Stratigraphic significance of the Cambrian ichnofauna of the Zanskar region,Ladakh Himalaya,India

The Cambrian succession in the Zanskar Basin of Tethys Himalaya contains an abundant ichnofossils like in the other Tethyan Himalayan successions. The ichnofossils are stratigraphically important as they occur below the trilobite body fossils and are useful to define the basal part of the Cambrian. The ichnofossil assemblage reported from the Zanskar Basin of Ladakh Himalaya is significant to demarcate the Early Cambrian age due to lack of other faunal elements so far. The body fossils of trilobites recorded from the overlying beds indicates the earliest part of the Middle Cambrian age. Sixteen ichnogenera identified include: Bifungites, Cruziana, Diplichnites, Dimorphichnus, Isopodichnus, Lockeia, Merostomichnites; Monomorphichnus, Psammichnites, Palaeophycus, Planolites, Rhizocorallium, Skolithos, Taphrhelminthopsis, Teichichnus, Trepitichnus and trilobite scratch marks etc. The ichnogenera reported so far from this part of the Tethyan Himalayan region belongs mostly to the traces of arthropod origin. The ichnofauna ranges in age from Lower Cambrian to late part of the Middle Cambrian. The ichnofaunal assemblage can be assigned to repichnial, cubichinial, pascichnial, to fodinichnial behaviour. The distribution of ichnofossils in the studied sections shows that the ichnofossils are predominately less in occurrence in the sections were trilobites dominates and higher in the successions the abundance of ichnofossils decreases.     

Additional late Middle Cambrian trilobites from Karsha Formation (Haimanta Group) Zanskar region of Zanskar-Spiti basin, Northwest Himalaya

Additional latest Middle Cambrian polymerid trilobites are recorded from the Teta Member (Karsha Formation) along the Tangze-Yogma-Kuru section, Kurgiakh Valley, southeastern Zanskar, Northwest Himalaya. It includes record of a new genus Pianspis Saito & Sakakuru, 1936 of family Lisianiidae and others associated fauna comprises of Neoanomocarella asiatica (Lisianiidae), Parablackwelderia sp., (Damesellidae) and two undetermined pygidium and one unidentified cranidia. All these specimens are collected from the level representing the Lejopyge acantha Zone of Guzhangian Stage (Cambrian Series 3) of the Middle Cambrian. The fauna is mainly correlated with the South China, North Korea, Kazakhastan, Uzbekistan, Siberia and Australia.     

祁连和秦岭地区寒武纪和奥陶纪古生物区系关系的探讨

我国西北地区由不同地体组成,构造运动复杂。由于缺乏充分的古生物学证据,古生物区系的识别和归属一直是有待解决的问题。本文在前人工作的基础上,依据各类动物群特征的分析,较详细地探讨了我国祁连和秦岭地区相关地层区中-晚寒武世和奥陶纪动物群的性质及其古生物类型的归属,确定了过渡型动物群在祁连和秦岭地区的大体分布范围。同时,结合地球动力学和古地理学研究成果的分析,探讨了导致在祁连和秦岭地区中-晚寒武世和奥陶纪时存在着与华南相似过渡类型动物群的两种可能原因。一种可能是中-晚寒武世和奥陶纪时塔里木板块位于扬子板块西边,中朝板块位于扬子板块之北,祁连和秦岭可能位于扬子和塔里木板块之间,存在与华南及华北动物群交流的基本条件,产生了大体相近的动物群。另一种可能是中-晚寒武世和奥陶纪时祁连和秦岭地区存在一些原位于扬子板块北部边缘的地体,在这些沿扬子板块北缘分布的地体上发育了与南部边缘相似的生物群或生物类型。中生代之前,沿扬子板块主体和北部边缘地体之间的走滑活动,可能使扬子板块主体向东移动了几百公里,而其边缘那些属于祁连和秦岭的地体向东移动的幅度不大,从而造成现今祁连和秦岭地区存在与扬子板块东南部地区大体相似的具过渡性质的生物群或生物类型,而扬子板块北部边缘又缺少边缘过渡相带的现状。     

Integrated ichnological and sedimentological studies of the Parahio Formation (Cambrian) of the Zanskar region (Zanskar-Spiti Basin), Northwest Himalaya

Lower-Middle Cambrian (Lungwangmiaoan-Hsuchuangian stage) Parahio Formation (Kunzam La Formation) exposed at Purni Village, Niri-Tsarap Chu Valley of Zanskar region of Tethyan Himalaya has yielded a high diversity and abundant ichnofossils with myriapod trackways. It includes Cruziana isp., Phycodes palmatum, Diplocraterion isp., Diplichnites isp., Dimorphichnus isp., Tapherhelminthopsis cf. circularis, Teichichnus isp., Monomorphicnus isp., Lockeia isp., Skolithos isp., Planolites isp., Palaeophycus isp., Isopodichnus isp., and arthropod appendage marks. Integration of ichnological and sedimentological studies for measured part of the Parahio Formation (Cambrian) reveals that alternating energy conditions due to recurring storm events were superimposed on low energy fair-weather sediments. The post-depositional Arenicolites ichnofacies, preserved within the high-energy storm beds of shoreface deposits, commonly overlies the fair-weather assemblage of pre-depositional Cruziana ichnofacies of relatively more offshore deposits, which contains more diverse and varied behavioral signatures of various deposit feeders. The traces of the Cruziana ichnofacies are abruptly replaced by traces of Arenicolites ichnofacies and show reiteration throughout the measured part of the Parahio Formation. The ichnofabric indices in these deposits range from ii3–ii5. No discrete sets of tiers are observed; perhaps there were two coeval communities. Ichnologically, an idealized wave-dominated progradational cycle in the Parahio Formation is characterized, from base to top, by a Cruziana ichnofacies (lower-upper offshore to offshore transition) and storm related Arenicolites ichnofacies (shoreface). Each progradational cycle reflects a progressive increase in sand content, degree of oxygenation, hydrodynamic energy and dearth of food. These environmental factors controlled the vertical distribution of trace fossils in Parahio Formation. The occurrence of Cruziana ichnoassociation much below the Middle Cambrian trilobite horizon in the Parahio Formation restricts the age of this part of the formation to the pre-Middle Cambrian, probably series 2 of Cambrian system.     

A middle Cambrian shallow water trilobite fauna from the Comstock Formation,near Queenstown,western Tasmania

A limestone lens within the Comstock Formation, from low in the Tyndall Group near the Lyell Comstock Mine near Queenstown, western Tasmania, contains a small trilobite fauna. The trilobites include Ammagnostus cf. laiwuensis, Kootenia sp., Sudanomocarina? sp. and a possible member of the Monkaspidae. These suggest an age somewhere in the range from the upper Ptychagnostus atavus Zone to the Lejopyge laevigata Zone, with regional stratigraphic correlations suggesting the latter zone. This is only the second Tasmanian Cambrian fossil assemblage described from limestone; almost all previously described Tasmanian Cambrian faunas are found in siltstones and shales.     

Early Cambrian palaeobiogeography of the Zhenba–Fangxian Block (South China): Independent terrane or part of the Yangtze Platform?

Early Cambrian small skeletal fossils (SSFs) are studied and revised from the Zhenba–Fangxian Block of the transitional zone between the Yangtze Block and the South Qinling Terrane. The study reveals a diverse fauna with 47 species of various biological affinities, including the new species Gapparodus gapparites sp. nov. The SSFs are assigned to the newly defined Cambroclavus fangxianensis–Rhombocorniculum cancellatum Assemblage Zone. Based on the investigated SSF fauna from Zhenba County, Southeast Shaanxi of China and published data, a palaeobiogeographic study is carried out for the Cambrian Stage 3 (equivalent to the Atdabanian–Botoman of Siberia). A hierarchical Pearson similarity cluster analysis of 295 species from 32 regions of the world indicates a distinct palaeobiogeographic pattern with seven faunal provinces. The result is mostly consistent with existing palaeogeographic reconstructions for the early Cambrian. However, it is also shown that the SSF assemblages of the Zhenba–Fangxian Block have low similarity with those of the Yangtze Block. Instead, they share high similarity with those from Armorica, Tarim and the Karatau–Naryn terranes (South Kazakhstan/North Kyrgyzstan). The Yangtze Block has a unique SSF assemblage dissimilar to most of other regions. The Terreneuvian–Cambrian Stage 3 sedimentary sequence of the Zhenba–Fangxian Block is more consistent with that of the South Qinling Terrane. Besides, sedimentary Ediacaran manganese ore deposits and Cambrian barite/witherite deposits have unique distribution pattern on the Zhenba–Fangxian Block. Derived from the profound dissimilarities in faunal composition, sedimentary sequence and distribution of sedimentary ore deposits, we hypothesize that during the Neoproterozoic–Cambrian transition, the Zhenba–Fangxian Block might have been an independent terrane and more distant from the Yangtze Block. The palaeobiogeographic analysis of SSFs also indicates a closer alliance between Avalonia and Siberia. It corroborates the palaeogeographic reconstruction of North China at the margin of Gondwana, in the vicinity of Australia, Antarctica, and Armorica.     

Peri-Gondwana Late Early–Middle Cambrian trilobites from the Kuhbanan Formation in Dahu section,Kerman Province,Iran

Outcropped of the Kuhbanan Formation at Dahu, near Zarand, about 63 km north of Kerman, Iran contains peri-Gondwana trilobites. In this study, 185 trilobite samples including six species and genera were identified and described from Dahu section. This trilobite’s assemblage including Redlichia noetlingi, Redlichia sp., Kermanella kuhbananensis, Kermanella lata lata, Kermanella lata minuta, Iranoleesia pisiformis, and Iranaspis sp. based on occurrence of the trilobite fauna a late Early to Middle Cambrian (Series 2–3) is suggested for this strata. These trilobite fauna help confirm conclusions from recent geological studies that place the Kerman Basin of Iran during the Cambrian.     

峡东寒武纪三叶虫化石新材料

三峡东部中寒武统化石稀少。本文对在宜昌三斗坪中寒武世覃家庙组二段下部含鲕粒灰岩、细晶灰岩中所采获的三叶虫Xingrenaspis sp.进行了研究和描述,并对含化石地层时代进行了讨论。此次发现,丰富了峡东中寒武世的化石资料,也为本区中寒武世地层划分对比提供了新的依据。     

Cambrian stratigraphy of Kashmir and its boundary problems

The biostratigraphy of the Cambrian of Kashmir based on trilobite assemblage zones and trace fossils has been attempted. Faunal gaps occur in lower part of Lower Cambrian, upper Lower Cambrian, lower Middle Cambrian and upper part of Upper Cambrian. In the Lower Cambrian Cruziana—Rusophycus and Redlichia Zones, in the Middle Cambrian Solenopleura—Tonkinella, Tonkinella—Anomocare, Anomocare—Bailiella and Bolaspidella Zones, and in the Upper Cambrian Chuangia and Dikelocephalites Zones are recognized. The position of intrasystem boundaries is also discussed. While the various taxa have lithological preferences and are not necessarily found in a continuous sequence, an attempt has been made to interpret the ranges of different genera. The affinities of the fauna with that of other Cambrian basins of the world are discussed. It is concluded that the bulk of the fauna is of a cosmopolitan nature.     

The oldest bivalved arthropods from the early Cambrian of East Gondwana: Systematics,biostratigraphy and biogeography

The oldest bradoriid fauna from Australia, occurring in the lower Cambrian Ajax and Wirrapowie limestones of the Flinders Ranges, South Australia consists of eleven taxa, including one new genus and species, Quadricona madonnae gen. et sp. nov. and two new species, Liangshanella circumbolina sp. nov. and Zepaera jagoi sp. nov. In the Ajax Limestone, Liangshanella circumbolina sp. nov. occurs c. 20 m below the FAD of the zonal trilobite Abadiella huoi. This pre-trilobitic occurrence represents the oldest bivalved arthropod hitherto known from East Gondwana and suggests a lower Cambrian (Series 2, Stage 3) age for the assemblage. The recognition of distinct bradoriid assemblages associated with the Abadiella huoi (Atdabanian), Pararaia tatei, P. bunyerooensis and P. janeae (all Botoman) trilobite biozones in South Australia indicates great potential for future regional biostratigraphic correlation. Quantitative biogeographic analysis including new taxonomic data from the lower Cambrian of South Australia, highlights the strong endemism displayed by early Cambrian bradoriid communities and strengthens the close faunal affinities with South China and Antarctica.     

Trilobite trace fossils and their stratigraphical significance in the Cambrian sequence of the Holy Cross Mountains,Poland

Abundant and varied trilobite trace fossils are recognized and described in the Cambrian sequence of the Holy Cross Mountains. Cruziana and Rusophycus are stratigraphically significant and some of the described traces may be used as local index fossils. The first recognition of the Cruziana barbata ichnozone in Poland is made and it occurs in the uppermost Middle Cambrian. A new position for the Middle/Upper Cambrian boundary is established. Three new ichnospecies are described: Cruziana magna, C. regularis and Rusophycus crebus.     

Planktic Foraminiferal Biostratigraphy of the Cretaceous Oceanic Red Beds in Kangmar,Southern Tibet,China

The planktic foraminifera of the Chuangde Formation (Upper Cretaceous Oceanic Red Beds, CORBs) as exposed at Tianbadong section, Kangmar, southern Tibet has been firstly studied for a detailed for a detailed biostratigraphy elaboration. A rich and well-preserved planktic foraminifera were recovered from the Chuangde Formation of the Tianbadong section and the Globotruncanita elevata, Globotruncana ventricosa, Radotruncana calcarata, Globotruncanella havanensis, Globotruncana aegyptiaca, Gansserina gansseri and Abathomphalus mayaroensis zones have been recognized. The planktic foraminiferal assemblage points to an early Campanian to Maastrichitian age for the CORBs of the eastern North Tethyan Himalayan sub-belt, which also provides a better understanding of the shifting progress of the Indian Plate to the north and the evolution of the Neotethyan ocean. The lithostratigraphy of the Chuangde Formation of the Tianbadong section comprises two lithological sequences observed in ascending succession: a lower unit (the Shale Member) mainly composed of purple (cherry-red, violet-red) shales with interbedded siltstones and siliceous rocks; and an upper unit (the Limestone Member) of variegated limestones. The strata of the Chuangde Formation in the Tianbadong section are similar to CORBs in other parts of the northern Tethyan Himalaya area of Asia (Gyangze, Sa’gya, Sangdanlin, northern Zanskar, etc.). The fossil contents of the Chuangde Formation in the sections (CORBs) studied provide a means of correlation with the zonation schemes for those of the northern Tethyan Himalayan sub-belt and the Upper Cretaceous of the southern Tethyan Himalayan sub-belt. Paleogeographic reconstruction for the Late Cretaceous indicates that the Upper Cretaceous Chuangde Formation (CORBs) and correlatable strata in northern Zanskar were representative of slope to basinal deposits, which were situated in the northern Tethyan Belt. Correlatable Cretaceous strata in Spiti and Gamba situated in the southern Tethyan Belt in contrast were deposited in shelf environments along the Tethyan Himalayan passive margin. CORBs are most likely formed by the oxidation of Fe(II)-enriched, anoxic deep ocean water near the chemocline that separated the oxic oceanic surface from the anoxic.     

Stratigraphic Implications of Skeletal Microfossils from the Cambrian of Korea: A Preliminary Report

Diverse small shelly fossils and other associated fossils were recently recovered from the Cambrian Hanaeri section, southwestern Mungyeong, Korea. The fauna includes conoidal problematica, poriferans (sponge spicules), coeloscleritophorans (chancelloriids), brachiopods, monoplacophorans, trilobite (?) fragments, echinoderms, and conodonts. A preliminary assessment of faunal associations, stratigraphic implications, and the correlation of these skeletal fossils is given, based on occurrences of Actinotheca cf. mira (He), Microcornus sp., Torellela laevigata (Linnarsson), Torellela sp., Archiasterella quadratina Lee, Chancelloria sp., Lingulella sp., Prototreta sp., Phakelodus tenuis (Müller), Phakelodus elongates (An), Hertzina sp., and Furnishina sp.     

Thermal model for the Zanskar Himalaya

ABSTRACT Crustal thickening along the northern margin of the Indian plate, following the 50 Ma collision along the Indus Suture Zone in Ladakh, caused widespread high-temperature, medium-pressure Barrovian facies series metamorphism and anatexis. In the Zanskar Himalaya metamorphic isograds are inverted and structurally telescoped along the Main Central Thrust (MCT) Zone at the base of the High Himalayan slab. Along the Zanskar valley at the top of the slab, isograds are the right way-up and are also telescoped along northeast-dipping normal faults of the Zanskar Shear Zone (ZSZ), which are related to culmination collapse behind the Miocene Himalayan thrust front. Between the MCT and the ZSZ a metamorphic-anatectic core within sillimanite grade rocks contains abundant leucogranite-granite crustal melts of probable Himalayan age. A thermal model based on a crustal-scale cross-section across the Zanskar Himalaya suggests that M₁ isograds, developed during early Himalayan Barrovian metamorphism, were overprinted during high-grade MCT-related anatexis and folded around a large-scale recumbent fold developed in the hanging wall of the MCT.     

The oldest trilobites in Cambria: Early Cambrian trilobite faunas from the Llanberis Slates Formation,Gwynedd, North Wales

Historical published works on the fossil fauna of the Llanberis Slates Formation in Gwynedd, North Wales, concentrated on the endemic trilobite Pseudatops viola (Woodward, 1888), and placed the formation towards the top of Stage 3 (Series 2) of the Cambrian chronostratigraphy.The impression given was that the fauna is not particularly diverse and fossils are rare. However, recent collecting has produced a comprehensive fauna of sponges, hyolithids and diverse arthropods including over 250 trilobite specimens from two localities of the uppermost Green Slate horizon. Among them are examples of the endemic trilobite Pseudatops viola, sufficiently well preserved to enable a reappraisal of its status. The most abundant trilobite is a form of Strenuella (cf. strenua), but the collection includes previously unrecorded eodiscoid trilobites and specimens representing unrecorded and potentially new taxa. These additions result in the most comprehensive lower Cambrian trilobite assemblage yet described from Wales, and place it in the lower part of Stage 4, Series 2 in standard global terms.     

Trilobite Biostratigraphy of the lower Paleozoic (Cambrian–Ordovician) Joseon Supergroup, Taebaeksan Basin, Korea

In Korea,trilobites are among the most intensively studied fossil groups in the past century and provide invaluable information about lower Paleozoic stratigraphy,paleogeography,and tectonics of the Korean Peninsula. Trilobites occur in the lower Paleozoic Joseon Supergroup of the Taebaeksan Basin which was part of the Sino-Korean Craton in the Paleozoic. The Joseon Supergroup is divided into the Taebaek,Yeongwol,and Mungyeong groups. The Taebaek and Yeongwol groups are richly fossiliferous,while the Mungyeong Group is poorly fossiliferous. Contrasting trilobite faunal contents of the Taebaek and Yeongwol groups resulted in two separate biostratigraphic schemes for the Cambrian–Ordovician of the Taebaeksan Basin. A total of 22 biozones or fossiliferous horizons were recognized in the Taebaek Group; 19 zones were established in the Yeongwol Group; and four biozones were known from the Mungyeong Group. These trilobite biozones of the Taebaeksan Basin indicate the Joseon Supergroup ranges in age from the Cambrian Series 2 to Middle Ordovician and can be correlated well with the formations of North China,South China,and Australia.     

Systematics, biometry of the species Opsidiscus from the Middle Cambrian succession of the Spiti basin, India

The Middle Cambrian succession of the Parahio Valley in general and Debsakhad section in particular is dominated by trilobites. Genus Opsidiscus, along with other trilobite forms dominates the lower Middle Cambrian succession. In the present study two new species of Opsidiscus, Opsidiscus wadiai and Opsidiscus srikantiai are reported from the lower Middle Cambrian succession of Debsakhad section. The species were differentiated on their morphological features and their multivariate analysis. The multivariate analysis applied here is used as a supplement method to qualitative analysis in order to differentiate between the cranidial characters of Opsidiscus. The qualitative study of each species studied individually shows more or less close affiliation as is observed by different quantitative methods. The presence of Opsidiscus has a great stratigraphic significance in this region, as they first appear from informal Stage 5 of Series 3 and goes up to the Drumian Stage of Series 3 of the Cambrian System in the Debsakhad section, which helps to correlate this section with other well known sections of the Middle Cambrian.     

Structural evolution and sequence of thrusting in the High Himalayan, Tibetan—Tethys and Indus suture zones of Zanskar and Ladakh, Western Himalaya

The timing of motion on major thrusts in the Western Himalaya shows an extremely complex sequence that spans approximately 70 Ma from the latest Cretaceous throughout the Tertiary. Three major phases of thrusting can be distinguished. The earliest phase (T1) is associated with emplacement of Tethyan basin thrust sheets (Lamayuru sediments and Spontang ophiolite) south and south-westwards onto the submerged northern passive margin of India (75-60 Ma). Collision between India and Asia occurred at 50-36 Ma and was followed immediately by the major phase (T2) of crustal shortening involving large-scale south and south-westward directed thrusting of the complete Palaeozoic, Mesozoic and Late Tertiary Tibetan—Tethys zone rocks. Preliminary balanced cross-sections show a minimum shortening of 126 km of these rocks across the Zanskar Range. The late collision phase (T3) involved re-thrusting of the previously stacked pile (breaching or leap-frog thrusting) reversing the earlier stacking order in places, and widespread steepening, overturning and backthrusting along the whole northern margin of the Tibetan—Tethys zone and throughout the Indus suture zone.     

Newly Discovered Fish Faunas from the Early Triassic, Karoo Basin, South Africa, and their Correlative Implications

Recent collecting in exposures of the lowermost Burgersdorp Formation (Beaufort Group), of the Karoo Basin of South Africa, has revealed a previously unknown fish fauna from the Early Triassic (Scythian), lowermost Cynognathus Assemblage Zone (CAZ), which forms an important component of the total vertebrate assemblage. The newly discovered fish material includes lungfish, saurichthyids, and a large microfauna that includes numerous isolated chondrichthyan teeth, two fin spine fragments, and actinopterygian scales and teeth. The latest fish finds, together with the lowermost Cynognathus Assemblage Zone vertebrate faunas, make this Karoo Basin Assemblage Zone one of the most diverse Early Triassic faunal assemblages, comparable in faunal diversity to those from the Czatkowice Formation (Poland) and the Arcadia Formation (Australia). The presence of the lungfish Ptychoceratodus phillipsi in the early Middle Triassic Cynognathus Assemblage Zone (Subzone B), and in the underlying latest Early Triassic Cynognathus Assemblage Zone (Subzone A), indicates that these lungfish could serve as range index fossils within the CAZ, and thus are potentially useful biostratigraphic markers across the Early-Middle Triassic boundary. Furthermore the ‘new’ fish fauna provides a vital marine realm link in particular with the faunas of Madagascar and Australia, that is unavailable using the tetrapod faunal elements of the lower CAZ.     

Cambro‐Ordovician trace fossils from the SW‐Norwegian Caledonides

A diverse trace fossil association is described for the first time from low‐grade metamorphic rocks of the SW‐Norwegian Caledonides. The investigated cliff sections with autochthonous to parautochthonous metasediments comprise a coarsening‐ and thickening‐upward succession interpreted as prograding delta deposits. Sedimentary features indicate a tide‐influenced environment. Twenty‐one ichnospecies have been identified and assigned to the Cruziana and Skolithos ichnofacies, including the oldest record of Beaconites capronus and Macaronichnus segregatis. Cruziana ichnostratigraphy (sensu lato), previously rarely used in other palaeocontinents than Gondwana, allows an age determination for these metasediments of Middle Cambrian to Lower Ordovician as indicated by the presence of Cruziana barbata, C. furcifera, C. rugosa, C. semiplicata, Didymaulichnus rouaulti and Rusophycus ramellensis. Baltica was geographically the most isolated from the other three large continents (Gondwana, Laurentia and Siberia) during Cambro‐Ordovician time, and provinciality of faunal assemblages (e.g. brachiopods, conodonts) has been proved and is also supposed for trilobites by some authors. However, although the Cruziana ichnospecies result from a high specialization of their tracemakers, and therefore only a small group of trilobite species is eligible for its origin, the ichnospecies reported from Baltica occur also on other palaeocontinents and do not support the assumption of trilobite provincialism. Copyright © 2004 John Wiley & Sons, Ltd.