Sea area and allow correlations with other reference records in a wider area.     

Weichselian and Holocene palaeoenvironmental history of the Bol'shoy Lyakhovsky Island, New Siberian Archipelago, Arctic Siberia

Cryolithological, ground ice and fossil bioindicator (pollen, diatoms, plant macrofossils, rhizopods, insects, mammal bones) records from Bol'shoy Lyakhovsky Island permafrost sequences (73°20′N, 141°30′E) document the environmental history in the region for the past c. 115 kyr. Vegetation similar to modern subarctic tundra communities prevailed during the Eemian/Early Weichselian transition with a climate warmer than the present. Sparse tundra‐like vegetation and harsher climate conditions were predominant during the Early Weichselian. The Middle Weichselian deposits contain peat and peaty soil horizons with bioindicators documenting climate amelioration. Although dwarf willows grew in more protected places, tundra and steppe vegetation prevailed. Climate conditions became colder and drier c. 30 kyr BP. No sediments dated between c. 28.5 and 12.05 ¹⁴C kyr BP were found, which may reflect active erosion during that time. Herb and shrubby vegetation were predominant 11.6–11.3 ¹⁴C kyr BP. Summer temperatures were c. 4 °C higher than today. Typical arctic environments prevailed around 10.5 ¹⁴C kyr BP. Shrub alder and dwarf birch tundra were predominant between c. 9 and 7.6 kyr BP. Reconstructed summer temperatures were at least 4 °C higher than present. However, insect remains reflect that steppe‐like habitats existed until c. 8 kyr BP. After 7.6 kyr BP, shrubs gradually disappeared and the vegetation cover became similar to that of modern tundra. Pollen and beetles indicate a severe arctic environment c. 3.7 kyr BP. However, Betula nana, absent on the island today, was still present. Together with our previous study on Bol'shoy Lyakhovsky Island covering the period between about 200 and 115 kyr, a comprehensive terrestrial palaeoenvironmental data set from this area in western Beringia is now available for the past two glacial–interglacial cycles.     

Climatic bisection of the last interglacial warm period in the Polar North Atlantic

New multiproxy marine data of the Eemian interglacial (MIS5e) from the Norwegian Sea manifest a cold event with near-glacial surface ocean summer temperatures (3–4 °C). This mid-Eemian cooling divided the otherwise relatively warm interglacial climate and was associated with widespread expansions of winter sea-ice and polar water masses due to changes in atmospheric circulation and ocean stability. While the data also verify a late rather than early last interglacial warm peak, which is in general disharmony with northern hemisphere insolation maximum and the regional climatic progression of the early Holocene, the cold event itself was likely instrumental for delaying the last interglacial climate development in the Polar North when compared with regions farther south. Such a ‘climatic decoupling’ of the Polar region may bear profound implications for the employment of Eemian conditions to help evaluate the present and future state of the Arctic cryosphere during a warming interglacial.     

Amino acid ratios in Quaternary molluscs and foraminifera from western Norway: correlation, geochronology and paleotemperature estimates

Isoleucine epimerization (alle/Ue) ratios in the pelecypod Mya truncata and benthic foraminifer Cibicides lobalulus from emerged marine units in western Norway allow construction of a regional relative chronostratigraphy for the Ecmian and Weichselian. Two in situ interglacial sections are considered correlative by the similar biostratigraphy and alle/Ile ratios in C. lobalulus. Overlying sediments at the two sites are of both marine and glacial origin. Neither site contains a complete Weichselian record, but allelic ratios, lithostratigraphy and fauna! changes suggest at least four stadial and three interstadial events occurred along the western Norwegian coast during Early and Middle Weichselian time. Kinetic data defining the relationship between the isoleucine epimerization rate constant and temperature for the species studied allow the estimation of paleotemperatures for samples of known age. Accepting published age estimates for the Eemian interglacial beds, the average Weichselian temperature in western Norway is calculated to have been ca. 4°C below the average Holocene temperature, whereas the last interglacial was 1 to 2°C warmer that the Holocene. The limited temperature depression over this region during the Weichselian implies that coastal western Norway was ice-covered only about 30% of this period, and that Atlantic water, although not necessarily in a warm surface current as today, entered the Norwegian Sea during much of marine isotope stage 5 and intermittently during stage 3. Interpolated amino acid ages date interstadial events at ca. 94 ka, 78 ka and 52 ka, B.P., whereas glacial events are dated ca. 103 ka and bracketed by limiting dates between 78 and 89 ka, between 52 and 63 ka and less than 36 ka B.P.     

Late Quaternary history of the Kap Mackenzie area,northeast Greenland

Wagner, B., Bennike, O., Cremer, H. & Klug, M. 2010: Late Quaternary history of the Kap Mackenzie area, northeast Greenland. Boreas, Vol. 39, pp. 492–504. 10.1111/j.1502‐3885.2010.00148.x. ISSN 0300‐9483. The Kap Mackenzie area on the outer coast of northeast Greenland was glaciated during the last glacial stage, and pre‐Holocene shell material was brought to the area. Dating of marine shells indicates that deglaciation occurred in the earliest Holocene, before 10 800 cal. a BP. The marine limit is around 53 m a.s.l. In the wake of the deglaciation, a glaciomarine fauna characterized the area, but after c. one millennium a more species‐rich marine fauna took over. This fauna included Mytilus edulis and Mysella sovaliki, which do not live in the region at present; the latter is new to the Holocene fauna of northeast Greenland. The oldest M. edulis sample is dated to c. 9500 cal. a BP, which is the earliest date for the species from the region and indicates that the Holocene thermal maximum began earlier in the region than previously documented. This is supported by driftwood dated to c. 9650 cal. a BP, which is the earliest driftwood date so far from northeastern Greenland and implies that the coastal area was at least partly free of sea ice in summer. As indicated by former studies, the Storegga tsunami hit the Kap Mackenzie area at c. 8100 cal. a BP. Loon Lake, at 18 m a.s.l., was isolated from the sea at c. 6200 cal. a BP, which is distinctly later than expected from existing relative sea‐level curves for the region.     

Interglacial remains of caribou (Rangifer tarandus) and lemming (Dicrostonyx torquatus(?)) from North Greenland

A caribou antler from Kap København in Peary Land has been radiocarbon dated at > 40,000 B.P. A corresponding infinite date was obtained on allochthonous plant and animal remains, including five lemming droppings from Skalhøjene in Warming Land. These remains are indicative of high arctic conditions much like the present environment of the region. Because the caribou antler and the remains from Skalhojene point to biotic and climatic conditions no more severe than modern conditions, an interglacial age is suggested, and we propose that they belong to the Sangamonian Stage. The finds indicate that evolution of high arctic subspecies such as Rangifer tarandus pearyi and Dicrostonyx torquatus groenlandicus probably took place in pre-Wisconsinan times. It is unlikely that caribou and lemming survived in North Greenland during the last full glacial period. They probably re-immigrated in the early Holocene.     

Retreat of the Smith Sound Ice Stream in the Early Holocene

Nares Strait, a major connection between the Arctic Ocean and Baffin Bay, was blocked by coalescent Innuitian and Greenland ice sheets during the last glaciation. This paper focuses on the events and processes leading to the opening of the strait and the environmental response to establishment of the Arctic‐Atlantic throughflow. The study is based on sedimentological, mineralogical and foraminiferal analyses of radiocarbon‐dated cores 2001LSSL‐0014PC and TC from northern Baffin Bay. Radiocarbon dates on benthic foraminifera were calibrated with ΔR = 220±20 years. Basal compact pebbly mud is interpreted as a subglacial deposit formed by glacial overriding of unconsolidated marine sediments. It is overlain by ice‐proximal (red/grey laminated, ice‐proximal glaciomarine unit barren of foraminifera and containing >2 mm clasts interpreted as ice‐rafted debris) to ice‐distal (calcareous, grey pebbly mud with foraminifera indicative of a stratified water column with chilled Atlantic Water fauna and species associated with perennial and then seasonal sea ice cover) glacial marine sediment units. The age model indicates ice retreat into Smith Sound as early as c. 11.7 and as late as c. 11.2 cal. ka BP followed by progressively more distal glaciomarine conditions as the ice margin retreated toward the Kennedy Channel. We hypothesize that a distinct IRD layer deposited between 9.3 and 9 (9.4–8.9 1σ) cal. ka BP marks the break‐up of ice in Kennedy Channel resulting in the opening of Nares Strait as an Arctic‐Atlantic throughflow. Overlying foraminiferal assemblages indicate enhanced marine productivity consistent with entry of nutrient‐rich Arctic Surface Water. A pronounced rise in agglutinated foraminifers and sand‐sized diatoms, and loss of detrital calcite characterize the uppermost bioturbated mud, which was deposited after 4.8 (3.67–5.55 1σ) cal. ka BP. The timing of the transition is poorly resolved as it coincides with the slow sedimentation rates that ensued after the ice margins retreated onto land.     

Tracer transport in the Greenland Ice Sheet: constraints on ice cores and glacial history

The climate history and dynamics of the Greenland Ice Sheet are studied using a coupled model of the depositional provenance and transport of glacier ice, allowing simultaneous prediction of the detailed isotopic stratigraphy of ice cores at all the major Greenland sites. Adopting a novel method for reconstructing the age–depth relationship, we greatly improve the accuracy of semi-Lagrangian tracer tracking schemes and can readily incorporate an age-dependent ice rheology. The larger aim of our study is to impose new constraints on the glacial history of the Greenland Ice Sheet. Leading sources of uncertainty in the climate and dynamic history are encapsulated in a small number of parameters: the temperature and elevation isotopic sensitivities, the glacial–interglacial precipitation contrast and the effective viscosity of ice in the flow law. Comparing predicted and observed ice layering at ice core sites, we establish plausible ranges for the key model parameters, identify climate and dynamic histories that are mutually consistent and recover the past depositional elevation of ice cores to ease interpretation of their climatic records. With the coupled three-dimensional model of ice dynamics and provenance transport we propose a method to place all the ice core records on a common time scale and use discrepancies to adjust the reconstructed climate history. Analysis of simulated GRIP ice layering and borehole temperature profiles confirms that the GRIP record is sensitive to the dynamic as well as to the climatic history, but not enough to strongly limit speculation on the state of the Greenland Ice Sheet during the Eemian. In contrast, our study indicates that the Dye 3 and Camp Century ice cores are extremely sensitive to ice dynamics and greatly constrain Eemian ice sheet reconstructions. We suggest that the maximum Eemian sea-level contribution of the ice sheet was in the range of 3.5–4.5 m.     

Climatic reconstructions from the Eemian/Early Weichselian transition in Central Europe based on the coleopteran record from Gröbern, Germany

At Grobern (51°52'N, 12°6'E, altitude 94–98 m a.s.l., 50km north of Leipzig), a succession of lacustrine sediments has yielded a fossil coleopterous fauna permitting the reconstruction of climatic conditions through out the Eemian/Early Weichselian transition. In accord with other indicators, the Coleoptera show that this period was characterized by three major climatic oscillations during which the thermal climate fluctuated between conditions of arctic severity and more temperate conditions. During the cold episodes, the beetle faunas were dominated by species which today have exclusively northern or even Asiatic ranges. In the intervening warm periods these cold-adapted species were absent and the faunas included temperate species some of which are tree-dependent. Quantitative palaeotemperature estimates using the Mutual Climatic Range method, show that the mean July temperatures during the relatively colder intervals were 5°C to 6°C lower than during the warmer periods, but the depression of winter temperatures during the colder periods was much greater than this. Because different species of Coleoptera have different thresholds of temperature tolerance, their indications of climatic change do not always coincide with the changes in the lithology or pollen.     

Different types of ending to the last two interglacials in western Europe

Tidal deposits dating from the penultimate interglacial (Oxygen Isotope Stage 7) are preserved on a bank of the River Seine at Tancarville. The foraminiferal assemblages indicate a gradual cooling towards Arctic conditions, which corresponds to the onset of the penultimate cold period. This cooling started before or during the drop in sea level in Europe, in contrast to the situation at the end of the last interglacial (Eemian) when Europe remained warm even during the period of falling sea level.     

Palaeoceanographic changes in the North Atlantic during the Mid‐Pleistocene Transition (MIS 31–19) as inferred from planktonic foraminiferal and calcium carbonate records

Marine sediments from the Integrated Ocean Drilling Project (IODP) Site U1314 (56.36°N, 27.88°W), in the subpolar North Atlantic, were studied for their planktonic foraminifera, calcium carbonate content, and Neogloboqudrina pachyderma sinistral (sin.) δ¹³C records in order to reconstruct surface and intermediate conditions in this region during the Mid‐Pleistocene Transition (MPT). Variations in the palaeoceanography and regional dynamics of the Arctic Front were estimated by comparing CaCO₃ content, planktonic foraminiferal species abundances, carbon isotopes and ice‐rafted debris (IRD) data from Site U1314 with published data from other North Atlantic sites. Site U1314 exhibited high abundances of the polar planktonic foraminifera N. pachyderma sin. and low CaCO₃ content until Marine Isotope Stage (MIS) 26, indicating a relatively southeastward position of the Arctic Front (AF) and penetration of colder and low‐salinity surface arctic water‐masses. Changing conditions after MIS 25, with oscillations in the position of the AF, caused an increase in the northward export of the warmer North Atlantic Current (NAC), indicated by greater abundances of non‐polar planktonic foraminifera and higher CaCO₃. The N. pachyderma sin. δ¹³C data indicate good ventilation of the upper part of the intermediate water layer in the eastern North Atlantic during both glacial and interglacial stages, except during Terminations 24/23, 22/21 and 20/1. In addition, for N. pachyderma (sin.) we distinguished two morphotypes: non‐encrusted and heavily encrusted test. Results indicate that increases in the encrusted morphotype and lower planktonic foraminiferal diversity are related to the intensification of glacial conditions (lower sea‐surface temperatures, sea‐ice formation) during MIS 22 and 20.     

Paleoclimatology and paleo-oceanography of the Norwegian and Greenland seas: glacial-interglacial contrasts

Synoptically mapped faunal abundance and faunal composition data, derived from a suite of 24 Norwegian Sea cores, were used to derive sea-surface temperatures for the last glacial maximum (18,000 B.P.), the last interglacial (120,000 B.P.), and isotope stage 5a (82,000 B.P.). Surface circulation and ice cover reconstructions for these three times, deduced from the sea-surface temperatures, suggest the following conclusions: (1) During glacial periods, Norwegian Sea surface circulation formed a single, sluggish, counterclockwise gyre that was caused by wind drag on the ubiquitous sea ice cover; (2) the last interglacial was characterized by a circulation pattern similar to that of today except that the two counterclockwise gyres were displaced toward the east and were more vigorous than they are today. This circulation pattern forced the Norwegian Current into a position close to the coast of Norway and permitted formation of a strong east-west temperature gradient close to the Scandinavian landmass; (3) interglacial periods prior to 120,000 B.P. had similar climatic conditions to the 82,000 B.P. level and were characterized by a weak two-gyre circulation pattern. The southern gyre, driven by wind stress in summer months, was ice covered in winters. The northern gyre had little open water even in summers and was primarily formed by wind drag on sea ice. Atmospheric modifications resulting from these circulation patterns and sea ice conditions produced varying climatic conditions in Scandinavia during interglacials prior to the Holocene. The climate was probably warmer and moister during the last interglacial (Eemian) than it is today. Other interglacials during the last 450,000 years, but prior to the Eemian, were probably colder and drier as the Norwegian Sea was not an important source of heat and moisture.