古尔班通古特沙漠植被与环境关系分析

The Gurbantunggut Desert is the largest stable and semi-stable sand desert in China, yet few data exist on vegetation pattern and species-environment relationships for these diverse desert landscapes. The sand dunes of the survey area are mainly of the longi- tudinal form from north to south, but checkerboard-shaped and honeycomb-shaped forms are also present, with the height of 10-50 m. We measured vegetation and soil data on north-south transects and compared them with vegetation and soil data on east-west transects. Analysis revealed that the varying trend and strength of the species diversity, dominance and cover of the plant communities in the longitudinal and transverse directions across the landscape are significantly different. The results of CCA ordination show that the microhabitats of soil organic matter （OM）, soil salts （TS）, sorting index （ σ ）, topsoil water-content （SM） and pH relate to the differences in vegetation observed as differences in species assemblage from salt-intolerant ephemerals, micro-subshrubs and subshrubs to salt-tolerant shrubs and micro-arbors. The terrain （alt.） and soil texture （the contents of Mz and Ф 1） affect the spatial differentiation of many species. However, this spatial differentiation is not so marked on transects running longitudinally with the landscape, in the same direction as the dunes. The species of the desert vegetation have formed three assemblages under the action of habitat gradients, relating to three sections running transversely across the landscape, at right angles to the direction of the dune crests. In the mid-east section of the study area the topography is higher, with sand-lands or dune-slopes with coarse particles. Here the dominant vegetation comprises shrubs and subshrubs of Seriphidium santofium and Ephedra distachya, with large numbers of ephemeral and ephemeroid plants of Senecio subdentatus, and Carex physodes in spring and summer. On the soil of the dune-slopes in the mid-west of the study area, with coarse partic     

古尔班通古特沙漠生境对植物群落格局的影响

沿87^o37'09"~88^o24'04"E纵穿古尔班通古特沙漠,对44^o14'04"~45^o41'52"N的范围进行11个样地的地貌景观调查、植被分布参数采集和土壤取样。研究区统计到的植物种数占古尔班通古特沙漠文献资料中植物种数的1/3,出现的植物群落构建了该沙漠大面积成片分布的荒漠植被景观。调查发现,随着宏观生境的空间变化,植物群落作相应的演替。在此基础上,利用因子分析 (PCA)和相关分析 (CA) 查明了沙漠风沙土理化属性,如土壤养分、盐分、水分、碱性 (pH) 等微生境的异质性仅对沙漠植被草本层片的多样性 (PIE草) 产生显著影响。同时,这些微生境因相互拮抗使沙漠环境处于暂时的脆弱平衡之中。     

古尔班通古特沙漠纵向沙垄植被空间异质性

在古尔班通古特沙漠南部纵向沙垄上按5 m×10 m的网格测度植被样方100个,采集浅层风沙土样品184件。调查发现,乔灌木层片和草本层片的分布及相互关系明显受沙垄地形影响,并且在干旱环境下种群间的竞争和空间上的互补使两层片生态优势度具有显著的相关性。地统计分析结果显示,纵向沙垄上植被空间异质性表现为木本层片的物种多样性呈斑块状分布,草本层片物种多样性呈现平行沙垄的带状结构和斑块结构叠加的特点,而植被盖度则在垂直沙垄走向的方向上具有强烈的梯度性变化。和沙垄土壤理化性质空间格局特征相对比,植被盖度及物种多样性与风沙土浅层含水量在自相关空间（变程）及空间异质性程度上比较接近。这是纵向沙垄小生态系统中草本植物和浅层土壤水分耦合关系在空间上的反映,也是木本植物对水分和土壤资源在小尺度（m）上竞争及中小尺度（10～100 m）上互补的结果。     

春夏季放牧对古尔班通古特沙漠南部土壤种子库和地上植被的影响

位于新疆北部准噶尔盆地腹地的古尔班通古特沙漠,是世界上温带荒漠生物多样性的精华和最为丰富的典型区域。历史上这里就是北疆羊和部分马匹重要的冬牧场,但近年来的放牧活动在春夏季也开始进入沙漠腹地,使地表性质与植被覆盖发生了明显改变。应用野外随机取样与室内萌发实验相结合的方法,对比研究了古尔班通古特沙漠南部春夏季放牧和冬季放牧沙垄土壤种子库差异及其与地上植被的关系。结果表明：冬季放牧沙垄种子库种数为24种,而春夏季放牧沙垄有20种,后者明显减少了植物种数。冬季放牧沙垄种子库密度为(353.2±87)粒·m-2,而春夏季放牧沙垄为(189.1±23)粒·m-2,比前者减少了88%。两类沙垄土壤种子库的生活型组成都以一年生植物为主,相似性系数为0.711。相同沙垄不同部位的物种相似性系数在0.429～0.600之间,异质性较大,尤其表现在除垄顶之外的其他部位。冬季放牧沙垄和春夏季放牧沙垄地上植被中分别有植物36种和26种,各生活型所有物种均为后者低于前者,其中短命、类短命物种数量减少幅度最大。春夏季放牧沙垄地上植被的物种多样性和均匀度降低,而优势度升高。回归分析显示,冬季放牧沙垄定植群落有21%的变异来源于土壤种子库,而春夏季放牧沙垄有51%的变异来源于土壤种子库,可见不同季节的放牧干扰显著影响土壤种子库及其与地上植被的相关关系。     

古尔班通古特沙漠风沙土微量元素对植被格局的影响

横穿古尔班通古特沙漠,测度了沙漠植被盖度、物种多样性和丰富度等格局参数及微地形、土壤理化性质。在此基础上,借助ICP/MS仪测定了沙漠表层（0~10 cm）和次表层（10~30 cm）风沙土中Cu、Zn、Mo、Fe、Co、Ni、Mn、V、Se、B微量元素含量。利用t-检验和CCA排序的方法,分析了微量元素在风沙土中的空间分布特征和对植被格局的影响。结果显示,地势较高的东部Ni和Mn的含量高于其他地段。地势较低、土壤中水分、有机质、全盐含量较高的西部,Mo、Cu、Zn、Fe、Se含量较高。Fe的高含量值主要出现在分选差、有机质较多的风沙土中。另外,绝大多数微量元素在表层风沙土中的平均含量高于浅层。对比不同微量元素成分条件下的乔灌木和草本层片植物组成和结构特征,发现乔灌木层片基本格局所受影响不明显,草本层片则在沙漠生境下已形成的植物群丛内部发生分化,出现多个小的植物组合。     

Distribution of plant species and species-soil relationship in the east central Gurbantunggut Desert,China

The distribution of plant species and relationships between species and soil factors in the east central part of Gurbantunggut Desert was studied to provide more insight into the flora and determine differences in vegetation across various parts of the desert. Two-way Cluster Analysis showed that the vegetation in the area could be divided into three groups, the first group was dominated by the shrub species, Ephedra przewalskii and the grass species, Carex physodes mainly in areas of flat grounds and gentle slopes; the second group was dominated by C. physodes, Artemisia songorica and A. xerophytica mainly on the slope of sand dunes and the third group was dominated by the shrub species, Haloxylon persicum mainly on the top of sand dunes. There was no difference in plant density between Groups 1 and 2 but there was a significant decrease in Group 3. Soil water under vegetation Group 3 was much lower than that in the other two groups at all soil depths. The EC, organic matter, total P and soluble Na, Ca and Mg varied very similarly with soil water. Canonical correspondence analysis (CCA) satisfactorily assessed the species-soil relations in the area. The distribution of plant species was strongly correlated with the soil factors of water content, organic matter, EC and nutrients. The variations in species occurrence explained by the three CCA axes were about 70%, indicating that some explanatory site variables may exist outside our studied parameters. Soil texture is suggested to be included in future studies to improve the explanation of CCA.     

中国古尔班通古特沙漠东部中心地区植物分布与植物—土壤关系(英文)

The distribution of plant species and relationships between species and soil factors in the east central part of Gurbantunggut Desert was studied to provide more insight into the flora and determine differences in vegetation across various parts of the desert. Two-way Cluster Analysis showed that the vegetation in the area could be divided into three groups, the first group was dominated by the shrub species, Ephedra przewalskii and the grass species, Carex physodes mainly in areas of flat grounds and gentle slopes; the second group was dominated by C. physodes, Artemisia songorica and A. xerophytica mainly on the slope of sand dunes and the third group was dominated by the shrub species, Haloxylon persicum mainly on the top of sand dunes. There was no difference in plant density between Groups 1 and 2 but there was a significant decrease in Group 3. Soil water under vegetation Group 3 was much lower than that in the other two groups at all soil depths. The EC, organic matter, total P and soluble Na, Ca and Mg varied very similarly with soil water. Canonical correspondence analysis(CCA) satisfactorily assessed the species-soil relations in the area. The distribution of plant species was strongly correlated with the soil factors of water content, organic matter, EC and nutrients. The variations in species occurrence explained by the three CCA axes were about 70%, indicating that some explanatory site variables may exist outside our studied parameters. Soil texture is suggested to be included in future studies to improve the explanation of CCA.     

新疆准噶尔盆地典型荒漠区不同景观植被对土壤养分的影响

在群落演替的不同阶段,土壤与植被之间存在不同的互动关系。在新疆准噶尔盆地南缘,选择荒漠-绿洲交错区、沙漠边缘、沙漠腹地典型景观类型取样,监测不同景观条件下土壤水分、土壤有机质、全氮、全磷和植被盖度。结果表明,不同景观植被盖度、土壤水分、土壤养分含量存在显著的差异性(P0.5)。土壤水分含量、植被盖度、土壤养分含量表现为荒漠-绿洲交错区沙漠边缘沙漠腹地的变化趋势;土壤含水量由荒漠-绿洲交错带10.4%递减到沙漠边缘5.8%、沙漠腹地3.5%;植被盖度、土壤养分含量随土壤类型由荒漠灰钙土向风沙土过渡,呈现显著的下降趋势。在相同土壤属性情况下,植被盖度与土壤有机质、土壤全氮之间存在显著的线性正相关关系,表明不同区域内植被在一定程度上影响土壤养分的积累。     

古尔班通古特沙漠南缘干季土壤水分含量空间异质性

土壤水分含量是荒漠植被发育的主要制约因子。对古尔班通古特沙漠南缘个体、群落、丘间地0~100 cm干季土壤水分含量的空间异质性进行分析。结果表明：（1）同尺度下,土壤水分含量随土层深度增加呈显著升高趋势,且表层土壤水分含量的变异系数高于其他土层。（2）个体尺度,梭梭树干基部周围土壤水分含量在垂直和水平方向存在格局分异。垂直方向,土壤水分含量随土层深度增加呈升高趋势。水平方向,随距树干基部距离增加,坡顶土壤水分含量呈升高趋势,坡中和坡底土壤水分含量呈降低趋势,但没有显著差异。（3）梭梭群落尺度上土壤水分含量异质性较强,且呈斑块状分布。（4）地形是影响丘间地尺度土壤水分含量空间分异的主要因素,形成坡底土壤水分含量最高、坡中次之、坡顶最低的空间分布格局。     

古尔班通古特沙漠土壤养分空间分异与干扰的关系

对干扰条件下古尔班通古特沙漠土壤养分空间分异特征进行了研究。分析土壤养分的变化趋势,及干扰与植被和土壤养分之间的关系。研究表明:沙漠边缘至沙漠腹地,全N、全P和全K含量、EC、总盐,可溶性离子HCO-3、SO2-4及Ca2+呈增加趋势。CO2-3、Mg2+、 Cl-、Na+4种离子含量较低,变化规律不明显。除全K含量随公路里程呈较连续的增加外,其他养分条件在公路里程25—75 km ,80—125 km和大于125 km的范围内的变化趋势都呈现高\?低\?显著增高的波动过程,分析表明这种波动性的变化是由干扰造成的。干扰因子数与生物结皮盖度呈显著负相关,但与植物种类与植物盖度都达到了显著正相关的水平。除与pH值呈正相关外,干扰因子数与土壤养分指标都呈负相关,其中与全K、全P、全N和EC的负相关接近于显著或极显著水平。随公路里程增加,生物结皮呈现从地衣苔藓藻类地衣地衣苔藓的过渡,这与土壤养分条件和干扰因子数相对应。人类活动是导致彩克沙漠公路里程80—125 km范围沙漠土壤养分条件的降低的主要原因,受水源和居住地的影响,该段沙漠人类放牧活动影响的最大范围为里程125—145 km之间。人类在沙漠活动的加强会增加沙漠生态干扰源和干扰强度,最终降低沙漠土壤的养分条件。     

巴丹吉林沙漠周边植被特征和物种多样性

通过对巴丹吉林沙漠周边地区的野外调查,以重要值为测度指标,计算物种多样性指数、均匀度指数和优势度指数,对巴丹吉林沙漠周边地区荒漠植物群落物种多样性进行分析。结果表明:(1)该区域共计植物37种,各区域重要值大于0.1的物种中,灌木和半灌木占优势;(2)8个区域的Shannon-Wiener指数、Simpson指数、丰富度指数和优势度指数差异显著,区域5(北部)和7(西部)因含有单物种群落,Shannon-Wiener指数和Simpson指数显著低于其他区域;(3)荒漠植物群落结构层次中,8个区域的灌木层片每个多样性指数均没有显著性差异,草本层片在8个区域中多样性水平差异显著,由于荒漠区植被物种稀少、数量少,灌木层和草本层并未表现出显著差异;(4)巴丹吉林沙漠物种多样性水平较低,生态系统脆弱,对巴丹吉林沙漠周边地区生态系统重建时应该关注各种生态型植被的恢复重建。     

The spatial distribution patterns of biological soil crusts in the Gurbantunggut Desert, Northern Xinjiang, China

The Gurbantunggut Desert, the largest fixed and semi-fixed desert in China, is characterized by a predominant coverage of lichen-dominated biological soil crusts, which serve an indispensable role in sand fixation. Two findings of biological soil crusts have been disclosed from previous field observations: first, distribution of biological soil crusts is selective upon locations; second, species composition varies significantly for the biological soil crusts that are at different developing stages. In this study, a strategy was developed to investigate the spatial distribution of biological soil crusts by coupling remote sensing data and field measurements. A crust index for the Landsat ETM+ data has been developed and applied to detect the lichen-dominated biological soil crusts in the Gurbantunggut Desert. The results indicated the South of the desert encompassed the most abundant biological soil crusts. Besides, biological soil crusts were distributed in uniform density in the South of the desert whereas their distribution patterns become patchier in the rest of the desert. Finally, statistics from the classification revealed that biological soil crusts covered 28.7% of the land in the whole study area. However, it is worth mentioning that the crusts coverage may be underestimated given the fact that detection of crusts from the Landsat ETM+ imagery is viable only if crusts constitute more than 33% of the instantaneous field of view (IFOV) of the Landsat ETM+ sensor.     

生态水文阈值在中国沙区人工植被生态系统管理中的意义

中国北方风沙区是全球沙漠化发展和生态退化最为严重的地区之一。利用植被固沙是遏制沙化土地扩张和生态重建与恢复的主要途径,但人工植被的稳定维持是可持续性风沙治理所长期面临的挑战。作为人工植被生态系统管理的依据,对生态水文阈值的认识及界定具有重要的理论和实践意义。本文以固沙植被中人工种植的灌木和乔木（仅限东部沙区）的盖度与深层土壤水分含量互馈互调作用的长期变化为依据,基于对中国不同气候带沙区典型人工植被系统生态和水文过程的模拟,通过建立生态水文模型初步界定了不同沙区适宜于人工植被生态系统管理的生态水文阈值,对现有沙区固沙植被的生态系统管理和未来风沙区植被建设具有一定的借鉴意义。     

干旱荒漠区退耕地植被演替及土壤水分变化

采用时空互代法,以不同退耕年限退耕地为研究对象,对27个样方植被群落特征和土壤含水量变化特征进行描述性统计分析,使用多元线性回归探究植物演替和土壤水分之间的关系。结果表明：（1）退耕18a间,退耕地植被群落自然演替经历了一年生草本为绝对优势种一年生和多年生草本竞争存在灌木开始出现并逐渐成为优势种的演替过程,物种Margalef指数和Shannon-Wiener指数均出现增大的趋势。（2）土壤水分随着退耕年限的增加也有所增加,且同一退耕年限内,随着土壤深度增加,土壤水分亦逐渐增加。（3）土壤水分对物种Margalef指数、Shannon-Wiener指数具有极显著影响,相比于其他土层,深度为40～50cm的土层对植被生长和演替的贡献最为显著。     

黄土高原西部荒漠草原植被恢复的土壤水分管理研究

针对兰州地区荒漠草原带具有代表性的三种地带性植被：草本植被短花针茅、灌木植被珍珠猪毛菜、荒漠锦鸡儿,对其0~2.0 m的土壤含水量进行了比较,同时对草本和灌木植被的根系进行了测定。结果表明：天然植被对土壤含水量的影响在1 m以上。随着土壤深度的不断加深,土壤含水量呈减少趋势而后趋于稳定。短花针茅区1 m以上含水量比珍珠猪毛菜和荒漠锦鸡区平均高1.08和1.21个百分点;1 m以下土壤含水量变化情况是:珍珠猪毛菜区最高,荒漠锦鸡儿区最低。本文在分析了兰州地区荒漠草原区土壤水分动态的基础上,提出了这一地区植被恢复的对策。     

荒漠绿洲过渡带不同立地条件下物种多样性及其与土壤理化因子的关系

通过于2007-2011年对甘肃河西走廊中部荒漠绿洲过渡带不同立地条件下植被样方的调查,应用 Margalef丰富度指数、Pielou均匀度指数、Shannon-Weiner指数和Simpson指数研究了不同立地条件下物种多样性的变化,并采用典范对应分析方法(CCA)研究植物物种分布与土壤环境因子的关系。结果表明：荒漠绿洲过渡带丘间低地的植被盖度和Margalef、Simpson、Shannon-Wiener、Pielou指数值均最大,流动沙丘条件下最小;Hill多样性指数排序结果显示,随着土壤水分条件的变差,物种多样性同样表现出降低的趋势。按多样性从高到低的顺序,其相应的立地条件分别为丘间低地、缓平沙坡、固定沙丘、半固定沙丘和流动沙丘。选取的土壤因子对过渡带植被分布的影响程度大小为土壤含水量＞pH值＞全钾＞有机质＞速效氮＞全氮＞速效钾＞全磷＞土壤盐分＞速效磷。土壤含水量和pH是荒漠绿洲过渡带植被分布空间差异的最主要影响因子,前3个轴的环境解释率为98％,证明排序可信。     

Species diversity and its relation with soil factors under different site conditions in a desert-oasis ecotone

Vegetation and soil surveys were conducted under different site conditions in 2007–2011 to study species diversity using richness, evenness and diversity indices, in the middle portion of the Heihe River Basin. The relationship between species distribution and soil environmental factors was also studied by Canonical Correspondence Analysis (CCA). Results show that vegetation coverage and species diversity were the highest in the interdune lowland, and the lowest in the mobile dune. Results of the Hill’s index (diversity ordering) shows that species diversity is reduced along decreasing soil water content, and the order of species diversity was interdune lowland, flat slope, fixed dune, semifixed dune and mobile dune. The influence degree of soil factors on vegetation distribution was soil water content > pH > total K > organic matter > available N > total N > available K > total P > saline content > available P. Soil water content and pH were important factors significantly affecting spatial distribution difference of vegetation, the environmental explanation was 98%.     

古尔班通古特沙漠表层土壤凝结水水汽来源特征分析

水分条件是决定干旱沙漠区生态环境的关键因素,凝结水是干旱区植物和低等生物的重要水分来源。利用自制蒸渗计在春、夏、秋3个季节对古尔班通古特沙漠苔藓、地衣、藻类及流沙4种地表类型表层原状土壤凝结水形成进行了观测研究。结果表明,在2 cm、5 cm、10 cm、20 cm 4种高度原状土中,用纱网封底土壤表层2 cm和5 cm土壤的凝结水测定结果能够真实的代表古尔班通古特沙漠不同类型地表土壤凝结水形成特征;凝结水主要集中产生在土壤表层2 cm范围内;凝结水的水汽来源于空气和土壤且以空气来源为主,春季由于表层土壤含水率较高,来自于土壤的水汽所占的比重较高,地衣0~2 cm表层凝结水来源于土壤水汽补充的比例春季为35.5%,夏季和秋季分别降到15.5%和11.3%;秋季55 d的观测结果表明,凝结水形成总量随流沙、藻类、地衣和苔藓依次增加,分别为3.46 mm、4.07 mm、4.89 mm和5.15 mm,说明在干旱的沙漠地带,凝结水是除降水以外补充表层土壤水分最重要的水分来源。     

古尔班通古特沙漠草本植物生物量分配特征

草本植物层片是古尔班通古特沙漠早春生产力的主要贡献者,为深入了解荒漠草本植物在群落水平上对荒漠环境的整体适应性,分析了沙漠中部1 hm2样地中草本植物层片地上与地下生物量分配关系,并验证了草本植物层片生物量与根冠比、物种丰富度以及植株密度之间的关系.结果 表明:(1)类短命植物囊果苔草(Carex physodes)是...     

古尔班通古特沙漠典型灌木群落土壤化学计量特征及其影响因素

对古尔班通古特沙漠3种典型灌木群落77个样地的0～10 cm层土壤及环境因子进行调查取样,分析了土壤有机碳（C）、全氮（N）、全磷（P）及全钾（K）的化学计量特征及其影响因素。结果表明：土壤C、N、P、K含量均值分别为1.057、0.133、0.319、14.11 g·kg^-1,C∶N、C∶P、C∶K、N∶P、N∶K、P∶K值分别为8.124、3.486、0.099、0.434、0.013、0.034。本研究土壤C、N、P养分元素含量及其化学计量比均远低于全球及全国平均水平。C∶N和P含量具有较高的稳定性（CV=19.26%和22.46%）,而K、C∶K、N∶K和P∶K变异性较强（CV=52.17%～80.30%）。土壤化学计量特征在不同群落类型间、不同结皮类型间均有一定差异。C-N、C-K、P-K之间以及土壤养分与其化学计量比之间具有显著的非线性耦合关系,可用二次函数、幂函数或指数函数等非线性函数表征。相关性分析和CCA排序分析表明,除C∶N不受所调查环境因子影响外,土壤化学计量特征主要受经度、纬度、年降水量、生物结皮发育等级、电导率及海拔的影响。