Biodiversity and conservation in the Tibetan Plateau

The Tibetan Plateau (Qinghai-Xizang Plateau) is a unique biogeographic region in the world, where various landscapes, altitudinal belts, alpine ecosystems, and endangered and endemic species have been developed. A total of 26 altitudinal belts, 28 spectra of altitudinal belts, 12,000 species of vascular plant, 5,000 species of epiphytes, 210 species of mammals, and 532 species of birds have been recorded. The plateau is also one of the centers of species formation and differentiation in the world To protect the biodiversity of the plateau, about 80 nature reserves have been designated, of which 45 are national or provincial, covering about 22% of the plateau area. Most of the nature reserves are distributed in the southeastern plateau. Recently, the Chinese government has initiated the “Natural Forests Protection Project of China,” mainly in the upper reaches of the Yangtze and Yellow rivers. “No logging” policies have been made and implemented for these areas.     

长江上游山地垂直带谱及其空间分布模式

长江上游包括青藏高原东南部、秦巴山地、四川盆地与云贵高原部分地区,在地理、地貌、气候、生物多样性方面都表现得极为复杂和丰富多彩,在世界山地中也占有举足轻重的位置。特别是复杂多样的山地垂直带谱更是欧亚大陆乃至世界山地垂直带研究中至关重要的组成部分。在地学信息图谱和数字山地垂直带体系的基础上,本文系统地收集和分析了长江上游共50个山地垂直带谱所体现的空间规律,河源区、横断山区、秦巴山区及贵州高原的垂直带谱类型多样并各具特色,且在经度和纬度方向又具有统一的分布规律,如雪线、林线、针叶林及阔叶林等的分布界线变化规律比较符合二次曲线规律,验证了大陆尺度上山地垂直带二次曲线模式假说。另外,山地垂直带分布规律又具有尺度效应,中小尺度上地形的影响作用表现得极为显著。     

青藏高原植被垂直带与气候因子的空间关系

集成了青藏高原气候区149个山地植被垂直带数据,利用国家基本气象台站自建站以来到2001年的地面观测日气象数据,计算了地面的温暖(WI)、寒冷(CI)、湿润(MI)、吉良龙夫(Kira)干湿指数、干燥度(Idm)等水热指数,运用GIS的空间分析模块,模拟了青藏高原水热条件的空间分布形势,探讨山地植被垂直带谱分布规律与制约因子的定量指标.结果表明:在高原的东北部、西北边缘,以荒漠、荒漠草原、山地森林、山地草原、灌丛、草甸为组合的半干旱、干旱结构向高原腹地以高寒草原、高山草甸、荒漠带组合的高寒干旱带谱结构的变化;东南、南部边缘,以温暖湿润为特征的以森林带为优势带谱组合结构逐渐向寒冷的高原中心变化;高原的地势效应,致使的水热形势旱现从中央向边缘变化的趋势是致使青藏高原植被垂直带谱分布的重要原因.     

Implications of mass elevation effect for the altitudinal patterns of global ecology

The varied altitudinal gradient of climate and vegetation is further complicated by mass elevation effect (MEE), especially in high and extensive mountain regions. However, this effect and its implications for mountain altitudinal belts have not been well studied until recently. This paper provides an overview of the research carried out in the past 5 years. MEE is virtually the heating effect of mountain massifs and can be defined as the temperature difference on a given elevation between inside and outside of a mountain mass. It can be digitally modelled with three factors of intra-mountain base elevation (MBE), latitude and hygrometric continentality; MBE usually acts as the primary factor for the magnitude of MEE and, to a great extent, could represent MEE. MEE leads to higher treelines in the interior than in the outside of mountain masses. It makes montane forests to grow at 4800–4900 m and snowlines to develop at about 6000 m in the southern Tibetan Plateau and the central Andes, and large areas of forests to live above 3500 m in a lot of high mountains of the world. The altitudinal distribution of global treelines can be modelled with high precision when taking into account MEE and the result shows that MEE contributes the most to treeline distribution pattern. Without MEE, forests could only develop upmost to about 3500 m above sea level and the world ecological pattern would be much simpler. The quantification of MEE should be further improved with higher resolution data and its global implications are to be further revealed.     

青藏高原维管植物物种丰富度分布的情景模拟

如何充分利用离散的观测数据,通过对维管植物物种分布丰富度及其与生境因子之间的相互作用和影响机理的定量分析,实现维管植物物种丰富度的空间分布及其情景模拟,是目前生物多样性研究前沿和核心内容之一。针对这一问题,在实现青藏高原37个国家自然保护区的维管植物物种数量收集和边界数据矢量化的基础上,分别进行维管植物物种数量与土地覆盖类型、环境因子和景观生态指数等三大类生境因子之间的相关关系的定量计算和对比分析,筛选和确定最佳相关分析方程,进而构建青藏高原维管植物物种丰富度的空间模拟分析模型。该模型中,维管植物物种丰富度与生境因子之间的复相关系数为0.94,模型验证结果表明,青藏高原的维管植物物种的平均丰富度为496.79种/100 km²,其空间分布格局整体上呈东南向西北逐渐减少趋势;另外,除柴达木盆地荒漠区域以外,维管植物物种的空间分布随海拔的升高而减少。基于CMIP5 RCP 2.6、RCP 4.5和RCP 8.5三种气候情景模拟获得的青藏高原维管植物物种丰富度未来情景结果显示,在T0-T4（2010-2100）时段内,青藏高原维管植物物种丰富度整体将呈减少趋势。RCP 8.5情景下青藏高原维管植物物种丰富度的变化幅度最大,而RCP 2.6情景下的维管植物物种丰富度的变化幅度最小。研究表明,本文构建的模型能够对青藏高原维管植物物种丰富度的空间分布格局及其未来情景进行模拟分析,模拟结果可为青藏高原生物多样性及其对气候变化响应的综合评估和情景模拟提供方法和技术支持。     

秦岭中部山地落叶阔叶林超级垂直带的发现与意义

山地垂直带谱是气候和植被水平地带变化和更替的缩影,垂直带的带幅、带间过渡方式、带内结构和垂直带组合方式都表现出高度的异质性和复杂性。本文发现在中国南北过渡带中部太白山发育了世界上最宽的山地垂直带——山地落叶阔叶林垂直带。该垂直带从基带到典型垂直带再到先锋性垂直带皆为山地落叶阔叶林,3种本来可以独立存在的垂直带,连续分布形成了包含3个栎林亚带、2个桦林亚带的“三层五亚带”超级垂直带,远远超过正常情况下山地垂直带1000 m的阈值,且其上限达到了海拔2800 m。它的形成与秦岭所处的过渡性地理位置、秦岭中部垂直带谱的完整性、丰富的落叶木本植物种群及其形成的强大群落竞争优势等因素紧密相关。超级垂直带的发现有多方面的意义：它是中国南北过渡带又一重要的标志性自然地理特征;它表明山地垂直带在特殊的山地环境中可以具有非常复杂的内部结构和宽大带幅,这扩展了我们对山地垂直带谱结构及机理认识的广度,对于创建山地垂直带谱结构理论具有十分重要的意义;超级垂直带的发现,也说明中国南北过渡带还有很多科学内容有待我们去探索和发现,希望本文能起到抛砖引玉的作用,引起学界对超级垂直带形成的气候和生物多样性因素、地理过渡带的结构和生态效应等重大问题进行深入研究。     

天山数字垂直带谱体系与研究

对6个全国性山地垂直带谱体系进行评述。认为区域性的数字垂直带谱的详细研究也是建立中国山地垂直带信息图谱的重要一环。只有通过这样的工作，才能进一步发现问题和解决问题，才能逐步趋于完成山地垂直带谱集大成的工作。分析了天山垂直带谱形成的因素，建立了包括北坡、南坡、西部伊犁谷地、天山腹地(巴音布鲁克盆地)的天山数字垂直带谱体系，分析了各垂直带的特点，总结了垂直带谱的区域分异规律。     

珠穆朗玛峰国家级自然保护区湿地分布研究

利用2000～2009年覆盖珠穆朗玛峰国家级自然保护区的MOD09A1和MOD13Q1数据集的相关数据,该保护区1：5万DEM,并结合2008年9月采集的野外湿地数据,进行了基于增强型植被指数（EVI）、改进型调整植被指数（MSAVI）、地表水分指数（LSWI）、海拔和坡度5个参数的保护区湿地分布特征研究。结果表明,珠穆朗玛峰国家级自然保护区的湿地类型主要为河流、湖泊、沼泽和沼泽化草甸;湿地主要分布在海拔4100～4700m且坡度≤4°的佩枯错、浪强错、多布扎等湖泊密集的高原湖盆地区、佩枯错与浪强错之间的平原地带和保护区中部及北部坡度较缓的河谷地区,大部分湿地分布在保护区的实验区内;2000～2008年保护区湿地以季相变化为主,6～9月（湿季）湿地面积约占保护区总面积的11．2％;6～9月湿地面积年际变化幅度小于保护区总面积的0．1％;坡度和海拔是影响保护区湿地空间分布的关键因素和提取该地区湿地空间分布范围的有效参数。     

"Greatest lake period"|and its palaeo-environment on the Tibetan Plateau

The “greatest lake period” means that the lakes are in the stage of their maximum areas. As the paleo lake shorelines are widely distributed in the lake basins on the Tibetan Plateau, the lake areas during the “greatest lake period” may be inferred by the last highest lake shorelines. They are several, even tens times larger than that at present. According to the analyses of tens of lakes on the Plateau, most dating data fell into the range of 40-25 ka BP, some lasted to 20 ka BP. It was corresponded to the stage 3 of marine isotope and interstitial of last glaciation. The occurrence of maximum areas of lakes marked the very humid period on the Plateau and was also related to the stronger summer monsoon during that period.     

青藏高原沼泽合理利用与生态环境保育

青藏高原有沼泽约491.3万hm2,是世界上最高的湿地。通过概要介绍高原沼泽湿地基本类型及其分布,重点阐述沼泽湿地利用、环境变化现状与存在的问题,为了保护高原生态环境,并持续利用湿地资源,提出了初步的看法和建议。     

中国山地垂直带的数字集成与基本规律分析

首次归纳出我国63类垂直带(包括31类基带)；在ArcGIS 8．3平台上,以山地垂直带谱“数字引擎”为内核,成功研制了“中国山地垂直带谱信息系统(1．0)”,将我国约239个山地垂直带谱进行了数字集成,使我们对我国垂直带谱数据的掌握从残缺上升为相对完备。该研究工作不仅为我国山地垂直带谱规律的数字分析提供了强大的手段,也为世界山地垂直带谱的数字集成提供了必要的理论和技术方法。还利用该系统,对重要垂直带界线如雪线、林线、高山草甸进行了空间规律分析。结果表明,除高山草甸经向变化缺乏明显规律外,其他垂直带界线的经、纬向变化的二次曲线(抛物线)模式比其他模式具有更高的相关系数。因而,抛物线模式应是我国山地垂直带宏观尺度上的基本空间模式。今后垂直带谱研究应在界线精度、带谱结构的地理学／生态学解释等方面继续深入。     

太白山土壤种子库储量与物种多样性的垂直格局

通过野外植被调查和室内试验,从1 220份土样中,挑选土壤中的种子,进行分类统计,研究秦岭太白山南坡土壤种子库储量与物种多样性沿海拔梯度的变化。研究结果如下：(1) 太白山南坡土壤种子库中的植物种为172种;从总体上看,土壤种子库中的物种丰富度 (S) 随着海拔的上升呈下降趋势。土壤种子库生物多样性特征表现为在2个海拔段发生较明显的变化：一是从低海拔到中等海拔 (1 500~2 500 m),二是从中等海拔到高海拔 (2 500~3 500 m)。在?琢多样性方面,生态优势度在海拔2 500 m处最大;在海拔2 400 m以下 (包含2 400 m) 的样地中,种子库生态优势度大于海拔2 500 m以上 (含2 500 m) 的样地种子库的生态优势度。 H′(Shannon-Wiener指数)与生态优势度的变化趋势相反。(2) 太白山南坡土壤种子库储量最大值出现在海拔2 600 m的样地,为2.24×104 Ind./m²;种子库储量最小值出现在海拔3 500 m的样地,为4.43×102 Ind./m²。储量沿海拔梯度的变化趋势表现为：在海拔2 600 m以下,种子库储量沿海拔的上升呈逐渐增加的趋势;在海拔2 600 m以上,种子库储量沿海拔的上升呈逐渐下降的趋势。(3) 土壤种子库的种子储量与种子密度的变化规律基本一致。而用单位面积土壤中的种子储量来表示种子库的大小特征比用种子密度来表示更为实用和方便,而且采集具有不同性质的土壤研究种子库,考虑到了土壤性质等因素的影响,更能客观反映土壤种子库的特点。     

Tibetan Plateau: Geodynamics and Environmental Evolution--The cooperative projects based upon the memorandum of CAS and DFG

The Tibetan Plateau (TP) plays a unique role in Earth System Sciences. It represents a key area to understand not only basic geodynamic processes linked with the formation and uplift of mountains and plateaus, but also the interaction between plateau uplift and environmental changes. Over the last 50 million years the formation of the TP has considerably influenced the global climate and monsoon system. Moreover, the TP proves to be extremely sensitive to present-day global change phenomena. Based upon the foundation of the new Institute of Tibetan Plateau Research (ITP) by the Chinese Academy of Sciences (CAS) and through the Memorandum signed by the CAS and DFG (Deutschen Forschungsgemeinschaft), both CAS and DFG provide opportunities to intensify TP research and to develop coordinated research programs. “The Tibetan Plateau – Geodynamics and Environmental Evolution” consisting of one big projects funded by CAS and five projects funded by DFG that cover the pre- and early-collision history of the TP, the Palaeogene/Neogene uplift and climatic dynamics as well as the Late Quaternary and recent environmental and climatic changes on the TP. The projects are linked through several levels of interactions.     

山体基面高度对欧亚大陆东南部林线分布的影响——山体效应定量化研究

根据收集到173个林线数据,采用纬度、经度和基面高度的三元一次方程拟合欧亚大陆东南部林线分布,计算各自的标准回归系数和贡献率,以此来确定山体基面高度(山体效应的简明表达形式)对林线分布高度的影响。结果表明,纬度、经度和山体基面高度对林线分布高度的贡献率分别为30.60%、26.53%、42.87%。以北纬32o为界线,对其以北、以南区域也分别进行了分析,基面高度的贡献率达到24.10%和39.11%。分析不同尺度和区域山体基面高度作用于林线的贡献率不难发现:在欧亚大陆东南部以基面高度代表的山体效应对于林线高度的影响显著,明显地超过了纬度和经度。基面高度的作用受气候条件和海陆位置影响较小,不论大陆内部或沿海,基面高度分异对山地垂直带分异的影响都相对独立和稳定。该结果定量地表明了山体效应对林线分布高度的重要作用。     

青藏高原东南部地貌边界与金沙江水系发育

4000米山顶面高程等值线,可定为高原东南部与外围地区间的地貌界线。低于山顶面的高原面的形成与上新世一早更新世古谷地、盆地的加积相关。元谋运动以来,地貌边界带的发展与高原整体的隆升同步,1500米大河河床高程等值线环绕高原分布,现代水系相继发育。金沙江与长江贯通后东流入海,这是中国大陆地势最终转变为西高东低的标志。     

青藏高原自然地理研究的进展

本文综述青藏高原自然地理研究所取得的主要进展,诸如晚新生代以来高原剧烈抬升引起的自然环境巨大变化,上新世的古地理环境和高原隆起,湖泊和水系的演变,第四纪冰川作用,全新世古地理环境演化以及高原隆起对自然环境和过程的影响等。指出了青藏高原具有的独特自然环境类型和特征,阐述了山地垂直自然带谱类型的比较研究,三维空间的地域分异,自然区划以及对独特地生态现象和区域的研究成果。     

黄土高原典型塬区土壤热状况研究

在陆-气相互作用中,土壤热状况（土壤温度、土壤导热率等）和土壤湿度等陆面状况对大气环流和气候变化都有着重要影响。黄土高原横跨干旱、半干旱及半湿润地区,为我国第二大高原,幅员辽阔。该复杂下垫面上的陆-气相互作用不仅直接影响到黄土高原地区的气候和环境变化,而且对东亚、乃至全球的气候和环境变化都可能产生重要影响。而对黄土高原区域的土壤热状况及土壤温度的研究是黄土高原陆-气相互作用研究的重要组成部分。分析了黄土高原典型塬区不同下垫面的土壤温度状况,分析了造成各种下垫面温度分布和变化不同的原因,得到如下结论：在近地层,随着土壤深度的增加,土壤温度振幅逐渐减小,40 cm土壤温度相对以上各层变化不明显。就季节变化而言,土壤温度在1 a中有两次稳定状态。第一次出现在4月上旬,其值约为6 ℃左右;第二次出现在11月中旬,温度值为14 ℃。相对于全年土壤温度而言,在12月到次年2月有一个低温中心,温度低于零度;7～8月间有一个暖中心。各层土壤温度在1月份是最低的,其后一路上升,4、5月份是土壤温度快速上升期,至8月上旬土壤温度达到最大值,为土壤升温期;其后温度开始下降。土壤温度梯度具有明显的日变化特征,夜间,土壤的热量是从深层传向地表的,而随着太阳高度角的加大,土壤温度梯度转为负值,深层土壤从地表获得能量,到了傍晚19时左右,温度梯度又转为正值;土壤温度梯度的变幅在有植被时要明显小于无植被时。各站的日平均土壤导热率,柴寺、塬下和中心站分别是1.43,1.24,1.17 W·m^-1·k^-1,土壤物理性质和土壤质地的不同是各站土壤温度分布和土壤热传导率存在差异的原因之一。     

Human’s digital footprints on the Qinghai-Tibet Plateau: Variations during festivals and impacts on nature reserves

Mobile internet and wireless communication technologies have produced unprecedented location-aware data. Such big geospatial data can be used as a proxy measure of the ‘digital footprints' left by us on the planet and provide a valuable opportunity to understand the dynamic and short-term human disturbance on the nature at fine scales. This study investigated the spatiotemporal variations of human's digital footprints on the Qinghai-Tibet Plateau using smartphone-users-generated Tencent's location request data. The results showed that human's digital footprints cover less than 5% of Qinghai and Tibet, exhibiting either a U-shaped or an N-shaped temporal change pattern during the major festivals. Spatial changes of the digital footprints manifested a transition process from dispersion to concentration in Xining and Lhasa. Human disturbance assessment of seven large nature reserves on the plateau showed that the Qinghai Lake is the most disturbed one as shown by 14.6% of its area is stained with human digital footprints and the areal average of footprint intensity is 1.59, and the disturbance was significantly escalated during the National Day holiday. By contrast, the Qangtang and Hoh Xil are the least affected nature reserves with the two indices less than 1% and 0.1, respectively.     

横断山区垂直带谱的分布模式与坡向效应

根据收集到的横断山区山地垂直带谱数据,对山地垂直带的坡向效应和空间分布规律进行了分析和研究.结果表明:1)主要的垂直带和垂直带界线如林线、暗针叶林带、雪线等的纬度和经度地带性分布规律明显并且分布模式都相似,纬向上呈开口向下的二次曲线分布模式,经向上呈开口向上的二次曲线分布模式,两者共同形成"双曲抛物面"分布模式,充分反映了横断山区的环境与生态的复杂性和独特性,也进一步丰富和发展了山地垂直带谱的二次曲线假说; 2)横断山区山地垂直带谱表现出明显的基于水分驱动的坡向效应,主要表现为同一山体的东、西坡往往具有不同的基带和带谱结构,相同类型的带谱出现的海拔和分布范围不同,迎风坡表现出较为湿润的类型和带谱结构,而背风坡则表现出更为干旱的类型和组成结构;横断山区的坡向效应主要是由于山体对当地盛行季风的影响,造成迎风坡和背风坡水热条件相差很大,从而发育不同的山地垂直带谱类型.从横断山区山地垂直带谱的空间分布规律来看, 28°～29°N、98°～101°E范围内,即大致在澜沧江以东-雅砻江以西,山地垂直带谱普遍表现出干热的特点,为横断山区干热气候的核心地带.但如何定量分析山地的坡向效应尚有待于进一步的研究和讨论.此外、数据质量和数据误差也对分析的结果,尤其是空间分布模式的数学模拟结果产生一定的影响,在以后的研究中尚需进一步完善.     

基于NPP数据和样区对比法的青藏高原自然保护区保护成效分析（英文）

Twenty-one typical coupled large samples were chosen from areas within and surrounding nature reserves on the Tibetan Plateau using the large sample comparison method(LSCM).To evaluate the effectiveness of the nature reserves in protecting the ecological environment,the alpine grassland net primary production(NPP) of these coupled samples were compared and the differences between them before and after their establishment as protected areas were analyzed.The results showed that:(1) With respect to the alpine grassland NPP,the ecological and environmental conditions of most nature reserves were more fragile than those of the surrounding areas and also lower than the average values for the Tibetan Plateau.(2) Of the 11 typical nature reserves selected,the positive trend in the NPP for Manzetang was the most significant,whereas there was no obvious trend in Taxkorgan.With the exception of Selincuo,the annual NPP growth rate in the nature reserves covered by alpine meadow and wetland was higher than that in nature reserves consisting of alpine steppe and alpine desert.(3) There were notable findings in 21 typical coupled samples:(a) After the establishment of the nature reserves,the annual rate of increase in the NPP in 76% of samples inside nature reserves and 82% of samples inside national nature reserves was higher than that of the corresponding samples outside nature reserves.(b) The effectiveness of ecological protection of the Mid-Kunlun,Changshagongma,Zoige and Selincuo(Selin Co) nature reserves was significant; the effectiveness of protection was relatively sig-nificant in most parts of the Sanjiangyuan and Qiangtang nature reserves,whereas in south-east Manzetang and north Taxkorgan the protection effectiveness was not obvious.(c) The ecological protection effectiveness was significant in nature reserves consisting of alpine meadow,but was weak in nature reserves covered by alpine steppe.This study also shows that the advantage of large sample comparison method in evaluating regional ecology change.Careful design of the samples used,to ensure comparability between the samples,is crucial to the success of this LSCM.