Distribution of crinoidal remains in the Upper Cretaceous of the U.S.S.R.

A list of all Upper Cretaceous sea-lilies known from the U.S.S.R. is given. Their stratigraphic and geographic distributions are discussed. Three paleogeographic subprovinces, the south-eastern, the northern and the south-western, can be distinguished in the Mediterranean paleobiogeographic province of the U.S.S.R. based on the development and distribution of the Upper Cretaceous crinoids.     

Ornithomimidae (Dinosauria: Theropoda) from the Bissekty Formation (Upper Cretaceous: Turonian) of Uzbekistan

The stratigraphically oldest remains of ornithomimid theropod dinosaurs are known from the Cenomanian Khodzhakul Formation and the Turonian Bissekty Formation of Uzbekistan. The ornithomimid from the Bissekty Formation is documented by more than 800 isolated bones that represent much of the skeleton. It shows at least three unambiguous synapomorphies of Ornithomimidae: length of anterior cervical centra three to five times greater than transverse width; low and rounded fibular crest of tibia; metatarsal III pinched between metatarsals II and IV; and proximal end of metatarsal III not visible in anterior view. Phylogenetic analysis, based on a dataset with 568 morphological characters and including all known ornithomimosaurian taxa, places the ornithomimid from the Bissekty Formation near the base of the ornithomimid radiation, between Archaeornithomimus asiaticus and Sinornithomimus dongi.     

Vertebrate remains from the Upper Cretaceous (Santonian) Ajka Coal Formation,western Hungary

Vertebrate remains from the Upper Cretaceous (Santonian) Ajka Coal Formation (Bakony Mountains, western Hungary) are described. Macro- and microfossils collected from two boreholes and from isolated chunks of sediment/matrix dumped on spoil heaps of the Jókai Mine represent pycnodontiform and lepisosteiform fishes, bothremydid turtles, the mosasauroid Pannoniasaurus inexpectatus, the crocodyliforms cf. Theriosuchus, Iharkutosuchus makadii and cf. Allodaposuchus, as well as ankylosaurian and theropod dinosaurs. This unit was deposited in a swampy lacustrine environment, in contrast with the neighbouring and contemporaneous floodplain deposit of the vertebrate-bearing Csehbánya Formation at Iharkút. Despite significant environmental differences, the faunal composition of the Ajka Coal Formation assemblage completely overlaps with that of the Csehbánya Formation, suggesting the occurrence of the same semi-aquatic and terrestrial species in both settings. The ankylosaurian remains further strengthen the previous view that ankylosaurus preferred wetland habitats such as fluvial systems and coastal regions.     

A new genus of Tanyderidae (Insecta: Diptera) from Myanmar amber,Upper Cretaceous

A new genus and species, Similinannotanyderus lii gen. et sp. nov., is described from Myanmar amber. It differs from other genera of Tanyderidae based on special characters of wing venation and male genitalia. This new taxon broadens the diversity of Tanyderidae in the Cretaceous, and its morphological characters enhance our understanding of the development and evolution of the primitive crane flies.     

New abelisaurid remains from the Anacleto Formation (Upper Cretaceous), Patagonia,Argentina

New theropod remains with abelisaurid affinities from the Upper Cretaceous (Anacleto Formation, lower Campanian), NW Patagonia, Argentina, are here described. The specimen (MPCN-PV 69) consists of a partial premaxilla, fragmentary vertebrae, proximal portion of both humeri, distal portion of the pubis, and an incomplete pedal ungual. Characters linking with Abelisauridae are a premaxilla with a subquadrangular body, externally ornamented, and paradental plates with a striated surface; and humerus with bulbous proximal head, conical internal tuberosity, and reduced greater tubercle. The humerus is similar to those of Carnotaurus and Aucasaurus, due to the presence of a bulbous head and a discontinuity between the head and the internal tuberosity, but also differs from both taxa in the more distal location of the greater tubercle with respect to the internal tuberosity. Aucasaurus also comes from Anacleto Formation, but differences in the humeri suggest that MPCN-PV 69 is a different taxon. The phylogenetic analysis performed supports the affiliation to Abelisauridae, but fails to determinate a more precise relationship with others abelisaurids. However, a majority rule consensus of the analysis shows a position within Brachyrostra. Despite being fragmentary, MPCN-PV 69 probably represents a new abelisaurid from the Anacleto Formation, thus increasing the knowledge and diversity of Late Cretaceous South American abelisaurids.     

Therizinosauroidea (Dinosauria: Theropoda) from the Upper Cretaceous of Uzbekistan

Skeletal remains of indeterminate therizinosauroid dinosaurs are present in the Cenomanian Khodzhakul Formation and common in the Turonian Bissekty Formation of Uzbekistan. At least two taxa are present in the Bissekty Formation based on different frontal and humerus morphotypes. Phylogenetic analysis based on a dataset with 348 morphological characters and including all known therizinosaurian taxa places the Bissekty taxa as derived non-therizinosaurid therizinosauroids. The Bissekty therizinosauroids are more derived than Alxasaurus elesitaiensis in the extensive pneumatization of the postcranial axial skeleton, the absence of teeth in the anterior portion of the dentary, the weak development or even absence of pits for the collateral ligaments on the manual phalanges, and the subtriangular medial aspect of the distal end of the humerus, with the entepicondyle positioned well medial to the ulnar condyle. They are less derived than Therizinosauridae in the presence of a basisphenoid recess, the absence of pneumatic openings on the anterior caudal vertebrae, the distal end of metacarpal III not being ginglymoid in dorsal view, and the separation of the femoral head from the neck of that bone by a raised ventral rim.     

Drill holes and shell repair in brachiopods from a Late Cretaceous (Maastrichtian) oyster reef,North Canterbury,New Zealand

Several hundred brachiopods were separated from a bulk sample taken from a single locality of the Late Cretaceous Ostrea Bed at the top of the Broken River Formation in the Weka Pass area of North Canterbury, New Zealand. The specimens were divided among five different species and examined for drill holes and shell repair following failed predatory attacks. The results show that one or more unknown predators and/or parasites apparently selectively preyed on the rhynchonellide Wekarhynchia cataracta, with approximately 16.2% of complete shells showing evidence of attack. The larger, coarsely-ribbed terebratulide Ostreathyris allani was seemingly immune to drilling, while sample sizes for the three other brachiopod species were too small to allow even rudimentary estimates of predation/parasitism rates.     

Lamniform sharks from the Cenomanian (Upper Cretaceous) of Venezuela

Sampling of Cenomanian fossil-rich horizons within the La Luna Formation of two localities in the Zulia and Trujillo states (northern Venezuela) yielded numerous shark teeth belonging to various species within the order Lamniformes (Mackerel sharks). Twelve lamniform species were identified including three new species (Squalicorax lalunaensis sp. nov., Squalicorax moodyi sp. nov., Acutalamna karsteni gen. et sp. nov.) and the genus Microcarcharias gen. nov. is proposed to accommodate with the peculiar morphology of the small-sized odontaspidid M. saskatchewanensis. Other taxa reported here include Cretoxyrhina mantelli, Cretolamna sp., cf. Nanocorax sp. and five Squalicorax species left in open nomenclature. This is the first report of chondrichthyans from the mid-Cretaceous of Venezuela and one of the few records of this group from the Cenomanian of South America. The composition of these assemblages suggests some degree of endemism in the La Luna Sea but also possible connexions with the Western Interior Seaway. One of the most striking features of these assemblages is the high anacoracid diversity (eight species) despite the corresponding outer shelf/upper slope palaeoenvironments of the La Luna Formation. The high diversity of these opportunistic predators is probably related to the high diversity of medium to large marine vertebrates that provided food resources.     

A new Plattenkalk Konservat Lagerstätte in the Upper Cretaceous of Gara Sbaa, south-eastern Morocco

Laminated, dolomitised marine limestones within a transgressive fluvio-lagoonal-carbonate platform sequence at Gara Sbaa in the Kem Kem region of south eastern Morocco yield well preserved marine fishes and crustaceans. A rarer terrestrial component includes delicate fern fronds, insects and a lizard. Sedimentological structures and stratigraphic context indicate initial shallow waters with in a carbonate lagoon with benthic microbial mats followed by deepening. High precision dating has yet to be accomplished, but a Late Cenomanian or Early Turonian age is indicated on the basis of faunal and sequence-stratigraphic considerations. The fish assemblage has affinities with mid-Cretaceous ichthyofaunas in South America and Lebanon, including taxa in common at generic level.     

Rhizocorallium in estuarine Ingersoll shale (Upper Cretaceous Eutaw Formation,Eastern Alabama coastal plain)

The ichnofossil Rhizocorallium—an oblique to horizontal U-shaped spreiten burrow—has been recognized in a broad range of Phanerozoic continental, marginal marine, and shallow to deep marine facies. Nonetheless, occurrences of this ichnotaxon in Cretaceous estuarine deposits are rarely described. Herein we document Rhizocorallium in the Ingersoll shale, a carbonaceous clay lens within an estuarine succession contained in the Upper Cretaceous (Santonian) Eutaw Formation exposed in the eastern U.S. Gulf coastal plain. The Ingersoll Rhizocorallium specimens, assigned to R. commune var. irregulare based on their basic morphology, horizontal disposition, dimensions, and actively filled spreiten, record a change in depositional conditions and substrate consistency linked to transgression. The Ingersoll shale accumulated rapidly in a restricted tidal channel in a bayhead delta setting and hosts an impoverished resident ichnofauna reflecting soupground conditions. In contrast, attributes of Rhizocorallium (e.g., burrow wall bioglyphs) and associated Thalassinoides in the upper part of the Ingersoll shale reflect firm substrates (i.e., Glossifungites ichnofacies). Firmground development and colonization reflect burial dewatering and subsequent exhumation by bay-shoreline ravinement of clays and the concomitant shift to estuarine central bay deposition. The biological affinity of the Rhizocorallium trace makers cannot be confidently established. However, based on relatively straight and seemingly current-aligned axes, passive burrow fills, lack of associated fecal pellets, and emplacement in firm clay, these structures were likely produced by suspension-feeding organisms—either crustaceans or worms—that took advantage of wave- or tide-induced currents in shallow areas near the estuary head.     

A new species of Platanus from the Cenomanian(Upper Cretaceous) in eastern Heilongjiang,China

The stratigraphic division and sequence of the Upper Cretaceous sediments in eastern Heilongjiang Province,China,have been ambiguous and controversial,mainly due to a lack of biostratigraphically useful fossils and related radiometric dating.A new species of angiospermous fossil plant.Platanus heilongjiangensis sp.nov.,from Qitaihe in eastern Heilongjiang has been found in sediments conformably above which zircons from a rhyolitic tuff has been dated by U-Pb radiometric methods as 96.2± 1.7 Ma.indicating that the Upper Houshigou Formation is of Cenomanian age.This discovery not only provides new data to improve our stratigraphic understanding of the Houshigou Formation,but also shows that Platanus flourished in the early Late Cretaceous floras of the region.This new study also indicates active volcanism taking place in the eastern Heilongjiang region during the Cenomanian of the Late Cretaceous.     

A new titanosaur (Dinosauria,Sauropoda) from the Upper Cretaceous of Lo Hueco (Cuenca,Spain)

The upper Campanian-lower Maastrichtian site of Lo Hueco (Cuenca, Spain) has provided a set of well-preserved partial skeletons in anatomical connection or with a low dispersion of their skeletal elements. One partial skeleton is herein described and a new titanosaurian sauropod is established, Lohuecotitan pandafilandi. This titanosaur is diagnosed by eight autapomorphic features: dorsally and ventrally widened or bifurcated posterior centrodiapophyseal lamina in anterior and middle dorsal vertebrae; short postspinal lamina with a transversely expanded distal end represented by smooth scars in the dorsal vertebrae; anteriormost caudals with the medial spinoprezygapophyseal and medial spinopostzygapophyseal laminae ventrally connected with the prespinal and postspinal laminae, respectively; anterior caudal neural spines with a dorsal projection of the prespinal and postspinal laminae; anterior caudal neural spines bears a “greek-cross”-like cross-section; middle caudal centra having two round and rough structures in the dorsal edge of the posterior articulation, which extends to the dorsal surface of the centrum; the articular ends of the rami of the haemal arches are divided in two articular surfaces; and tuberosity between the anterior and the lateral trochanter of the fibula. The herein performed phylogenetic analysis considered L. pandafilandi as a member of Lithostrotia more derived than Malawisaurus. The known palaeodiversity of the Late Cretaceous Ibero-Armorican titanosaurs is increasing, and further analyses focused on this group will be necessary to better understand the evolutionary history of European titanosaurs and to clarify their relationships within Titanosauria.     

A new tyrannosaurine specimen (Theropoda: Tyrannosauroidea) with insect borings from the Upper Cretaceous Honglishan Formation of Northwestern China

A large theropod ilium was recently collected from the Upper Cretaceous Honglishan Formation in the Sangequan area of the northern Junggar Basin, Xinjiang, China, which represents the first officially reported dinosaur fossil from this formation. Several morphological features, including robust supracetabular ridge, reduced supracetabular crest, concave anterior margin of the pubic peduncle, ventral flange on the pubic peduncle, converging dorsal surface of the iliac blades, laterally visible cuppedicus shelf, and ventral flange on the posterior surface of pubic peduncle, suggest that this specimen can be referred to Tyrannosaurinae, and furthermore, a few differences between this specimen and other tyrannosaurines in particular the contemporary Asian tyrannosaurine Tarbosaurus suggest that IVPP V22757 may represent a new tyrannosaurine species. However, in the absence of extensive data that would make it possible to properly evaluate these differences, we refrain from naming a new taxon based on this specimen. Some insect borings are also identified in this specimen, and are referable to the ichnogenus Cubiculum, which is interpreted as the insect pupichnia. This new fossil documents the presence of a gigantic theropod in the Upper Cretaceous of Junggar Basin, adding new information on its poorly studied ecosystems.     

Troodontidae (Dinosauria: Theropoda) from the Upper Cretaceous of Uzbekistan

To date three taxa of troodontid theropod dinosaurs have been recognized from Upper Cretaceous strata in two regions of the Kyzylkum Desert in Uzbekistan. The Cenomanian Khodzhakul Formation in the southwestern Kyzylkum Desert has yielded isolated serrated teeth and some postcranial bones of an indeterminate troodontid. In the central Kyzylkum Desert troodontids are known from the Cenomanian Dzharakuduk Formation (Urbacodon itemirensis) and the Turonian Bissekty Formation (Urbacodon sp.). Urbacodon itemirensis is known from a single dentary whereas Urbacodon sp. is represented by isolated teeth, maxilla and dentary fragments, a partial braincase, and some postcranial bones. The troodontid affinities of Urbacodon are supported by several synapomorphies: presence of a subotic recess; reduced basal tubera placed directly under the occipital condyle; maxilla participating in the margin of the external naris; nutrient foramina on dentary situated within a deep lateral groove; dentary without distinct interdental plates; large number of small dentary and maxillary teeth; teeth constricted between root and crown; anterior dentary teeth smaller, more numerous, more closely spaced than those in the middle of the tooth row, and implanted in a groove; posterior dorsal vertebrae with tall and posterodorsally tapering neural spines; and presence of a midline sulcus on the neural arches of distal caudals. Among Troodontidae, Urbacodon resembles Byronosaurus, Gobivenator, and Xixiasaurus in the absence of serrations on the tooth crowns and having premaxillary teeth that are D-shaped in cross-section. However, phylogenetic analysis did not recover a clade of Asiatic troodontids with unserrated teeth.     

A diversified vertebrate ichnite fauna from the Feitianshan Formation (Lower Cretaceous) of southwestern Sichuan,China

The Bajiu tracksite exposed on a fine sandstone surface at the top of the Feitianshan Formation, close the contact with the overlying Xiaoba Formation, reveals a saurischian dominated ichnofauna consisting of four sauropod and at least eight theropod trackways. The sauropod trackways display a medium-wide gauge pattern characteristic of titanosauriforms, while the theropod trackways can be subdivided into two distinct morphotypes. One is similar to the ichnogenera Eubrontes and Megalosauripus, the other (cf. Dromaeopodus) appears to represent a functionally didactyl dromaeosaur where pedal digit II is represented only by an oval basal pad. A single swim track is possibly attributable to a turtle or crocodylian. Such sauropod-theropod dominated ichnofaunas are consistently typical of the red-bed tracksites in the region and useful for characterizing a fauna that is otherwise poorly known from body fossils. The stratigraphic position and late occurrence of the (typical Jurassic) Eubrontes-Megalosauripus morphotype is well known from other Cretaceous localities in China and seems to reflect a peculiarity in the theropod faunas of East Asia.     

Hydrocarbon source rock variability within the Austin Chalk and Eagle Ford Shale (Upper Cretaceous), East Texas, U.S.A.

The Austin Chalk and Eagle Ford Shale are Upper Cretaceous deposits that extend across Texas from the northeast to southwest. These formations contain organic carbon enriched mudstones and chalks that were deposited during transgressions of the Cretaceous epeiric sea in North America. Recent workers in petroleum geochemistry have demonstrated that these organic enriched rocks possessed attributes common to oil source rocks. The present study of these Austin Chalk and Eagle Ford Shale rocks is from the perspective of organic petrology, and it augments the earlier geochemical work that documented source variability within units of these formations. As with the earlier work, the results of this study show that both formations contain intervals that are, when mature, capable of generating commercial quantities of liquid hydrocarbons. However, this work further revealed that Eagle Ford rocks not only exhibit greater or ganic carbon contents, but also have greater quantities of oil-prone kerogen (fluorescent amorphinite and exinite) when compared with rocks from the Austin Chalk. These source rock differences relate to levels or degrees of organic preservation. Dysaerobic to oxic depositional settings seem to be more characteristic of the Austin Chalk than of the Eagle Ford Shale. Such oxic environments do not consistently favor the preservation of organic matter. Usually, well-preserved kerogen forms under more anoxic conditions, such as those that occurred during deposition of some Eagle Ford units. These anoxic conditions suggest that the geographically more extensive Eagle Ford Shale is a more important source for oil than is the Austin Chalk.     

A possible azhdarchid pterosaur from the Crato Formation (Early Cretaceous,Aptian) of northeast Brazil

The partially articulated, distal portion of a left wing finger of a pterosaur from the Crato Formation of northeast Brazil has a T-shaped cross-section to the second and third phalanges. This cross-sectional shape is one of several characters diagnostic of the pterodactyloid pterosaur family Azhdarchidae (Unwin & Lu 1997). Until now, this family of pterosaurs was known exclusively from the Late Cretaceous. The specimen described here may be the first recorded azhdarchid from the southern hemisphere and the earliest recorded member of the family.     

Structural geology of the Upper Cretaceous Chalk Central Mass, Isle of Wight, U.K.

Remapping the Chalk of the Central Chalk Mass of the Isle of Wight between Carisbrooke (Newport), Calbourne and Shalcombe, including the Bowcombe Valley, has identified a complex series of tectonic ‘rolls’ and ‘flats’ in a region that has been interpreted to be a relay ramp between the Needles and the Sandown faults. A major new WNW trending fault at Cheverton throws the Chalk down by >50 m to the SW in a 80-100 m wide zone of faulting within which some chalk blocks have near vertical dips. The fault location and trend closely follows the edge of the Cranbourne-Fordingbridge High and could also reflect, for the first time, the surface expression of part of the Needles Fault, a major inversion reverse fault. Located along this fault zone deep Quaternary weathering of the Chalk and Quaternary gravel deposits are present. The trend of the Cheverton Fault brings it towards Gotten Leaze where a groundwater pumping station is located and groundwater springs regularly cause flooding on the Brighstone-Calbourne Road. Analyses of the jointing in the Chalk show that stratabound fracture patterns typical of the Chalk formations elsewhere in Southern England are present in the Central Mass. In addition, there are numerous small faults along which valleys have formed. Tectonic structure and lithology have had a profound influence on the geomorphology and groundwater flow in the Chalk in the Central Mass.     

New species Dalembia jiayinensis (Magnoliopsida) from the Upper Cretaceous Yong'ancun Formation,Heilongjiang, northern China

Leaves of Dalembia jiayinensis sp. nov. are reported from the Santonian Yong'ancun Formation exposed along the Heilongjiang (Amur) River in Jiayin County, Northeast China. The leaves are compound, odd-pinnate, consisting of five leaflets. Leaflets are pinnately-lobed, trilobate or unlobed, elliptic, oblong, elliptic-rhomboid or ovate in outline, with blunt or rounded apex and the leaflet base is variable being cuneate, slightly decurrent, truncate or slightly cordate and sometimes asymmetric. Margin is entire or lobed. Lobes are short, simple, decreasing upwards, with wide and rounded sinuses. Apices of the lobes are usually rounded, sometimes notched. Venation is pinnate or palmately-pinnate, craspedodromous or semi-craspedodromous. The new species D. jiayinensis extends both the geographical and stratigraphic ranges of the genus Dalembia. This is the first occurrence of Dalembia in China and the most southern and youngest (Santonian) occurrence of this genus.     

The cranial morphology and taxonomic status of Tarchia (Dinosauria: Ankylosauridae) from the Upper Cretaceous of Mongolia

The taxonomic status of specimens referred to Tarchia kielanae, T. gigantea, and Minotaurasaurus ramachandrani has been contested. The holotype of T. kielanae, Zaklad Paleobiologii (Institute of Paleobiology)-Polish Academy of Sciences (ZPAL) MgD I/111, is redescribed. It has features common to both Palaeontological Institute-Russian Academy of Sciences (PIN) 3142/250, long considered Tarchia, and to M. ramachandrani but which are lacking in Saichania chulsanensis. The specimen PIN 3142/250 is not referable to Saichania but instead represents a new species, Tarchia teresae sp. nov. Similarly, the holotype skull of M. ramachandrani is not referable to T. kielanae, so Minotaurasaurus is a valid taxon. Tarchia is more derived than either Saichania or Minotaurasaurus while sharing features of both.