Mud volcanoes and gas hydrates in the Anaximander mountains (Eastern Mediterranean Sea)

Detailed multibeam, sedimentological, and geophysical surveys provide ample new data to confirm that the Anaximander Mountains (Eastern Mediterranean) are an important area for active mud volcanism and gas hydrate formation. More than 3000 km of multibeam track length was acquired during two recent missions and 80 gravity and box cores were recovered. Morphology and backscatter data of the study area have better resolution than previous surveys, and very detailed morphology maps have been made of the known targeted mud volcanoes (Amsterdam, Kazan and Kula), especially the Amsterdam “crater” and the related mud breccia flows. Gas hydrates collected repeatedly from a large area of Amsterdam mud volcano at a sub-bottom depth of around 0.3–1.5 m resemble compacted snow and have a rather flaky form. New gas hydrate sites were found at Amsterdam mud volcano, including the mud flow sloping off to the south. Gas hydrates sampled for the first time at Kazan mud volcano are dispersed throughout the core samples deeper than 0.3 m and display a ‘rice’-like appearance. Relative chronology and AMS dating of interbedded pelagic sediments (Late Holocene hemipelagic, sapropel layer S1 and ash layers) within the mud flows indicate that successive eruptions of Kula mud volcano have a periodicity of about 5–10 kyrs. New mud volcanoes identified on the basis of multibeam backscatter intensity were sampled, documented as active and named “Athina” and “Thessaloniki”. Gas hydrates were sampled also in Thessaloniki mud volcano, the shallowest (1264 m) among all the active Mediterranean sites, at the boundary of the gas hydrate stability zone. Biostratigraphical analyses of mud breccia clasts indicated that the source of the subsurface sedimentary sequences consists of Late Cretaceous limestones, Paleocene siliciclastic rocks, Eocene biogenic limestones and Miocene mudstones. Rough estimations of the total capacity of the Anaximander mud volcanoes in methane gas are 2.56–6.40 km³.     

地中海海岭泥火山的构造特征及其油气意义

地中海海岭泥火山属于典型的活动陆缘泥火山特征,上面覆盖着富含有机质、高沉积速率的沉积物盖层。受板块构造挤压作用的影响,增生楔内断裂和盐底辟构造发育,为地层深部高压的流体和气体的向上运移提供通道。同时,渗透性差的海底碳酸盐盖层和墨西拿期蒸发盐又是理想的油气和天然气水合物捕获体,已经成为海底油气、天然气水合物评估的重要标志地貌。台湾增生楔和地中海海岭有着相似的活动陆缘地质构造特征,具有类似的成油环境及沉积物中流体、气体的迁移机制。通过对比研究,可进一步了解活动板块陆缘增生楔的油气生储意义。     

Geophysical evidence of mud diapirism on the Mediterranean Ridge accretionary complex

Mud volcanoes, mud cones, and mud ridges have been identified on the inner portion of the crestal area, and possibly on the inner escarpment, of the Mediterranean Ridge accretionary complex. Four areas containing one or more mud diapirs have been investigated through bathymetric profiling, single channel seismic reflection profiling, heat flow measurements, and coring. A sequence of events is identified in the evolution of the mud diapirs: initially the expulsion on the seafloor of gasrich mud produces a seafloor depression outlined in the seismic record by downward dip of the host sediment reflectors towards the mud conduit; subsequent eruptions of fluid mud may create a flat topped mud volcano with step-like profile; finally, the intrusion of viscous mud produces a mud cone.The origin of the diapirs is deep within the Mediterranean Ridge. Although a minimum depth of about 400 m below the seafloor has been computed from the hydrostatic balance between the diapiric sediments and the host sediments, a maximum depth, suggested by geometric considerations, ranges between 5.3 and 7 km. The presence of thermogenic gas in the diapiric sediments suggests a better constrained origin depth of at least 2.2 km.The heat flow measured within the Olimpi mud diapir field and along a transect orthogonal to the diapiric field is low, ranging between 16 ± 5 and 41 ± 6 mW m^–2. Due to the presence of gas, the thermal conductivity of the diapiric sediments is lower than that of the host hemipelagic oozes (0.6–0.9 and 1.0–1.15 W m^–1 K^–1 respectively).We consider the distribution of mud diapirs to be controlled by the presence of tectonic features such as reverse faults or thrusts (inner escarpment) that develop where the thickness of the Late Miocene evaporites appears to be minimum. An upward migration through time of the position of the décollement within the stratigraphic column from the Upper Oligocene (diapiric sediments) to the Upper Miocene (present position) is identified.     

Uca marionis Mud Volcanoes: A Unique Ichnological Tool from the Bay of Bengal Coast of India for Ready Assessment of Beach Stability

Formation of burrow-head mud volcanoes by the mud-loving amphibious crab Uca marionis (Alcock) selectively on the either sides of estuary mouths along the Bay of Bengal coast, eastern India, requires peculiar substratal and biological conditions where soft muddy bottom sediments of the coastal mudflats are covered by a thin blanket of relatively rigid beach sands that provide a false substrate stability. Shallow-seated mud layers, being within the substrate penetration power, allow opportunistic burrowing by the crab and consequent oozing out of internal mud slur to form conical heaps of mud resembling volcanic cones. Removal of this thin sand cover prompts very fast erosion of the bottom mud and consequent widening of the estuary mouth and rapid beach erosion. A thick and rigid sand cover restricts deeper burrowing by the crabs.

Uca marionis mud volcanoes, by virtue of limited power of vertical penetration down to required muddy substrate and requirement of special substrate conditions and geomorphic position along the coastal zone, are considered as unique ichnological tools that readily identify and precisely demarcate highly unstable or erosion-prone beach sectors, and thus help in planning, protection and execution of various coastal developmental programs. Wide zoogeographic distribution of Uca in tropical-subtropical sea coasts greatly enhances the application potential of this new ichnological tool. Fossilized biogenic mud volcanoes in post-Jurassic geologic records would ease taxonomic identification of the ancient trace producers and coastal marine palaeogeomorphic interpretations.     

Macrobenthic Communities of the Eastern Mediterranean Continental Shelf

Abstract. The results of a year long study in which soft-bottom epibenthic invertebrates were collected by grab, dredge and beam-trawl, along the southern Mediterranean coast of Israel, are described in this paper. The classificatory analysis used for both normal and inverse analyses used two measures of dissimilarity - Canberra metric and Bray-Curtis, and both group-average and nearest-neighbour clustering. The results were displayed as dendrograms. Four site groups and five species groups characterised a total of 58 site samples and 245 species. The prominent species in each of the five species groups are mentioned. The applicability of "community concepts" and the effects of sediment properties on community structure are discussed.     

Macrofauna Communities in the Eastern Mediterranean Deep Sea

Abstract. During two expeditions with RV ‘Meteor’ in summer 1993 and winter 1997/98 the structural and functional diversity of the benthic system of the highly oligotrophic eastern Mediterranean deep sea was investigated. The macrofauna communities were dominated by polychaetes even at the deepest stations. The fauna at shallow stations was dominated by surface deposit feeders, whereas subsurface deposit feeders and predators generally increased with depth. A high percentage of suspension‐feeding Porifera was found in the Levantine Basin. Mean abundance and number of taxa of both expeditions were significantly correlated to depth and distance to the nearest coast as well as to the total organic carbon (TOC) content in sediments. Numbers of taxa and abundance decreased generally with depth, although lowest numbers were not found at the deepest stations but in the extremely oligotrophic Levantine and Ierapetra Basin. Biomass measured during the second cruise was extremely low in the Ierapetra Basin and comparable to other extreme oligotrophic seas. The significant correlations found for TOC contents and macrofauna with distance to coast during both expeditions apparently reflect the role of hydrographically governed transport of organic matter produced in coastal regions into greater and extreme depths of the Mediterranean Sea. Seasonal differences in macrofauna communities due to seasonal differences in food supply were not found. However, recent large‐scale hydrographic changes (Eastern Mediterranean Transient, EMT) might change the oligotrophy and, thus, the structure of the benthic communities in the Eastern Mediterranean deep sea.     

The Levant Slumps and the Phoenician Structures: collapse features along the continental margin of the southeastern Mediterranean Sea

Two distinct series of slumps deform the upper part of the sedimentary sequence along the continental margin of the Levant. One series is found along the base of the continental slope, where it overlies the disrupted eastern edge of the Messinian evaporites. The second series of slumps transects the continental margin from the shelf break to the Levant Basin. It seemed that the two series were triggered by two unrelated, though contemporaneous, processes. The shore-parallel slumps were initiated by basinwards flow of the Messinian salt, that carried along the overlying Plio-Quaternary sediments. Seawater that percolated along the detachment faults dissolved the underlying salt to form distinctly disrupted structures. The slope-normal slumps are located on top of large canyons that cut into the pre-Messinian sedimentary rocks. A layer of salt is found in the canyons, and the Plio-Quaternary sediments were deposited on that layer. The slumps are bounded by large, NW-trending faults where post-Messinian faulted offset was measured. We presume that the flow of the salt in the canyons also drives the slope-normal slumps. Thus thin-skinned halokynetic processes generated the composite post-Tortonian structural patterns of the Levant margin. The Phoenician Structures are a prime example of the collapse of a distal continental margin due to the dissolution of a massive salt layer.     

Impact of the Eastern Mediterranean Transient on the distribution of anthropogenic CO2 and first estimate of acidification for the Mediterranean Sea

In the Mediterranean Sea the carbon chemistry is poorly known. However, the impact of the regional and large-scale anthropogenic pressures on this fragile environment rapidly modifies the distribution of the carbonate system key properties like C_T (total dissolved inorganic carbon), A_T (total alkalinity), C_ANT (anthropogenic CO₂), and pH. This leads inexorably to the acidification of its waters. In order to improve our knowledge, we first develop interpolation procedures to estimate C_T and A_T from oxygen, salinity, and temperature data using all available data from the EU/MEDAR/MEDATLAS II database. The acceptable levels of precision obtained for these estimates (6.11 ??mol-kg⁻¹ for C_T and 6.08 ??mol kg⁻¹ for A_T) allow us to draw the distribution of C_ANT (with an uncertainty of 6.75 ??mol kg⁻¹) using the Tracer combining Oxygen, inorganic Carbon, and total Alkalinity (TrOCA) approach. The results indicate that: 1) all Mediterranean water bodies are contaminated by anthropogenic carbon; 2) the lowest concentration of C_ANT is 37.5 ??mol kg⁻¹; and 3) the western basin is more contaminated than the Eastern basin. After reconstructing the distribution of key properties (C_T, A_T, C_ANT) for four periods of time (between 1986 and 2001) along a west-east section throughout the whole Mediterranean Sea, we analyze the impact of the Eastern Mediterranean Transient (EMT). Not only has the concentration of C_ANT increased (especially in the intermediate and the bottom layers of the eastern basin, during and after the EMT), but also the distribution of all properties has been considerably perturbed. This is discussed in detail. For the first time, the level of acidification is estimated for the Mediterranean Sea. Our results indicate that for the year 2001 all waters (even the deepest) have been acidified by values ranging from −0.14 to −0.05 pH unit since the beginning of the industrial era, which is clearly higher than elsewhere in the open ocean. Given that the pH of seawater may affect a very large number of chemical and biological processes, our results stress the necessity to develop new programs of research to understand and then predict the evolution of the carbonate system properties in the Mediterranean Sea.     

Role of the shale tectonics on the evolution of the Eastern Venezuelan Cenozoic thrust and fold belt

This paper presents a structural and stratigraphic analysis of the foreland-fold-belt of the Eastern Venezuelan Basin and the main conclusions about shale tectonic mechanisms in the area. The deformation of the foreland-fold-belt has been investigated analyzing the growth strata architecture preserved on the structure fold limbs. Three contractional episodes are proposed for the Eastern Venezuelan Basin: 1) Oligocene to middle Miocene, 2) late Miocene to Pliocene and 3) Pleistocene. The first episode produced contractional listric faults inside the shale and long displacement blind thrusts in the underlying Cretaceous units. The second episode produced the deformation of the Cenozoic strata into overlapping east-west-trending, convex northward anticlines that covers more than 200 kilometers in length and 40 kilometers wide, break-through normal faults product of a high sedimentary load that overcomes contraction and the formation of short-displacement blind thrusts in the underlying Cretaceous units. The last episode is related to an oblique compression and the formation of high angle extensional faults with dextral movement and NW-SE strike. The role of the shale tectonics in the evolution implies that shale deforms in two stages: 1) folding and 2) normal faulting of the crest of the anticline (Break through normal faulting). Folding controlled the sediment distribution during most of the Neogene strata, while the normal faulting of the anticlines represent basin potential for hydrocarbon. The best potential hydrocarbon plays in the basin are related to oblique-collision restricted basins and controlled by break-through normal faults and the presence of NW-SE strike faults that connect the HC source with the reservoirs. Results from this research imply that the role of sedimentation is fundamental for the overburden sand distribution and tectonic constrain of the folds.     

Vulnerability assessment of the impact of sea-level rise and flooding on the Moroccan coast: The case of the Mediterranean eastern zone

The eastern part of the Mediterranean coast of Morocco is physically and socio-economically vulnerable to accelerated sea-level rise, due to its low topography and its high ecological and touristic value. Assessment of the potential land loss by inundation has been based on empirical approaches using a minimum inundation level of 2 m and a maximum inundation level of 7 m, where scenarios for future sea-level rise range from 200 to 860 mm, with a ‘best estimate’ of 490 mm. The socio-economic impacts have been based on two possible alternative futures: (1) a ‘worst-case’ scenario, obtained by combining the ‘economic development first’ scenario with the maximum inundation level; and (2) a ‘best-case’ scenario, by combining the ‘sustainability first’ scenario with the minimum inundation level. Inundation analysis, based on Geographical Information Systems and a modelling approach to erosion, has identified both locations and the socioeconomic sectors that are most at risk to accelerated sea-level rise.     

Progressive deformation of an evaporite-bearing accretionary complex: SeaMARC I,SeaBeam and piston-core observations from the Mediterranean Ridge

The Mediterranean Ridge is an arcuate ridge of deformed sediment caught up in the convergent plate margin between the African plate and the Aegean. An intensive campaign of SeaMARC I and SeaBeam surveys followed by piston coring has been conducted along the contact between undeformed turbidites of the Sirte Abyssal Plain and folded and faulted sediments of the Mediterranean Ridge. Along the outer edge of the Ridge, surficial sediments have been deformed into sinusoidal ridges and troughs (wavelengths 0.5–2 km, amplitude 20–150 m), which we interpret as folds. In plan view, the ridge and the trough fabric parallels the NW-SE trending regional contours, suggesting that the folds formed in response to compression orthogonal to the Mediterranean Ridge. The outermost ridge is shedding a debris apron out onto the abyssal plain, implying that uplift and deformation are ongoing. We show that the geometry of the outermost folds can be produced by elastic bending of a packet of 5–10 relatively strong layers, each 10–20 m thick, interbedded between weaker layers; we equate the strong layers with gypsum beds in the Messinian upper evaporites. Folding the seafloor from a flat layer into the observed ridge and trough topography would shorten the layer by less than 2%. Two percent shortening (equals two percent thickening) is insufficient to create the observed relief of the Mediterranean Ridge even if the entire sediment column down to basement were involved; we infer that additional shortening/thickening is accommodated by thrust faulting above a decollement at the top of the Messinian salt layer. At distances > 15 km from the deformation front and more than 500 m from the abyssal plain, sharp-edged, fine-grained side-scan lineations with very little vertical relief cut across the kilometer-scale ridge and trough topography. These fine-grained lineations fall in two groups trending N/S to NNE/SSW and ~ENE. We interpret these lineaments as traces of conjugate strike-slip faults formed in the same compressional regime which formed the NW/SE trending folds. The onset of strike-slip faulting may coincide with the cessation of imbricate thrust fan development above the initial salt-controlled decollement surface. The following characteristics of the Mediterranean Ridge are attributed to the presence of evaporites in the incoming sedimentary section: (1) initial deformation by folding rather than thrust faulting; (2) narrow taper; (3) rapid rate of outward growth; (4) karstification.     

Seismic indicators of focused fluid flow and cross-evaporitic seepage in the Eastern Mediterranean

We present for the first time a synthesis of the evidence of focused fluid flow in the Eastern Mediterranean, providing an updated record that includes recent and past occurrences through the last ca. 6 My of evolution of the basin. We do this by adding the interpretation of a previously unpublished regional 2D seismic dataset to the existing occurrences of focused fluid flow reported in the literature. Our interpretation shows a high number (141) of focused fluid flow features, which span the stratigraphic interval from late Miocene to Recent. Of these features, (82) are at the seabed, and (59) are buried. The previous published record is more difficult to quantify, but in comparison shows an overwhelming majority of seabed features, with only rare examples of buried features.The spectrum of the buried and seabed features covers pockmarks, pipes, mud volcanoes, clastic intrusions and collapse structures. Clustering of the fluid flow features is observed at different times in specific areas, including the Nile Cone, and the Levant, Herodotus, Cyprus and Latakia basins. With the buried record, we are able to highlight the evolution of the leakage points through time. Focused fluid flow venting has been occurring since the onset of the Messinian Salinity Crisis and the start of basinwide deposition of evaporites. We focus in particular on seismic indicators of leakage through evaporites, and of sub-evaporitic source for fluids and remobilised sediments. We also discuss the role of the evaporites as a seal to ascending fluids, and in which circumstances they can be breached.Fluids (and associated remobilised sediments) are sourced from different intervals, from the sub- and supra-evaporitic section, and possibly within the evaporites. Only a minor proportion of the fluid flow features are certainly sourced from below the Messinian evaporites, and most of them are located in the Nile-Levant-Eratosthenes areas. The few examples of pathways that are able to cross thick, undeformed and well preserved evaporites are typically correlated to overpressure release and hydrofracturing. This confirms that the evaporites do act regionally as a very good seal as expected, while fluids are able to cross the evaporites only in their most extreme expression, i.e. in near-lithostathic overpressure conditions. This is confirmed by our observations made in the Eastern Mediterranean, where in the presence of relatively undisturbed evaporites, cross-evaporite vertical fluid pathways are only observed at the high end of the flux-pressure range, and involve sediment remobilisation. Maps combining these different elements can be used to detect areas potentially more prone to breaching.     

Tectonically-driven mud volcanism since the late Pliocene on the Calabrian accretionary prism,central Mediterranean Sea

Mud volcanoes recently discovered on the offshore Calabrian Arc are investigated at two sites 60 km apart, in water depths of 1650--2300 m, using swath bathymetry, 2D&3D multichannel seismic and cores. The seabed and subsurface data provide information on their formation and functioning in relation to tectonic activity during the rapid Plio-Quaternary advance of the accretionary prism. Fore-arc extension and thrust-belt compression are seen to have involved two main phases of activity, separated by a regional unconformity recording a mid-Pliocene (3.5–3.0 Ma) tectonic reorganization. The two sites of mud volcanism lie in contrasting tectonic settings (inner fore-arc basin vs central fold-and-thrust belt) and record differing forms of seabed extrusive activity (twin mud cones and a caldera vs a broad mud pie). At both sites, subsurface data show that mud volcanism took place throughout the second tectonic phase, since the late Pliocene; differing forms of mud extrusion were accompanied by subsidence to form depressions beneath and within extrusive edifices up to 1.5 km thick. The basal subsidence depressions point to sources within the succession of thrusts underlying the inner to central Arc, consistent with microfossils within cored mud breccias from both sites that are derived from strata as old as Late Cretaceous.     

Analysis of simrad EM12 multibeam bathymetry and acoustic backscatter data for seafloor mapping,exemplified at the Mid-Atlantic Ridge at 45° N

The EM12 multibeam echosounder can record acoustic backscatter information as well as high resolution bathymetry. The dataset presented, from the axis of the Mid-Atlantic Ridge at 45° N, was the first EM12 survey of a mid-ocean ridge. This paper presents methods for utilising the backscatter information. Data processing enables the production of a mosaic of acoustic backscatter, and visualisation techniques are investigated to provide initial qualitative views of the combined backscatter and bathymetry datasets. The co-registration of the backscatter and bathymetry data enables quantitative analysis of their relationships. Various sites of different geological type have been selected and their angular acoustic backscattering relationships estimated, including the effect on backscatter of incidence angle, its regional variability with bottom type and the influence of bottom slope. Incidence angles and bottom type are shown to affect backscatter to a similar degree, while slopes appear to contribute little. The geometry of hull-mounted systems, such as the EM12, is significantly different from that of conventional sidescan sonars, such as GLORIA, and the backscatter images from the two types differ in various respects. Because of the wide variations in incidence angle that are common with hull-mounted systems, and the importance of incidence angle in determining backscatter strength, it is vital to consider the effect of incidence angle during interpretation.     

Mud volcanism and fluid geochemistry in the Cheleken peninsula,western Turkmenistan

A geological and geochemical study has been carried out to investigate the relationships between major mud volcano structures and deep fluid migration in the Cheleken peninsula, in the South Caspian Basin. The fluid geochemistry allowed the origin and migration of the saline waters and the hydrocarbons to be deduced along with the regional source and reservoir rocks. The emitted waters formed by the mixing of deep highly saline water from the main source rocks of the Maykop Fm with the Caspian-like pore water contained in the Pliocene reservoirs. The water composition is very similar to that emitted by the mud volcanoes in Azerbaijan, allowing comparisons to be done between the reservoirs in the western and eastern sides of the South Caspian Basin. The associated oil is derived from a mixed type II/III kerogen deposited in a sub-oxic marine environment and generated during the early oil window. The oil biomarkers indicate that the source rock is the Maykop Fm., as previously determined for the other areas of the South Caspian Basin.The spontaneous emissions, showing different morphologies, are mainly aligned along normal and transtensive fault systems, which provide effective pathways for rapid fluid ascent from deep reservoirs to the surface.     

Midlittoral polychaete communities in the Eastern Mediterranean Sea: new information from the implementation of the Natural Geography in Shore Areas (NaGISA) protocol and comparisons at local and regional scales

The aim of this study was to attempt to fill current knowledge gaps on midlittoral Mediterranean biodiversity at local and regional scales, by using benthic polychaetes as a model taxon. Two different data sets were analysed: (i) a quantitative data set from the two Natural Geography in Shore Areas (NaGISA) study sites in Crete and (ii) a qualitative data set from multiple sites across the Mediterranean. At the local scale, the results provide evidence that (i) discrete species communities are formed in midlittoral Mediterranean habitats, which vary by geographical location and year, depending on the scale of observation; (ii) macrophyte coverage and Chl‐a are the only environmental variables associated, albeit weakly, with the above pattern; (iii) although naturally disturbed, the Cretan NaGISA sites do not seem to experience any anthropogenic stress; (iv) environmental heterogeneity and history seem to be much less important in shaping the polychaete communities than inter‐specific interactions; however, it is not possible to specify at this stage whether local or regional processes or even their interactions may shape the polychaete communities. At the regional scale, the results indicate that (i) the only factor that seems to be involved in the regional pattern is the identity of the study providing the data sets, which implies variability and bias in how research projects are carried out, from the sampling design through to data collection and analysis; (ii) the Cretan NaGISA sites may be considered as representative of the habitat in the Mediterranean, under certain conditions.     

西南印度洋中脊岩石地球化学特征及其岩浆作用研究

作为超慢速扩张脊的代表,西南印度洋中脊(SWIR)因其独一无二的地形地貌特征、洋壳结构、洋壳增生机制、岩浆和热液活动以及深部动力学过程,近30年来成为国内外研究的热点区域。基于近年来对SWIR玄武岩、辉长岩及橄榄岩的岩石学和地球化学研究成果总结,重点探讨了沿SWIR轴向（大尺度）以及单个洋脊分段（小尺度）的岩石地球化学变化特征及其影响因素,阐述了SWIR的岩浆供应及洋壳增生模式。其中,在9°～16°E斜向扩张脊,以构造作用为主的洋脊扩张模式导致了更宽的洋壳增生带和显著的地球化学异常;而在50°～51°E脊段,发育了强烈的火山活动,其成因机制包括克洛泽热点与洋中脊相互作用、微热点、古老熔融事件的残留地幔再熔融等几种观点。此外,西南印度洋中脊龙旂热液区（～49.7°E）的最新研究表明,其热液循环路径与拆离断层的发育密不可分,热液流体循环最深可达莫霍面以下6 km。因此,在今后的一段时间,应进一步加强SWIR不同空间尺度地幔源区性质、洋中脊构造与岩浆作用过程、热点-洋中脊相互作用和岩浆-热液活动与成矿等主要科学问题的研究。     

Feeding Habits of Trisopterus minutus capelanus (Gadidae) Off the Eastern Coast of Spain (Western Mediterranean)

Abstract. The stomach contents of poor cod, Trisopterus minutus capelanus (Lacepède), were taken at monthly intervals off the eastern coast of the Gulf of Valencia (Spain). A total of 1276 were analyzed to determine diet according to fish size and season. The basic food consists of crustaceans (Mysidacea and Decapoda) and teleosts. Feeding habits varied with size: decapods and fishes were more abundant in the stomachs of larger specimens. Little seasonal variation in food habits was recorded.     

Eastern and Western Poor Cod (Trisopterus minutus capelanus) Populations in the Mediterranean Sea: Evidence from Allozyme and Minisatellite Loci

Abstract. Nine allozyme and two minisatellite loci were used to investigate potential genetic differentiation among three samples of Mediterranean poor cod, Trisopterus minutus capelanus, from the Gulf of Lion, the Tuscan Archipelago and the Aegean Sea. Both types of markers showed consistent results, with F_ST values of 0.0262 and 0.0296 (P < 0.0015, after Bonferroni correction for multiple tests) for allozymes and minisatellites, respectively. Allele frequency heterogeneity tests between pairs of samples showed a clear separation between the two western Mediterranean samples (Gulf of Lion, Tuscan Archipelago) and the eastern one (Aegean Sea). The results indicate that at least two reproductively isolated populations of poor cod occur in the Mediterranean.